156 lines
16 KiB
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156 lines
16 KiB
XML
<document ID-DOI="10.18590/euscorpius.2018.vol2018.iss261.1" ID-GBIF-Dataset="46c20b18-c0d5-46f5-8894-fd8c5df4055e" ID-ISSN="1536-9307" ID-Zenodo-Dep="7118814" ID-ZooBank="BDB28370-E60B-49B0-A2DA-F30D6E89D6D0" approvalRequired="1" approvalRequired_for_treatments="1" checkinTime="1638258814613" checkinUser="felipe" docAuthor="Lowe, Graeme" docDate="2018" docId="1F0A87B4FFA56F10FEAD73F1FC0F2D6C" docLanguage="en" docName="Euscorpius.261.1-73.pdf" docOrigin="Euscorpius 261" docSource="https://doi.org/10.18590/euscorpius.2018.vol2018.iss261.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.2:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="2" docTitle="Xenobuthus Lowe 2018, gen. nov." docType="treatment" docUuid="CC2E0A92-2CDF-4D3B-A4A0-66974E8A567D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="38" masterDocId="E333FFCCFF836F37FF92766DFFE22A0E" masterDocTitle="The genera Butheolus Simon, 1882 and Xenobuthus gen. nov. (Scorpiones: Buthidae) in Oman" masterLastPageNumber="73" masterPageNumber="1" pageNumber="37" updateTime="1677625032903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genera Butheolus Simon, 1882 and Xenobuthus gen. nov. (Scorpiones: Buthidae) in Oman</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lowe, Graeme</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>261</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://doi.org/10.18590/euscorpius.2018.vol2018.iss261.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.18590/euscorpius.2018.vol2018.iss261.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">46c20b18-c0d5-46f5-8894-fd8c5df4055e</mods:identifier>
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<mods:identifier type="ISSN">1536-9307</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7118814</mods:identifier>
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<mods:identifier type="ZooBank">BDB28370-E60B-49B0-A2DA-F30D6E89D6D0</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7118844" ID-GBIF-Taxon="201133412" ID-Zenodo-Dep="7118844" LSID="urn:lsid:zoobank.org:act:CC2E0A92-2CDF-4D3B-A4A0-66974E8A567D" httpUri="http://treatment.plazi.org/id/1F0A87B4FFA56F10FEAD73F1FC0F2D6C" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
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<subSubSection box="[319,674,1435,1462]" pageId="38" pageNumber="37" type="nomenclature">
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<paragraph blockId="38.[204,789,1435,1517]" box="[319,674,1435,1462]" pageId="38" pageNumber="37">
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<heading box="[319,674,1435,1462]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="37" reason="2">
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Genus
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<taxonomicName ID-CoL="88NK" ID-ENA="2935629" authority="Lowe, 2018" authorityName="Lowe" authorityYear="2018" box="[401,548,1435,1462]" class="Arachnida" family="Buthidae" genus="Xenobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[401,548,1435,1462]" italics="true" pageId="38" pageNumber="37">Xenobuthus</emphasis>
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</taxonomicName>
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,
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<emphasis bold="true" box="[563,674,1436,1462]" pageId="38" pageNumber="37">
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<taxonomicNameLabel box="[563,674,1436,1462]" pageId="38" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="38" pageNumber="37" type="description">
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<paragraph blockId="38.[204,789,1435,1517]" pageId="38" pageNumber="37">
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<emphasis bold="true" box="[204,789,1469,1490]" pageId="38" pageNumber="37">http://zoobank.org/urn:lsid:zoobank.org:act:CC2E0A92-</emphasis>
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<emphasis bold="true" box="[317,677,1496,1517]" pageId="38" pageNumber="37">2CDF-4D3B-A4A0-66974E8A567D</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37" type="diagnosis">
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<paragraph blockId="38.[192,802,1557,1917]" lastBlockId="39.[830,1440,1598,1651]" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
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DIAGNOSIS. Small to medium-sized buthids (
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<bibRefCitation author="Kovarik" firstAuthor="Kovarik" pageId="38" pageNumber="37" refId="ref33839" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp." type="book" year="2009">Kovařík, 2009</bibRefCitation>
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;
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<bibRefCitation author="SISSOM, W. D." box="[262,409,1587,1610]" editor="Polis, G. A." journalOrPublisher="Stanford University Press, Stanford, California" pageId="38" pageNumber="37" pagination="64 - 160" refId="ref35859" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in Polis, G. A. (ed.). The Biology of Scorpions. Stanford University Press, Stanford, California." title="Systematics, biogeography and paleontology" type="book chapter" volumeTitle="The Biology of Scorpions" year="1990">Sissom, 1990</bibRefCitation>
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), adults
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<quantity box="[507,624,1587,1610]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="38" pageNumber="37" unit="mm" value="42.5" valueMax="50.0" valueMin="35.0">35–50 mm</quantity>
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in length; carapace strongly trapezoidal, ratio of posterior W/ anterior
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<geoCoordinate box="[192,260,1649,1672]" degrees="2.1" direction="west" orientation="longitude" pageId="38" pageNumber="37" precision="5555" value="-2.1">W 2.1</geoCoordinate>
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–2.4, preocular area inclined downwards towards anterior margin, surface granular, without distinct carinae; tergites granular, tergites I–III lacking carinae or with weak median carina, IV–VI weakly tricarinate with median and paired lateral carinae; tergite VII with broad median carina, two pairs of lateral carinae; posterior margins of sternites III–VI armed with fringe of noncontiguous digitate denticles that are larger in males; metasoma segments nearly uniform in width and depth, robust with smooth or granulated carinae, sparsely or moderately granular on lateral and ventral intercarinal surfaces; metasoma I–III with 10 carinae, IV with 6 or 8 carinae, V with 3 or 5 carinae; ventrolateral carinae on metasoma V without enlarged lobate dentition; telson with bulbous vesicle, without subaculear tubercle, aculeus shorter than vesicle; pectines with fulcra; hemispermatophore with flagellum separated from a 3-lobed sperm hemiduct, basal lobe a broad, low, axially oriented, curved scoop; chelicerae with characteristic buthid pattern of dentition (
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<bibRefCitation author="Vachon" box="[1125,1276,1586,1609]" firstAuthor="Vachon" pageId="38" pageNumber="37" pagination="161 - 166" refId="ref36237" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
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), two denticles on ventral aspect of fixed finger; pedipalps orthobothriotaxic,
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<typeStatus box="[974,1020,1649,1672]" pageId="38" pageNumber="37">type</typeStatus>
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Aβ (
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<bibRefCitation author="Vachon" box="[1078,1222,1649,1672]" firstAuthor="Vachon" pageId="38" pageNumber="37" pagination="857 - 958" refId="ref36318" refString="VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
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,
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<bibRefCitation author="Vachon" box="[1237,1292,1649,1672]" firstAuthor="Vachon" pageId="38" pageNumber="37" pagination="1597 - 1599" refId="ref36380" refString="VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser. D Sciences Naturelles, 281 (21): 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
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), patella with
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<emphasis box="[831,843,1680,1703]" italics="true" pageId="38" pageNumber="37">d</emphasis>
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<subScript attach="left" box="[844,853,1690,1705]" fontSize="6" pageId="38" pageNumber="37">3</subScript>
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internal to dorsomedian carina, manus with
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<emphasis box="[1369,1385,1680,1703]" italics="true" pageId="38" pageNumber="37">V</emphasis>
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<subScript attach="left" box="[1385,1394,1690,1705]" fontSize="6" pageId="38" pageNumber="37">2</subScript>
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not strongly displaced internally,
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<emphasis box="[1152,1167,1710,1733]" italics="true" pageId="38" pageNumber="37">V</emphasis>
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<subScript attach="left" box="[1167,1176,1721,1736]" fontSize="6" pageId="38" pageNumber="37">1</subScript>
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–
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<emphasis box="[1189,1205,1710,1733]" italics="true" pageId="38" pageNumber="37">V</emphasis>
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<subScript attach="left" box="[1205,1214,1721,1736]" fontSize="6" pageId="38" pageNumber="37">2</subScript>
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axis nearly collinear with long axis of chela, chela fixed finger with
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<emphasis box="[1376,1403,1741,1764]" italics="true" pageId="38" pageNumber="37">db</emphasis>
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on proximal half of finger between
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<emphasis box="[1188,1223,1772,1795]" italics="true" pageId="38" pageNumber="37">esb</emphasis>
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and
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<emphasis box="[1280,1309,1772,1795]" italics="true" pageId="38" pageNumber="37">est</emphasis>
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,
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<emphasis box="[1326,1341,1772,1795]" italics="true" pageId="38" pageNumber="37">it</emphasis>
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near tip; pedipalps short, chelae small with carinae reduced or obsolete, dentate margins of fingers armed with 8–10 non-imbricated linear subrows of primary denticles; subrows flanked by mid-row internal and proximal external accessory denticles; movable finger with 3 enlarged subdistal denticles; males without recess or scalloping of dentate margins at base of pedipalp fingers, with chela manus wider than females; tergites lacking macrosetae; tibial spurs present on legs III–IV; basitarsi I–III with regular series of long macrosetae on retrosuperior, retroinferior and inferior margins; ventral surfaces of telotarsi with paired rows of macrosetae, often reduced to single row on legs I–III; prolateral and retrolateral tarsal spurs present on all legs; size of macrosetae on carapace, pedipalps, legs and metasoma not sexually dimorphic.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7118890" ID-Zenodo-Dep="7118890" httpUri="https://zenodo.org/record/7118890/files/figure.png" pageId="39" pageNumber="38" startId="39.[192,278,1433,1456]" targetBox="[204,1425,223,1377]" targetPageId="39">
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<paragraph blockId="39.[192,1441,1433,1555]" pageId="39" pageNumber="38">
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<emphasis bold="true" box="[192,392,1433,1456]" pageId="39" pageNumber="38">Figures 18 3–193:</emphasis>
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Metasoma, telson, pedipalp, trichobothrial map and leg tarsi of
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<taxonomicName authority="Hendrixson, 2006" authorityName="Hendrixson" authorityYear="2006" box="[1064,1437,1434,1455]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="villosus">
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<emphasis box="[1064,1244,1434,1455]" italics="true" pageId="39" pageNumber="38">Butheolus villosus</emphasis>
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<bibRefCitation author="Hendrixson" box="[1256,1437,1434,1455]" firstAuthor="Hendrixson" pageId="39" pageNumber="38" pagination="33 - 120" refId="ref33409" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." type="journal article" year="2006">Hendrixson, 2006</bibRefCitation>
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</taxonomicName>
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. Holotype female. UV fluorescence.
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<emphasis bold="true" box="[540,684,1459,1480]" pageId="39" pageNumber="38">Figs. 183–184.</emphasis>
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Metasoma and telson. Right lateral (183) and ventral (184) aspects. Scale bar: 2 mm.
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<emphasis bold="true" box="[262,408,1484,1505]" pageId="39" pageNumber="38">Figs. 185–189.</emphasis>
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Pedipalp and trichobothria. Dorsal (185) and external (186) aspect of chela; dorsal (187) aspect of femur; external (188) and dorsal (189) aspect of patella. Scale bar: 1 mm.
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<emphasis bold="true" box="[846,991,1509,1530]" pageId="39" pageNumber="38">Figs. 190–193.</emphasis>
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Tarsi and tibia of left leg I (190), II (191), III (192) and IV (193), retrolateral aspect. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<subSubSection pageId="39" pageNumber="38" type="etymology">
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<paragraph blockId="39.[830,1440,1686,1770]" pageId="39" pageNumber="38">
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ETYMOLOGY. Prefix from Greek ξένος (xénos) meaning alien or strange, i.e. a strange buthid that resembles
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<taxonomicName authorityName=": Karsch" authorityYear="1886" box="[830,937,1747,1770]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis box="[830,937,1747,1770]" italics="true" pageId="39" pageNumber="38">Butheolus</emphasis>
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</taxonomicName>
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in some features but differs from it in others.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="38" type="type_taxon">
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<paragraph blockId="39.[830,1440,1806,1890]" pageId="39" pageNumber="38">
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<collectionCode box="[830,1023,1806,1829]" pageId="39" pageNumber="38">
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<typeStatus box="[830,897,1806,1829]" pageId="39" pageNumber="38">TYPE</typeStatus>
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SPECIES
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</collectionCode>
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.
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<taxonomicName authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" box="[1045,1440,1806,1829]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="anthracinus">
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<emphasis box="[1045,1264,1806,1829]" italics="true" pageId="39" pageNumber="38">Buthus anthracinus</emphasis>
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<bibRefCitation author="Pocock" box="[1283,1440,1806,1829]" firstAuthor="Pocock" pageId="39" pageNumber="38" pagination="292 - 316" refId="ref35417" refString="POCOCK, R. I. 1895. On the Arachnida and Myriopoda obtained by Dr. Anderson's collector during Mr T. Brent's expedition to the Hadramaut, South Arabia, with a supplement upon the scorpions obtained by Dr. Anderson in Egypt and the Eastern Soudan. Journal of the Linnean Society, 25: 292 - 316." type="journal article" year="1895">Pocock, 1895</bibRefCitation>
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</taxonomicName>
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[=
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<taxonomicName authority="(Pocock, 1895)" authorityName="Lowe" authorityYear="2018" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[854,1304,1836,1859]" class="Arachnida" family="Buthidae" genus="Xenobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="anthracinus" status="comb. n.">
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<emphasis box="[854,1120,1836,1859]" italics="true" pageId="39" pageNumber="38">Xenobuthus anthracinus</emphasis>
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(
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<bibRefCitation author="Pocock" box="[1140,1297,1836,1859]" firstAuthor="Pocock" pageId="39" pageNumber="38" pagination="292 - 316" refId="ref35417" refString="POCOCK, R. I. 1895. On the Arachnida and Myriopoda obtained by Dr. Anderson's collector during Mr T. Brent's expedition to the Hadramaut, South Arabia, with a supplement upon the scorpions obtained by Dr. Anderson in Egypt and the Eastern Soudan. Journal of the Linnean Society, 25: 292 - 316." type="journal article" year="1895">Pocock, 1895</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis bold="true" box="[1321,1434,1836,1859]" pageId="39" pageNumber="38">
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<taxonomicNameLabel box="[1321,1426,1836,1859]" pageId="39" pageNumber="38" rank="species">comb. n.</taxonomicNameLabel>
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]
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</emphasis>
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, designated here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |