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<document id="98208EC5D9A5BF4E317FDA55069F518B" ID-CLB-Dataset="9404" ID-DOI="10.11646/zootaxa.4890.3.11" ID-GBIF-Dataset="7b2cc9fc-fca2-4360-9ffa-f58dd62c614e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4306184" ID-ZooBank="65B07234-45BE-4946-B30B-45D296DB8565" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607098017240" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="2C0587DCFF89BD20A5B7FACDFD95A826" docLanguage="en" docName="zootaxa.4890.3.11.pdf" docOrigin="Zootaxa 4890 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rutaraphes taiwanicus Jałoszyński 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="450" masterDocId="D03CFFA4FF88BD24A520FFBAFFF2AB33" masterDocTitle="A new species of Rutaraphes Jałoszyński in Taiwan (Coleoptera, Staphylinidae Scydmaeninae)" masterLastPageNumber="450" masterPageNumber="446" pageNumber="447" updateTime="1698847110607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D000828064712DEAD7F2E23505322EF8">A new species of Rutaraphes Jałoszyński in Taiwan (Coleoptera, Staphylinidae Scydmaeninae)</mods:title>
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<mods:namePart id="254E6E725616B61A80ABEE194B0EB21B">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="56518A49AC63362A6113F2228ED32BAF">2020</mods:date>
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<treatment id="2C0587DCFF89BD20A5B7FACDFD95A826" ID-DOI="http://doi.org/10.5281/zenodo.4323754" ID-GBIF-Taxon="170712375" ID-Zenodo-Dep="4323754" LSID="urn:lsid:plazi:treatment:2C0587DCFF89BD20A5B7FACDFD95A826" httpUri="http://treatment.plazi.org/id/2C0587DCFF89BD20A5B7FACDFD95A826" lastPageId="4" lastPageNumber="450" pageId="1" pageNumber="447">
<subSubSection id="ECB66541FF89BD25A5B7FACDFE05AEA1" box="[151,503,1399,1426]" pageId="1" pageNumber="447" type="nomenclature">
<paragraph id="A41336CAFF89BD25A5B7FACDFE05AEA1" blockId="1.[151,503,1399,1462]" box="[151,503,1399,1426]" pageId="1" pageNumber="447">
<heading id="FF5B81A6FF89BD25A5B7FACDFE05AEA1" bold="true" box="[151,503,1399,1426]" fontSize="11" level="1" pageId="1" pageNumber="447" reason="1">
<emphasis id="96D8EAD8FF89BD25A5B7FACDFE05AEA1" bold="true" box="[151,503,1399,1426]" pageId="1" pageNumber="447">
<taxonomicName id="63AC4D49FF89BD25A5B7FACDFE5FAEA1" ID-CoL="8Y97Q" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,429,1399,1426]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="taiwanicus" status="sp. nov.">
<emphasis id="96D8EAD8FF89BD25A5B7FACDFE5FAEA1" bold="true" box="[151,429,1399,1426]" italics="true" pageId="1" pageNumber="447">Rutaraphes taiwanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEB57A3FF89BD25A494FAC2FE05AEA1" box="[436,503,1400,1426]" pageId="1" pageNumber="447" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="ECB66541FF89BD25A5B7FA26FEF9AE85" box="[151,267,1436,1462]" pageId="1" pageNumber="447" type="description">
<paragraph id="A41336CAFF89BD25A5B7FA26FEF9AE85" blockId="1.[151,503,1399,1462]" box="[151,267,1436,1462]" pageId="1" pageNumber="447">
(
<figureCitation id="3C972A4FFF89BD25A5BFFA26FEF0AE85" box="[159,258,1436,1462]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 36" captionStart-2="FIGURE 7" captionStartId-0="2.[151,264,1732,1756]" captionStartId-1="3.[151,264,834,858]" captionStartId-2="3.[151,250,1832,1856]" captionTargetBox-0="[164,1423,394,1695]" captionTargetBox-1="[163,1425,186,805]" captionTargetBox-2="[164,1423,882,1808]" captionTargetId-0="figure-109@2.[151,1436,381,1709]" captionTargetId-1="figure-19@3.[158,1430,181,810]" captionTargetId-2="figure-42@3.[164,1423,882,1808]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURES 12. Rutaraphes taiwanicus sp. n. Dorsal habitus (1) and head of male holotype (2)." captionText-1="FIGURES 36. Rutaraphes taiwanicus sp. n. Aedeagus in ventral (3), left lateral (4), dorsal (5), and right lateral (6) views." captionText-2="FIGURE 7. Distribution and key characters (head of male and aedeagus) of Rutaraphes species." figureDoi-0="http://doi.org/10.5281/zenodo.4306186" figureDoi-1="http://doi.org/10.5281/zenodo.4306188" figureDoi-2="http://doi.org/10.5281/zenodo.4306192" httpUri-0="https://zenodo.org/record/4306186/files/figure.png" httpUri-1="https://zenodo.org/record/4306188/files/figure.png" httpUri-2="https://zenodo.org/record/4306192/files/figure.png" pageId="1" pageNumber="447">Figs 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF89BD25A5B7FA5EFC9FAD75" pageId="1" pageNumber="447" type="materials_examined">
<paragraph id="A41336CAFF89BD25A5B7FA5EFC9FAD75" blockId="1.[151,1437,1508,2039]" pageId="1" pageNumber="447">
<emphasis id="96D8EAD8FF89BD25A5B7FA5EFEAEAECD" bold="true" box="[151,348,1508,1534]" pageId="1" pageNumber="447">Material studied</emphasis>
.
<materialsCitation id="14C43C97FF89BD25A448FA5EFC9FAD75" ID-GBIF-Occurrence="2990014301" collectingDate="2013-08-27" collectionCode="TAIWAN, V, NMNS" elevation="2216" latitude="24.4964" location="Taipingshan forest" longLatPrecision="7" longitude="121.525" pageId="1" pageNumber="447" specimenCount="2" stateProvince="Yilan" typeStatus="holotype">
<emphasis id="96D8EAD8FF89BD25A448FA5EFE2CAECD" bold="true" box="[360,478,1508,1534]" pageId="1" pageNumber="447">
<typeStatus id="7B178868FF89BD25A448FA5EFE2BAECD" box="[360,473,1508,1534]" pageId="1" pageNumber="447" type="holotype">Holotype</typeStatus>
:
</emphasis>
<emphasis id="96D8EAD8FF89BD25A4DEFA5EFD73AECD" bold="true" box="[510,641,1508,1534]" pageId="1" pageNumber="447">
(
<collectionCode id="C2BDAE0FFF89BD25A725FA5EFD88AECD" box="[517,634,1508,1534]" pageId="1" pageNumber="447">TAIWAN</collectionCode>
)
</emphasis>
: two labels: “
<collectionCode id="C2BDAE0FFF89BD25A635FA5EFC75AECD" box="[789,903,1508,1534]" pageId="1" pageNumber="447">TAIWAN</collectionCode>
,
<collectingRegion id="6668F828FF89BD25A6B0FA5EFC22AECD" box="[912,976,1508,1534]" country="Taiwan" name="Yilan" pageId="1" pageNumber="447">Yilan</collectingRegion>
, /
<location id="A1736011FF89BD25A6C5FA5EFB44AECD" LSID="urn:lsid:plazi:treatment:2C0587DCFF89BD20A5B7FACDFD95A826:A1736011FF89BD25A6C5FA5EFB44AECD" box="[997,1206,1508,1534]" latitude="24.4964" longLatPrecision="7" longitude="121.525" name="Taipingshan forest" pageId="1" pageNumber="447" stateProvince="Yilan">Taipingshan forest</location>
, /
<geoCoordinate id="C198500DFF89BD25A1ECFA5EFAD5AECD" box="[1228,1319,1508,1534]" degrees="24.4964" direction="north" orientation="latitude" pageId="1" pageNumber="447" precision="5" value="24.4964">24.4964</geoCoordinate>
<geoCoordinate id="C198500DFF89BD25A00CFA5EFA6AAECC" box="[1324,1432,1508,1535]" degrees="121.5250" direction="east" orientation="longitude" pageId="1" pageNumber="447" precision="5" value="121.525">121.5250</geoCoordinate>
, /
<quantity id="63549B2FFF89BD25A58AF9B2FF09AD11" box="[170,251,1544,1570]" metricMagnitude="3" metricUnit="m" metricValue="2.216" pageId="1" pageNumber="447" unit="m" value="2216.0">
<elevation id="2F81D1F9FF89BD25A58AF9B2FF09AD11" box="[170,251,1544,1570]" metricMagnitude="3" metricUnit="m" metricValue="2.216" pageId="1" pageNumber="447" unit="m" value="2216.0">2216m</elevation>
</quantity>
,
<date id="D012100AFF89BD25A429F9B2FE61AD11" box="[265,403,1544,1570]" pageId="1" pageNumber="447" value="2013-08-27">
<collectingDate id="C056E9E2FF89BD25A429F9B2FE61AD11" box="[265,403,1544,1570]" pageId="1" pageNumber="447" value="2013-08-27">27.viii.2013</collectingDate>
</date>
, sift / TW13,
<collectionCode id="C2BDAE0FFF89BD25A71CF9B2FDA2AD11" box="[572,592,1544,1570]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="447" type="Museum">V</collectionCode>
.Grebennikov” [white, handwritten], “
<taxonomicName id="63AC4D49FF89BD25A123F9B2FB5CAD11" box="[1027,1198,1544,1570]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="genus">
<emphasis id="96D8EAD8FF89BD25A123F9B2FB5CAD11" box="[1027,1198,1544,1570]" italics="true" pageId="1" pageNumber="447">RUTARAPHES</emphasis>
</taxonomicName>
/
<taxonomicName id="63AC4D49FF89BD25A1EBF9B3FAB6AD11" authorityName="Jałoszyński" authorityYear="2020" box="[1227,1348,1545,1570]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="taiwanicus">
<emphasis id="96D8EAD8FF89BD25A1EBF9B3FAB6AD11" box="[1227,1348,1545,1570]" italics="true" pageId="1" pageNumber="447">taiwanicus</emphasis>
</taxonomicName>
m. / P. JAŁOSZYŃSKI, 2020 /
<typeStatus id="7B178868FF89BD25A48EF996FDAAAD75" box="[430,600,1580,1606]" pageId="1" pageNumber="447" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="C2BDAE0FFF89BD25A62AF996FC93AD75" box="[778,865,1580,1606]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="447">NMNS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF89BD25A5E7F9EAFEB4ADE5" pageId="1" pageNumber="447" type="diagnosis">
<paragraph id="A41336CAFF89BD25A5E7F9EAFEB4ADE5" blockId="1.[151,1437,1508,2039]" pageId="1" pageNumber="447">
<emphasis id="96D8EAD8FF89BD25A5E7F9EAFEB0AD59" bold="true" box="[199,322,1616,1642]" pageId="1" pageNumber="447">Diagnosis.</emphasis>
Vertex in male with pair of oval, narrowly separated impressions densely filled with short setae; pro- notum with two pairs of distinct antebasal pits, inner pair narrowly separated by tiny horn-like projection; aedeagus in lateral view lacking dorsal projection just above basal orifice, in ventral view conspicuously asymmetrical, al-most C-shaped.
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF89BD20A5E7F95AFE50AA9E" lastPageId="4" lastPageNumber="450" pageId="1" pageNumber="447" type="description">
<paragraph id="A41336CAFF89BD25A5E7F95AFC0BAC2D" blockId="1.[151,1437,1508,2039]" pageId="1" pageNumber="447">
<emphasis id="96D8EAD8FF89BD25A5E7F95AFEABADC9" bold="true" box="[199,345,1760,1786]" pageId="1" pageNumber="447">Description.</emphasis>
Body of male (
<figureCitation id="3C972A4FFF89BD25A72BF95AFDBFADC9" box="[523,589,1760,1786]" captionStart="FIGURES 12" captionStartId="2.[151,264,1732,1756]" captionTargetBox="[164,1423,394,1695]" captionTargetId="figure-109@2.[151,1436,381,1709]" captionTargetPageId="2" captionText="FIGURES 12. Rutaraphes taiwanicus sp. n. Dorsal habitus (1) and head of male holotype (2)." figureDoi="http://doi.org/10.5281/zenodo.4306186" httpUri="https://zenodo.org/record/4306186/files/figure.png" pageId="1" pageNumber="447">Fig. 1</figureCitation>
) slender, moderately convex, BL
<quantity id="63549B2FFF89BD25A6EFF95AFBC5ADC8" box="[975,1079,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="447" unit="mm" value="1.23">1.23 mm</quantity>
; pigmentation light brown with appendages indistinctly lighter than body, setae distinctly lighter than cuticle.
</paragraph>
<paragraph id="A41336CAFF89BD26A5E7F892FC3DAA56" blockId="1.[151,1437,1508,2039]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="448" pageId="1" pageNumber="447">
Head (
<figureCitation id="3C972A4FFF89BD25A431F892FEA7AC71" box="[273,341,1832,1858]" captionStart="FIGURES 12" captionStartId="2.[151,264,1732,1756]" captionTargetBox="[164,1423,394,1695]" captionTargetId="figure-109@2.[151,1436,381,1709]" captionTargetPageId="2" captionText="FIGURES 12. Rutaraphes taiwanicus sp. n. Dorsal habitus (1) and head of male holotype (2)." figureDoi="http://doi.org/10.5281/zenodo.4306186" httpUri="https://zenodo.org/record/4306186/files/figure.png" pageId="1" pageNumber="447">Fig. 2</figureCitation>
) flattened and subtriangular with long clypeus distinctly broadening anterad, broadest at eyes, HL
<quantity id="63549B2FFF89BD25A5B7F8F6FF0DAC54" box="[151,255,1868,1895]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="447" unit="mm" value="0.2">0.20 mm</quantity>
, HW
<quantity id="63549B2FFF89BD25A464F8F6FE5FAC54" box="[324,429,1868,1895]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="447" unit="mm" value="0.2">0.20 mm</quantity>
; tempora in dorsal view much shorter then eyes, strongly convergent posteriorly, vertex with well-marked posterior margin and pair of large, oval impressions filled densely with short setae, so that cuticle within impressions is not visible, impressions narrowly separated at middle by tiny horn-like tubercle, in lateral view distinctly projecting dorsoposterad; frons in front of impressions flattened; supraantennal tubercles distinctly elevated; eyes large, strongly convex, finely faceted, oval, lacking posterior emargination. Punctures on frons and vertex inconspicuous, very fine; setae unevenly distributed, mainly along posterior margin of vertex, laterad each vertexal impression and between supraantennal tubercles, short, sparse and suberect. Antennae slender, with indis-tinctly demarcated trimerous club, strikingly short, slightly shorter than head and pronotum combined, AnL
<quantity id="63549B2FFF8ABD26A04BFF65FF34AA2E" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="448" unit="mm" value="0.43">0.43 mm</quantity>
; scape and pedicel each strongly elongate, antennomere 3 indistinctly transverse, 4 about as long as broad, 5 indistinctly elongate, 6 about as long as broad, 7 barely noticeably transverse, 810 each distinctly transverse, 11 indistinctly shorter than 9 and 10 combined, about 1.5 × as long as broad.
</paragraph>
<caption id="F0D36642FF8ABD26A5B7F97EFB93ADEF" ID-DOI="http://doi.org/10.5281/zenodo.4306186" ID-Zenodo-Dep="4306186" box="[151,1121,1732,1756]" httpUri="https://zenodo.org/record/4306186/files/figure.png" pageId="2" pageNumber="448" startId="2.[151,264,1732,1756]" targetBox="[164,1423,394,1695]" targetPageId="2">
<paragraph id="A41336CAFF8ABD26A5B7F97EFB93ADEF" blockId="2.[151,1121,1732,1756]" box="[151,1121,1732,1756]" pageId="2" pageNumber="448">
<emphasis id="96D8EAD8FF8ABD26A5B7F97EFEC8ADEF" bold="true" box="[151,314,1732,1756]" pageId="2" pageNumber="448">FIGURES 12.</emphasis>
<taxonomicName id="63AC4D49FF8ABD26A461F97EFDD8ADEF" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[321,554,1732,1756]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="448" phylum="Arthropoda" rank="species" species="taiwanicus" status="sp. nov.">
<emphasis id="96D8EAD8FF8ABD26A461F97EFDD8ADEF" box="[321,554,1732,1756]" italics="true" pageId="2" pageNumber="448">Rutaraphes taiwanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEB57A3FF8ABD26A710F97EFD95ADE8" box="[560,615,1732,1755]" pageId="2" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
Dorsal habitus (1) and head of male holotype (2).
</paragraph>
</caption>
<paragraph id="A41336CAFF8ABD26A5E7F8B1FEB0ACEB" blockId="2.[151,1437,1802,2008]" pageId="2" pageNumber="448">
Pronotum broadest slightly anterior to middle but weakly narrowing posterad, PL
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, PW
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; anterior margin and sides in anterior half rounded, sides in posterior half nearly straight and with distinct, sharply marked lateral carinae in posterior third; posterior corners nearly right; posterior margin straight; base with two pairs of distinct, sharply marked pits: outer pits large and elongate, adjacent to lateral pronotal carinae, inner pits circular and separated only by narrow, slightly raised longitudinal carina. Punctures on disc fine, inconspicuous; setae dense, short, suberect.
</paragraph>
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<paragraph id="A41336CAFF8BBD27A5B7FCF8FA88A869" blockId="3.[151,1402,834,858]" box="[151,1402,834,858]" pageId="3" pageNumber="449">
<emphasis id="96D8EAD8FF8BBD27A5B7FCF8FEC9A869" bold="true" box="[151,315,834,858]" pageId="3" pageNumber="449">FIGURES 36.</emphasis>
<taxonomicName id="63AC4D49FF8BBD27A461FCF8FDD8A869" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[321,554,834,858]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="species" species="taiwanicus" status="sp. nov.">
<emphasis id="96D8EAD8FF8BBD27A461FCF8FDD8A869" box="[321,554,834,858]" italics="true" pageId="3" pageNumber="449">Rutaraphes taiwanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DEB57A3FF8BBD27A710FCF8FD95A86A" box="[560,615,834,857]" pageId="3" pageNumber="449" rank="species">sp. n.</taxonomicNameLabel>
Aedeagus in ventral (3), left lateral (4), dorsal (5), and right lateral (6) views.
</paragraph>
</caption>
<caption id="F0D36642FF8BBD27A5B7F892FB9EAC72" ID-DOI="http://doi.org/10.5281/zenodo.4306192" ID-Zenodo-Dep="4306192" box="[151,1132,1832,1857]" httpUri="https://zenodo.org/record/4306192/files/figure.png" pageId="3" pageNumber="449" startId="3.[151,250,1832,1856]" targetBox="[164,1423,882,1808]" targetPageId="3">
<paragraph id="A41336CAFF8BBD27A5B7F892FB9EAC72" blockId="3.[151,1132,1832,1857]" box="[151,1132,1832,1857]" pageId="3" pageNumber="449">
<emphasis id="96D8EAD8FF8BBD27A5B7F892FEE1AC72" bold="true" box="[151,275,1832,1857]" pageId="3" pageNumber="449">FIGURE 7.</emphasis>
Distribution and key characters (head of male and aedeagus) of
<taxonomicName id="63AC4D49FF8BBD27A681F893FBE4AC72" box="[929,1046,1833,1857]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="96D8EAD8FF8BBD27A681F893FBE4AC72" box="[929,1046,1833,1857]" italics="true" pageId="3" pageNumber="449">Rutaraphes</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="A41336CAFF8BBD27A5E7F8D5FC7CACC6" blockId="3.[151,1436,1903,2037]" pageId="3" pageNumber="449">
Elytra together oval, broadest slightly in front of middle, EL
<quantity id="63549B2FFF8BBD27A6A4F8D5FC1EACBA" box="[900,1004,1903,1929]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="449" unit="mm" value="0.73">0.73 mm</quantity>
, EW
<quantity id="63549B2FFF8BBD27A10DF8D5FB67ACBA" box="[1069,1173,1903,1929]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="449" unit="mm" value="0.48">0.48 mm</quantity>
, EI 1.53; humeral calli distinctly elevated and elongate; basal elytral pits situated in deep but small circular basal impressions; elytral api-ces separately rounded. Punctures more distinct than those on pronotum and dense, but superficial and with diffuse margins; setae similar to those on pronotum but indistinctly sparser.
</paragraph>
<paragraph id="A41336CAFF8CBD20A5E7FF2DFD70AB82" blockId="4.[151,1437,151,789]" box="[199,642,151,177]" pageId="4" pageNumber="450">Hind wings well developed, functional.</paragraph>
<paragraph id="A41336CAFF8CBD20A5E7FF01FD2AABE6" blockId="4.[151,1437,151,789]" box="[199,728,187,213]" pageId="4" pageNumber="450">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="A41336CAFF8CBD20A5E7FF65FAEFAABA" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="450">
Aedeagus (
<figureCitation id="3C972A4FFF8CBD20A466FF65FE5EABC9" box="[326,428,223,250]" captionStart="FIGURES 36" captionStartId="3.[151,264,834,858]" captionTargetBox="[163,1425,186,805]" captionTargetId="figure-19@3.[158,1430,181,810]" captionTargetPageId="3" captionText="FIGURES 36. Rutaraphes taiwanicus sp. n. Aedeagus in ventral (3), left lateral (4), dorsal (5), and right lateral (6) views." figureDoi="http://doi.org/10.5281/zenodo.4306188" httpUri="https://zenodo.org/record/4306188/files/figure.png" pageId="4" pageNumber="450">Figs 36</figureCitation>
) strikingly asymmetrical, so that it is difficult to define ventral and dorsal sides without knowing the orientation inside abdomen; AeL
<quantity id="63549B2FFF8CBD20A7BCFEB8FCF0AA2E" box="[668,770,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="450" unit="mm" value="0.25">0.25 mm</quantity>
; median lobe in ventral view (= abparameral view) strongly curved, nearly C-shaped, with diaphragm subapical and oblique in relation to the long axis of aedeagus, apical pro-jections shifted to the right side, asymmetrical and complicated, composed of twisted plates; dorsal wall of median lobe above basal orifice with narrow median longitudinal fin; basal orifice surrounded by thick collar.
</paragraph>
<paragraph id="A41336CAFF8CBD20A5E7FE29FE50AA9E" blockId="4.[151,1437,151,789]" box="[199,418,403,429]" pageId="4" pageNumber="450">
<emphasis id="96D8EAD8FF8CBD20A5E7FE29FEECAA9E" bold="true" box="[199,286,403,429]" pageId="4" pageNumber="450">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF8CBD20A5E7FE0CFD7AAAE2" box="[199,648,438,465]" pageId="4" pageNumber="450" type="distribution">
<paragraph id="A41336CAFF8CBD20A5E7FE0CFD7AAAE2" blockId="4.[151,1437,151,789]" box="[199,648,438,465]" pageId="4" pageNumber="450">
<emphasis id="96D8EAD8FF8CBD20A5E7FE0CFEABAAE3" bold="true" box="[199,345,438,464]" pageId="4" pageNumber="450">Distribution</emphasis>
(
<figureCitation id="3C972A4FFF8CBD20A449FE0DFE5FAAE2" box="[361,429,439,465]" captionStart="FIGURE 7" captionStartId="3.[151,250,1832,1856]" captionTargetBox="[164,1423,882,1808]" captionTargetId="figure-42@3.[164,1423,882,1808]" captionTargetPageId="3" captionText="FIGURE 7. Distribution and key characters (head of male and aedeagus) of Rutaraphes species." figureDoi="http://doi.org/10.5281/zenodo.4306192" httpUri="https://zenodo.org/record/4306192/files/figure.png" pageId="4" pageNumber="450">Fig. 7</figureCitation>
). Northern
<collectingCountry id="DCBB765AFF8CBD20A70EFE0DFD76AAE2" box="[558,644,439,465]" name="Taiwan" pageId="4" pageNumber="450">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF8CBD20A5E7FE61FD72AAC6" box="[199,640,475,501]" pageId="4" pageNumber="450" type="etymology">
<paragraph id="A41336CAFF8CBD20A5E7FE61FD72AAC6" blockId="4.[151,1437,151,789]" box="[199,640,475,501]" pageId="4" pageNumber="450">
<emphasis id="96D8EAD8FF8CBD20A5E7FE61FEBCAAC6" bold="true" box="[199,334,475,501]" pageId="4" pageNumber="450">Etymology.</emphasis>
After the island of
<collectingCountry id="DCBB765AFF8CBD20A706FE61FD8EAAC6" box="[550,636,475,501]" name="Taiwan" pageId="4" pageNumber="450">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB66541FF8CBD20A5E7FE45FD95A826" pageId="4" pageNumber="450" type="discussion">
<paragraph id="A41336CAFF8CBD20A5E7FE45FD95A826" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="450">
<emphasis id="96D8EAD8FF8CBD20A5E7FE45FEC9A92A" bold="true" box="[199,315,511,537]" pageId="4" pageNumber="450">Remarks.</emphasis>
<taxonomicName id="63AC4D49FF8CBD20A464FE45FDB5A92B" authorityName="Jałoszyński" authorityYear="2020" box="[324,583,511,537]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="taiwanicus">
<emphasis id="96D8EAD8FF8CBD20A464FE45FDB5A92B" box="[324,583,511,537]" italics="true" pageId="4" pageNumber="450">Rutaraphes taiwanicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="63AC4D49FF8CBD20A7C0FE45FC7DA92A" authorityName="Jaloszynski" authorityYear="2015" box="[736,911,511,537]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="shikokuensis">
<emphasis id="96D8EAD8FF8CBD20A7C0FE45FC7DA92A" box="[736,911,511,537]" italics="true" pageId="4" pageNumber="450">R. shikokuensis</emphasis>
</taxonomicName>
in male head modifications, with two distinct oval setal patches instead of a single median semicircular group of dense short setae (
<figureCitation id="3C972A4FFF8CBD20A16BFD99FB7DA90E" box="[1099,1167,547,573]" captionStart="FIGURE 7" captionStartId="3.[151,250,1832,1856]" captionTargetBox="[164,1423,882,1808]" captionTargetId="figure-42@3.[164,1423,882,1808]" captionTargetPageId="3" captionText="FIGURE 7. Distribution and key characters (head of male and aedeagus) of Rutaraphes species." figureDoi="http://doi.org/10.5281/zenodo.4306192" httpUri="https://zenodo.org/record/4306192/files/figure.png" pageId="4" pageNumber="450">Fig. 7</figureCitation>
). These two species dif-fer also clearly in the shape of the pronotum (more elongate and with distinctly sinuate sides in posterior half in
<taxonomicName id="63AC4D49FF8CBD20A0A4FDFDFED7A9B6" authorityName="Jaloszynski" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="shikokuensis">
<emphasis id="96D8EAD8FF8CBD20A0A4FDFDFED7A9B6" italics="true" pageId="4" pageNumber="450">R. shikokuensis</emphasis>
</taxonomicName>
), inner antebasal pronotal pits in
<taxonomicName id="63AC4D49FF8CBD20A7BCFDD1FCBBA9B6" authorityName="Jaloszynski" authorityYear="2015" box="[668,841,619,645]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="shikokuensis">
<emphasis id="96D8EAD8FF8CBD20A7BCFDD1FCBBA9B6" box="[668,841,619,645]" italics="true" pageId="4" pageNumber="450">R. shikokuensis</emphasis>
</taxonomicName>
separated by a round tubercle vs. narrow carina in
<taxonomicName id="63AC4D49FF8CBD20A0A4FDD1FEE2A99B" authorityName="Jałoszyński" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="taiwanicus">
<emphasis id="96D8EAD8FF8CBD20A0A4FDD1FEE2A99B" italics="true" pageId="4" pageNumber="450">R. taiwanicus</emphasis>
</taxonomicName>
, and the form of the aedeagus. In
<taxonomicName id="63AC4D49FF8CBD20A7ADFD35FCD7A99B" authorityName="Jałoszyński" authorityYear="2020" box="[653,805,655,680]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="taiwanicus">
<emphasis id="96D8EAD8FF8CBD20A7ADFD35FCD7A99B" box="[653,805,655,680]" italics="true" pageId="4" pageNumber="450">R. taiwanicus</emphasis>
</taxonomicName>
, the median lobe is even more asymmetrical than that in
<taxonomicName id="63AC4D49FF8CBD20A5B7FD09FEB7A9FE" authorityName="Jaloszynski" authorityYear="2015" box="[151,325,691,717]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="shikokuensis">
<emphasis id="96D8EAD8FF8CBD20A5B7FD09FEB7A9FE" box="[151,325,691,717]" italics="true" pageId="4" pageNumber="450">R. shikokuensis</emphasis>
</taxonomicName>
, strongly curved and with twisted, complicated apical projections (in
<taxonomicName id="63AC4D49FF8CBD20A177FD09FAF7A9FE" authorityName="Jaloszynski" authorityYear="2015" box="[1111,1285,691,717]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="shikokuensis">
<emphasis id="96D8EAD8FF8CBD20A177FD09FAF7A9FE" box="[1111,1285,691,717]" italics="true" pageId="4" pageNumber="450">R. shikokuensis</emphasis>
</taxonomicName>
the aedeagus in ventral view is weakly asymmetrical, and apical projections are entirely hidden behind median lobe, not project-ing distad beyond it, as in
<taxonomicName id="63AC4D49FF8CBD20A49EFD41FDA4A827" authorityName="Jałoszyński" authorityYear="2020" box="[446,598,763,788]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="450" phylum="Arthropoda" rank="species" species="taiwanicus">
<emphasis id="96D8EAD8FF8CBD20A49EFD41FDA4A827" box="[446,598,763,788]" italics="true" pageId="4" pageNumber="450">R. taiwanicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>