179 lines
17 KiB
XML
179 lines
17 KiB
XML
<document ID-CLB-Dataset="21365" ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. & Hofman, Sebastian" docDate="2020" docId="204FCF065FC851CBBF6D2424D500690E" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Caucasopsis olsavskyi Mumladze & Grego 2020, sp. nov." docType="treatment" docUuid="F220AA48-63FE-4C96-A2CB-05D7AFEEF821" docUuidSource="ZooBank" docVersion="7" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1732936627699" updateUser="ExternalLinkService">
|
|
<mods:mods id="3CCA14CBF0E5C6FB6126FB66CABBFBCF" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="895F3438562B9CA62D5BFEF6C955054A">
|
|
<mods:title id="B8F254FEE36A9599C663CF33F37BF005">Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="4CC31FA68A1CA10A3218DF36405A50D2" type="personal">
|
|
<mods:role id="0A11A86018313614467466F58C9BCAC3">
|
|
<mods:roleTerm id="3D6F161122C4B8945E2E2D20B865D146">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="DDE1EFF17D7A6CDB70BA0F656F813B0B">Grego, Jozef</mods:namePart>
|
|
<mods:affiliation id="13CB45E6CAA3FD4208553F708F3687F9">Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="5EF66E31598224F17C0FD54E1AF51094" type="personal">
|
|
<mods:role id="F642212719486B6FE6ED0A1A15FFF186">
|
|
<mods:roleTerm id="02BD51F6E0A1FCCF90A0281165876A4E">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="073C41BBAAF4927CDF6CFF6913FF922C">Mumladze, Levan</mods:namePart>
|
|
<mods:affiliation id="1CB1F06CB5A6110F7633799CEB9ABADA">Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
|
|
<mods:nameIdentifier id="AAD13D5A9903F7B1D4C5B6FE632E94AD" type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="EBCCBF3E919AFDD4ABEBFCBF45F92C16" type="personal">
|
|
<mods:role id="D92C9A2B3037446F19C49BBF540024BB">
|
|
<mods:roleTerm id="171C5FD78FC84FF52E79E2DF56042A9A">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="12A354FA01CD98A36C56219AC53E6DDA">Falniowski, Andrzej</mods:namePart>
|
|
<mods:affiliation id="CE9D4CD5DB28DEF871C4BA1C346E224F">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="CA83AA14AC6D9A709ECD44381CE767D3" type="personal">
|
|
<mods:role id="053DFA94BA0DC8C05305CCCC8F94B4F4">
|
|
<mods:roleTerm id="5D9411CEF9E81C3654174F908F38D99F">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="79127BE53535080D0853A3608660F3BA">Osikowski, Artur</mods:namePart>
|
|
<mods:affiliation id="E947A1A52ECC71C073E5335F60B83BE6">Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
|
|
<mods:nameIdentifier id="7ED0FBE4D729DEB752AB09B04CCA64D7" type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
|
|
<mods:nameIdentifier id="98A44311A8D2CDD13BC8E27F105C63F1" type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="C5679DF540EB8CB5776538A626E3DE22" type="personal">
|
|
<mods:role id="3BE270E7291640E112457F2B96C92028">
|
|
<mods:roleTerm id="86D694E691000CDFC61D0552877E7DB3">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="3FBC6C776729322680D5551E27982F18">Rysiewska, Aleksandra</mods:namePart>
|
|
<mods:affiliation id="DB3E822FD9CAD07BB018365A5F634988">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="59701B3BC9F56BE5F9E7E29C21BF39F7" type="personal">
|
|
<mods:role id="23176E5D2439B28AB61260D7EA1E3F2B">
|
|
<mods:roleTerm id="7C7CFE31637A13FF4FC4715F1C03BBA8">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="BDAAEB9EBBC99BDEB187ED91855CDC5D">Palatov, Dimitry M.</mods:namePart>
|
|
<mods:affiliation id="EF08F3DD8C5529C5017500633A85FD52">Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="80B2336FC0DC90ED3E7027A05FE369C6" type="personal">
|
|
<mods:role id="9205B5E8BC256214C29B7F973A5B916A">
|
|
<mods:roleTerm id="A63850733E68226B4A7175710D1490AB">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="B20343687BBD7AAEE3319E7DE047F64C">Hofman, Sebastian</mods:namePart>
|
|
<mods:affiliation id="25A3A5683FED79AD0E7B1D2F4A0D04A3">Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="F5BDA8C78E305DDD7C50AE99B7794953">text</mods:typeOfResource>
|
|
<mods:relatedItem id="ACF86245A4D079C6F7E5359181554A51" type="host">
|
|
<mods:titleInfo id="3F4CD655ADAAAF5686DD5ADB4D6B337A">
|
|
<mods:title id="AF52B87BAE00AE7445F7A408DA21E429">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="9521767879259B63412F34DA57CE7878">
|
|
<mods:date id="3D89DF5582EC01C460E9495AE0296E2C">2020</mods:date>
|
|
<mods:detail id="67A3F4806FE9014D88A85901718DD16A" type="volume">
|
|
<mods:number id="9E1639852ADA71842F58BD94111A71B8">955</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="C88B252617E952100C44AA965BDFFBFA" unit="page">
|
|
<mods:start id="D9212CCEB7823D76EF2A3C6DDC90FA4E">1</mods:start>
|
|
<mods:end id="68ED41A7ABEEBC4E7D1768A9F077270F">77</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="ADC87CD29E08474656ECA301E3AC3462">
|
|
<mods:url id="72FC2684EE997E8621C51CEC31BB25FD">http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="A5993A097B4A9D42049F9FFE853D9F2F">journal article</mods:classification>
|
|
<mods:identifier id="947A06EDFF35789371EC2565261C133A" type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
|
|
<mods:identifier id="3B4395D7C07968DD7EFF7726037A9A91" type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
|
|
<mods:identifier id="CEE605C84D63556B9E7E5A086B784B98" type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
|
|
<mods:identifier id="49752CCB050FB9F04E37F93ED05259DE" type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="204FCF065FC851CBBF6D2424D500690E" ID-GBIF-Taxon="166096056" LSID="urn:lsid:zoobank.org:act:F220AA48-63FE-4C96-A2CB-05D7AFEEF821" httpUri="http://treatment.plazi.org/id/204FCF065FC851CBBF6D2424D500690E" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_phylum="Mollusca" scope_subFamily="Sadlerianinae" scope_superFamily="Truncatelloidea">
|
|
<subSubSection id="9F028B93B60E230B78E99C068D1D2E61" pageId="0" pageNumber="1" type="nomenclature">
|
|
<paragraph id="701E5E25DBF1AF1AE81E92C358E929A4" pageId="0" pageNumber="1">
|
|
<taxonomicName id="FE730F8E4F32B9710A6C435122B2B248" ID-CoL="7QL4Z" LSID="204FCF06-5FC8-51CB-BF6D-2424D500690E" authority="Mumladze & Grego" authorityName="Mumladze & Grego" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis olsavskyi" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="olsavskyi" status="sp. nov.">Caucasopsis olsavskyi Mumladze & Grego</taxonomicName>
|
|
<taxonomicNameLabel id="AFF0157FA366DC4E896097EBAC987BED" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
|
<figureCitation id="A45E3E45EB2B38368126AEB25DD29B11" captionStart="Plate 9" captionStartId="F21" captionText="Plate 9. 1 Caucasopsis olsavskyi sp. nov., Samegrelo, Chkhorotsku, Nazodelavo Cave, holotype 2 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, holotype 3 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Kachara Cave 4 C. egrisi sp. nov., Samegrelo, Pirveli Balda, Motena Cave form 5, 6 C. egrisi sp. nov., Samegrelo, Pirveli Balda, spring in village: 5 holotype 6 paratype 7 Imeretiopsis prometheus sp. nov., Imereti, Kumistavi, Prometheus Cave, holotype 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni (Tskhal-Tsiteli) Cave Spring, holotype 9 I. nakeralaensis sp. nov., Imereti, Tikibuli, spring above Tikibuli-Nikortsminda road to Nakerala pass, holotype 10 C. letsurtsume sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave, robust morphotype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego." figureDoi="10.3897/zookeys.955.51983.plate9" httpUri="https://binary.pensoft.net/fig/439172" pageId="0" pageNumber="1">Plates 9 (1); 10(3)</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4CC364CEFA29A8D353819CE59C256744" pageId="0" pageNumber="1" type="type locality">
|
|
<paragraph id="2ECACC384D2E94ED34ABC967888F46CE" pageId="0" pageNumber="1">Type locality.</paragraph>
|
|
<paragraph id="B65096BEFE6B379B1312E0FD4EB5CE14" pageId="0" pageNumber="1">
|
|
Georgia • Samegrelo, Chkhorotsku, Nazodelavo Cave (ნაზოდელავოს
|
|
<normalizedToken id="C1B47322A78A781DC61446F875377C8C" originalValue="მღვიმე">მღვიმე</normalizedToken>
|
|
);
|
|
<geoCoordinate id="12FB68340B65589DC7021869C2B29BC5" degrees="42" direction="north" minutes="30" orientation="latitude" precision="15" seconds="18" value="42.505">42°30'18"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="1191F5B6099C05F9536E35A5D037AB19" degrees="42" direction="east" minutes="13" orientation="longitude" precision="15" seconds="15" value="42.220833">42°13'15"E</geoCoordinate>
|
|
; 275 m a.s.l.; sandy sediment in cave stream.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="16E3607D671D06C19B14DD14E7E653DC" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="0E2EE2FB71DE21B627676B0C50510048" pageId="0" pageNumber="1">Material.</paragraph>
|
|
<paragraph id="3A4CEC68519C8637DD9C9C5EA4359812" pageId="0" pageNumber="1">
|
|
<emphasis id="4E8BA03708F48854924B9D9A6785221C" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
|
: Georgia • 1 adult, dry; type locality; 11 May 2018; J. Grego, L. Mumladze and M.
|
|
<normalizedToken id="32E293E6865ACD4B8442E9661D05328F" originalValue="Olšavský">Olsavsky</normalizedToken>
|
|
leg.; ISU FM-T009-H.
|
|
<emphasis id="09EDBF39E0659AC5CBEFB265F71FEB5C" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
|
: Georgia • same as for holotype; ISU FM-T009-P1/ dry, NHMW 113369/1 dry, coll. JG F1053/2 dry, coll.
|
|
<normalizedToken id="1B9F41720A478A2B3D54F153B37BAD80" originalValue="Glöer">Gloeer</normalizedToken>
|
|
/1 dry.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="75046CCED85087E27B0218AA0F736768" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="27E642EDF530F21E2A1F773086709AB0" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="5FC77AAB74BD7240A10EA88023FFE720" pageId="0" pageNumber="1">
|
|
The new species differs from its closest relatives by its oval shell shape, proportionally smaller aperture more close-set to the columella and closed umbilicus. There is some similarity to the shell shape of
|
|
<taxonomicName id="96EF244A78E6BF47D2C4D8183FAD5B6C" lsidName="C. subovata" pageId="0" pageNumber="1" rank="species" species="subovata">
|
|
<emphasis id="0D7A1C827D7214C607F655D2269C9D2A" italics="true" pageId="0" pageNumber="1">C. subovata</emphasis>
|
|
</taxonomicName>
|
|
(Starobogatov, 1962) from Abkhazia, however, the broken subfossil type does not allow more detailed comparison, and the drawing of the author within the description was likely just a reconstruction of the broken holotype.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="590AC726E3A75D0E630BDDBCCC61434E" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="5A2C69D50676C4994BC44F5128416E45" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="FCA994F77B449DA218460C3AB3B503CB" pageId="0" pageNumber="1">
|
|
<emphasis id="0953D3984EFADB9FD74A20F85D6484D8" bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
|
: is 1.50-1.96 mm high, elongate ovate-conical with rounded whorls and blunt apex. Surface smooth, whitish, occasionally with inorganic incrustations. Aperture proportionally small, flat-ovoid shaped, situated below larger body whorl. Lateral profile of labral margin straight, columellar margin very weakly sinuated. Umbilicus closed.
|
|
</paragraph>
|
|
<paragraph id="1A49E9AA5DAEDDF85DAB02DDDC25A6DA" pageId="0" pageNumber="1">
|
|
<emphasis id="45BB615C8A818720F4764A22F4E7A4BD" bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
<paragraph id="77F83C0842FE9F11DB13F58B1E6F24ED" pageId="0" pageNumber="1">
|
|
<emphasis id="6D12F736300384C541419F27813ADFB7" bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
<paragraph id="CCF2A2033997507D52673512BC397F3A" pageId="0" pageNumber="1">
|
|
<emphasis id="05CABF719DAFE121E5AB448968BC4969" bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
|
|
: H-1.87 mm; W-0.85 mm; BH-0.94 mm; BW-0.77 mm; AH-0.60 mm; AW-0.45 mm; CA: 26°.
|
|
</paragraph>
|
|
<paragraph id="7C95953AB8F77B700A83669CB12EB616" pageId="0" pageNumber="1">
|
|
<emphasis id="4A47F5B4BFD4DFA834396C540D7FB61C" bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
|
|
: not known.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B77B0D395A30F6E6D37E4EE65419FECE" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="D7C6ADC7AD8608AEE4363D8E4D77C28A" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="633C94E0376897899870700D6CEC8329" pageId="0" pageNumber="1">
|
|
Named for our friend
|
|
<normalizedToken id="8964AE5701A8A9DB8FE26A005E0FCA72" originalValue="Mário">Mario</normalizedToken>
|
|
<normalizedToken id="9753DA07A14BC711A83F0ADF8A8E9707" originalValue="Olšavský">Olsavsky</normalizedToken>
|
|
, geologist and speleologist from
|
|
<normalizedToken id="9CC30234F5D9620EB9F7D1F57FD85A63" originalValue="Banská">Banska</normalizedToken>
|
|
Bystrica, Slovakia, who actively participated in the field trip to Georgia.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C46C489E3431A344518397186FACDDD0" pageId="0" pageNumber="1" type="habitat">
|
|
<paragraph id="550DA6164DE5585920A7C5CA1A4A4600" pageId="0" pageNumber="1">Habitat.</paragraph>
|
|
<paragraph id="DEB5C0BAF38E7A68C160157017A3158A" pageId="0" pageNumber="1">
|
|
Stygobiotic species. Empty shells were found at the sandy bottom of the cave stream inside a conglomerate cave. The empty shells were very scarce, as an undetermined
|
|
<taxonomicName id="168AE3F985F60E3E83CDC5BAF21C2C87" authorityName="Vinarski & Palatov" authorityYear="2019" class="Gastropoda" family="Hydrobiidae" genus="Tschernomorica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tschernomorica" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="23E4476DEB19AB8AE75AA475DF964512" italics="true" pageId="0" pageNumber="1">Tschernomorica</emphasis>
|
|
</taxonomicName>
|
|
sp. was more abundant in the type locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="01445423DBFA37441E076B4F7CAC57C5" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="4D825B17D152F2B8E369A11548E31FE1" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="735FF22B5509B13B40F69BC06052FC29" pageId="0" pageNumber="1">Only known from the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7CE0CC4D40DA9DC3C32DECE6794BADB9" pageId="0" pageNumber="1" type="conservation status">
|
|
<paragraph id="3076DC6AA20B7424761F2910D028878C" pageId="0" pageNumber="1">Conservation status.</paragraph>
|
|
<paragraph id="D89D31149985A687FA62AF13AC13AD55" pageId="0" pageNumber="1">The number of known locations (1) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |