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<document id="C0BB97345A50C1CE516AE0F67EE656EF" ID-CLB-Dataset="298742" ID-DOI="10.11646/zootaxa.5471.4.4" ID-GBIF-Dataset="cf49a9e9-837f-4e80-b5a1-1ac3ee1e6170" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190387" ID-ZooBank="7486CBBE-0329-40C2-94AC-680D8703F56F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718894049516" checkinUser="plazi" docAuthor="Kim, Dooyoung &amp; Suh, Sang Jae" docDate="2024" docId="135E87AA5218FFEC2997FDA6FCC04CBB" docLanguage="en" docName="zootaxa.5471.4.4.pdf" docOrigin="Zootaxa 5471 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer 1996" docType="treatment" docVersion="6" lastPageNumber="459" masterDocId="EF67FFD2521FFFE42900FFC0FFEE4C42" masterDocTitle="Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)" masterLastPageNumber="464" masterPageNumber="451" pageNumber="458" updateTime="1719009245391" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FC39E4B066333E24D0472149EAE28756">Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)</mods:title>
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<mods:namePart id="D6E4BD7D4BA0D35203A550B01EB40448">Kim, Dooyoung</mods:namePart>
<mods:affiliation id="585506CDFF2A226410335232E5CF940C">Department of Biomedical Convergence Science and Technology, School of Industrial Technology Advances, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<mods:namePart id="8B6285241689C1E9E5F1B53D5D2027A9">Suh, Sang Jae</mods:namePart>
<mods:affiliation id="33587A96AFA8175243DC4D0436B8E83F">Department of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea &amp; Institute of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<subSubSection id="D3ED65375218FFE32997FDA6FDF24EE4" pageId="7" pageNumber="458" type="nomenclature">
<paragraph id="9B4836BC5218FFE32997FDA6FC274EC2" blockId="7.[151,969,614,713]" box="[151,969,614,641]" pageId="7" pageNumber="458">
<heading id="C00081D05218FFE32997FDA6FC274EC2" bold="true" box="[151,969,614,641]" fontSize="11" level="1" pageId="7" pageNumber="458" reason="1">
<taxonomicName id="5CF74D3F5218FFE32997FDA6FC274EC2" ID-CoL="4Q3X4" authority="Park, Kwon &amp; Lafer, 1996" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[151,969,614,641]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32997FDA6FC274EC2" bold="true" box="[151,969,614,641]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE32997FDA6FEC34EC3" bold="true" box="[151,301,614,641]" italics="true" pageId="7" pageNumber="458">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5218FFE3283EFDA6FE3C4EC3" bold="true" box="[318,466,614,641]" italics="true" pageId="7" pageNumber="458">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5218FFE328E4FDA6FD994EC3" bold="true" box="[484,631,614,641]" italics="true" pageId="7" pageNumber="458">bifoveolatus</emphasis>
<bibRefCitation id="FF664B4D5218FFE32B7EFDA7FC274EC2" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[638,969,614,641]" pageId="7" pageNumber="458" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="9B4836BC5218FFE32997FD4BFDF24EE4" blockId="7.[151,969,614,713]" box="[151,540,651,678]" pageId="7" pageNumber="458">Korean name: Ī마길쭉먼지벌레</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE32997FD6FFEE64E8B" box="[151,264,687,713]" pageId="7" pageNumber="458" type="description">
<paragraph id="9B4836BC5218FFE32997FD6FFEE64E8B" blockId="7.[151,969,614,713]" box="[151,264,687,713]" pageId="7" pageNumber="458">
(
<figureCitation id="03CC2A395218FFE3299FFD6FFEEE4E8B" box="[159,256,687,713]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figure 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE32997FD33FB7C4F3A" pageId="7" pageNumber="458" type="reference_group">
<paragraph id="9B4836BC5218FFE32997FD33FC834F49" blockId="7.[151,877,755,779]" box="[151,877,755,779]" pageId="7" pageNumber="458">
<treatmentCitationGroup id="BBE711925218FFE32997FD33FC834F49" box="[151,877,755,779]" pageId="7" pageNumber="458">
<treatmentCitation id="1A5610AD5218FFE32997FD33FC874F49" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[151,873,755,779]" page="74" pageId="7" pageNumber="458" year="1996">
<taxonomicName id="5CF74D3F5218FFE32997FD33FC874F49" ID-CoL="4Q3X4" authority="Park, Kwon &amp; Lafer, 1996: 74" authorityName="Park, Kwon &amp; Lafer" authorityPageNumber="74" authorityYear="1996" box="[151,873,755,779]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32997FD33FEFB4F49" box="[151,277,755,779]" italics="true" pageId="7" pageNumber="458">Pterostichus</emphasis>
(
<emphasis id="A983EAAE5218FFE32824FD33FE4E4F49" box="[292,416,755,779]" italics="true" pageId="7" pageNumber="458">Micronialoe</emphasis>
)
<emphasis id="A983EAAE5218FFE328AFFD33FDC54F49" box="[431,555,755,779]" italics="true" pageId="7" pageNumber="458">bifoveolatus</emphasis>
<bibRefCitation id="FF664B4D5218FFE32B31FD33FC874F49" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[561,873,755,779]" pageId="7" pageNumber="458" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996: 74</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="9B4836BC5218FFE32997FCF9FB7C4F3A" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE32997FCF9FE6F4F11" bold="true" box="[151,385,825,851]" pageId="7" pageNumber="458">Specimen examined</emphasis>
.
<materialsCitation id="2B9F3CE15218FFE3288CFCFAFB654F3A" ID-GBIF-Occurrence="4898139313" collectingDate="2022-08-05" collectorName="Dooyoung Kim" country="South Korea" county="Sancheong-gun" elevation="530" latitude="35.37964" location="Honggye-ri" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" pageId="7" pageNumber="458" specimenCount="1" specimenCount-male="1" stateProvince="Gyeongnam">
<collectingCountry id="E3E0762C5218FFE3288CFCFAFE3E4F16" box="[396,464,826,852]" name="South Korea" pageId="7" pageNumber="458">Korea</collectingCountry>
<specimenCount id="8DF1FD355218FFE328E2FCF9FDE94F11" box="[482,519,825,851]" count="1" pageId="7" pageNumber="458" type="male">1♂</specimenCount>
;
<collectingRegion id="5933F85E5218FFE32B11FCF9FD1B4F11" box="[529,757,825,851]" country="South Korea" name="Gyeongnam" pageId="7" pageNumber="458">Gyeongsangnam-do</collectingRegion>
,
<collectingCounty id="72294E305218FFE32BFEFCF9FC414F11" box="[766,943,825,851]" pageId="7" pageNumber="458">Sancheong-gun</collectingCounty>
,
<collectingMunicipality id="7B2CACC65218FFE32AB7FCF9FB9F4F11" box="[951,1137,825,851]" pageId="7" pageNumber="458">Samjang-myeon</collectingMunicipality>
,
<location id="9E2860675218FFE32D79FCF9FB134F11" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675218FFE32D79FCF9FB134F11" box="[1145,1277,825,851]" country="South Korea" county="Sancheong-gun" latitude="35.37964" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" name="Honggye-ri" pageId="7" pageNumber="458" stateProvince="Gyeongnam">Honggye-ri</location>
,
<location id="9E2860675218FFE32C05FCF9FA724F11" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675218FFE32C05FCF9FA724F11" box="[1285,1436,825,851]" country="South Korea" county="Sancheong-gun" latitude="35.37964" longLatPrecision="1" longitude="127.853806" municipality="Samjang-myeon" name="Bammeorijae" pageId="7" pageNumber="458" stateProvince="Gyeongnam">Bammeorijae</location>
pass;
<geoCoordinate id="FEC3507B5218FFE329D6FC9DFE814F3A" box="[214,367,861,888]" degrees="35" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="458" precision="1" seconds="46.7" value="35.37964">35°22'46.7&quot;N</geoCoordinate>
<geoCoordinate id="FEC3507B5218FFE32876FC9DFDF34F3A" box="[374,541,861,888]" degrees="127" direction="east" minutes="51" orientation="longitude" pageId="7" pageNumber="458" precision="1" seconds="13.7" value="127.853806">127°51'13.7&quot;E</geoCoordinate>
; alt. ca
<quantity id="5C0F9B595218FFE32B72FC9DFD524F3A" box="[626,700,861,888]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="7" pageNumber="458" unit="m" value="530.0">
<elevation id="10DAD18F5218FFE32B72FC9DFD524F3A" box="[626,700,861,888]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="7" pageNumber="458" unit="m" value="530.0">530 m</elevation>
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;
<date id="EF49107C5218FFE32BC8FC9DFCB84F35" box="[712,854,861,888]" pageId="7" pageNumber="458" value="2022-08-05">
<collectingDate id="FF0DE9945218FFE32BC8FC9DFCB84F35" box="[712,854,861,888]" pageId="7" pageNumber="458" value="2022-08-05">5 Aug. 2022</collectingDate>
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;
<collectorName id="3602536A5218FFE32A60FC9EFBFE4F35" box="[864,1040,861,888]" pageId="7" pageNumber="458">Dooyoung Kim</collectorName>
leg.; KNU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFE329C7FC41FD1D4941" pageId="7" pageNumber="458" type="diagnosis">
<paragraph id="9B4836BC5218FFE329C7FC41FABB4FA1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FC41FEAC4FD9" bold="true" box="[199,322,897,923]" pageId="7" pageNumber="458">Diagnosis.</emphasis>
This species is easily distinguished from known species of the subgenus by: i) posteromedian area of male sternite VII not rugose and asymmetrically depressed, ii) median lobe of aedeagus with rounded apex in dorsal view, and iii) internal sac of aedeagus with two sclerotized areas. Other significant characters are as follows.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FC2EFD9748DA" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">Pronotum slightly sinuate on lateral margin before basal angle; with two pairs of basal foveae. Posteromedian area of male sternite VII with microsculpture consisting of irregular transverse meshes. Median lobe of aedeagus stout, left lateral margin below apical orifice distinctly convex; ventral carina well developed, widely rounded; sclerotized areas in internal sac consist of followings: i) C-shaped sclerite (CS) on median area and ii) wide triangular shaped ventral sclerite (VS) on apical area.</paragraph>
<paragraph id="9B4836BC5218FFE329C7FB61FD1D4941" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
This species may be closely allied to
<taxonomicName id="5CF74D3F5218FFE32B72FB62FC5E48FE" authority="Morita, 2005" authorityName="Morita" authorityYear="2005" box="[626,944,1185,1212]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE5218FFE32B72FB62FD6D48F9" box="[626,643,1186,1211]" italics="true" pageId="7" pageNumber="458">P</emphasis>
. (
<emphasis id="A983EAAE5218FFE32B9CFB62FD5D48F9" box="[668,691,1186,1211]" italics="true" pageId="7" pageNumber="458">M</emphasis>
.)
<emphasis id="A983EAAE5218FFE32BCCFB61FCFE48F9" box="[716,784,1185,1211]" italics="true" pageId="7" pageNumber="458">kaniei</emphasis>
<bibRefCitation id="FF664B4D5218FFE32A19FB61FC5E48FE" author="Morita, S." box="[793,944,1185,1212]" pageId="7" pageNumber="458" pagination="9 - 12" refId="ref7740" refString="Morita, S. (2005) A New Pterostichus (Coleoptera: Carabidae) from the Islands of Tsushima, West Japan. Entomological Review of Japan, 60 (1), 9 - 12." type="journal article" year="2005">Morita, 2005</bibRefCitation>
</taxonomicName>
, based on their common genital characters: i) median lobe of aedeagus stout, ii) apex moderately rounded in dorsal view, iii) left lateral margin below apical orifice convex, and iv) ventral carina widely rounded.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375218FFEC29C7FACDFB014C97" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458" type="description">
<paragraph id="9B4836BC5218FFE329C7FACDFC6B496A" blockId="7.[151,1437,825,2040]" box="[199,901,1293,1320]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FACDFE9B4965" bold="true" box="[199,373,1293,1319]" pageId="7" pageNumber="458">Redescription.</emphasis>
Body length:
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, width:
<quantity id="5C0F9B595218FFE32BF7FACDFC6F496A" box="[759,897,1293,1320]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="7" pageNumber="458" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.83.4 mm</quantity>
.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FAF1FF5449D1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
Coloration reddish black and shiny, elytra slightly iridescent; antennae, legs, and lateral margin of both pronotum and elytra reddish brown; palpi yellowish brown, with apices distinctly paler; ventral side reddish brown (
<figureCitation id="03CC2A395218FFE32C3AFA95FF4249D1" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7A, B</figureCitation>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FA5DFC43499E" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FA5DFEEA49F5" box="[199,260,1437,1463]" italics="true" pageId="7" pageNumber="458">Head</emphasis>
(
<figureCitation id="03CC2A395218FFE32818FA5DFE9B49F5" box="[280,373,1437,1463]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7C</figureCitation>
). Eye weakly convex; tempus well developed, moderately convex, as long as eye; frontal impression deep, moderately curved toward anterior supraorbital pore.
</paragraph>
<paragraph id="9B4836BC5218FFE329C7FA26FCB54AF1" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7FA26FED849BD" box="[199,310,1510,1535]" italics="true" pageId="7" pageNumber="458">Pronotum</emphasis>
(
<figureCitation id="03CC2A395218FFE32848FA25FE6149BD" box="[328,399,1509,1535]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7</figureCitation>
) widest near 2/5 from apex; apical angle narrowly rounded, slightly protruded anteriorly; lateral margin slightly sinuate before basal angle; basal angle obtuse; with two pairs of basal foveae; inner foveae deep and distinct, linear, almost parallel, with distinct punctures; outer foveae almost linear, slightly tilted outward; basal surface almost smooth medially; area between inner fovea and outer fovea with or without distinct punctures.
<materialsCitation id="2B9F3CE15218FFE32997F9B5FDF64ACD" ID-GBIF-Occurrence="4898139302" box="[151,536,1653,1680]" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32997F9B5FF2F4ACD" box="[151,193,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/HW = 1.61 (
<quantity id="5C0F9B595218FFE32854F9B5FE4D4ACD" box="[340,419,1653,1680]" metricMagnitude="-2" metricUnit="m" metricValue="4.0386" pageId="7" pageNumber="458" unit="in" value="1.59">1.59 in</quantity>
<typeStatus id="444C881E5218FFE328A9F9B5FDE14ACD" box="[425,527,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
,
<materialsCitation id="2B9F3CE15218FFE32B22F9B5FC7A4ACD" ID-GBIF-Occurrence="4898139312" box="[546,916,1653,1680]" collectionCode="PW, PL" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32B22F9B5FDA54ACD" box="[546,587,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/
<collectionCode id="FDE6AE795218FFE32B54F9B5FD9A4ACD" box="[596,628,1653,1679]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="458" type="Museum">PL</collectionCode>
= 1.11 (
<quantity id="5C0F9B595218FFE32BD0F9B5FCF14ACD" box="[720,799,1653,1679]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="458" unit="in" value="1.2">1.20 in</quantity>
<typeStatus id="444C881E5218FFE32A25F9B5FC624ACD" box="[805,908,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32A9FF9B5FAC44ACD" ID-GBIF-Occurrence="4898139307" box="[927,1322,1653,1680]" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32A9FF9B5FC274ACD" box="[927,969,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/PbW = 1.19 (
<quantity id="5C0F9B595218FFE32D65F9B5FB5A4ACD" box="[1125,1204,1653,1679]" metricMagnitude="-2" metricUnit="m" metricValue="2.9463999999999997" pageId="7" pageNumber="458" unit="in" value="1.16">1.16 in</quantity>
<typeStatus id="444C881E5218FFE32DBBF9B5FACC4ACD" box="[1211,1314,1653,1679]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32C34F9B5FE5D4AF1" ID-GBIF-Occurrence="4898139315" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
<collectionCode id="FDE6AE795218FFE32C34F9B5FAB04ACD" box="[1332,1374,1653,1679]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
/PaW = 1.46 (
<quantity id="5C0F9B595218FFE329EEF959FED34AF1" box="[238,317,1689,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.5814" pageId="7" pageNumber="458" unit="in" value="1.41">1.41 in</quantity>
<typeStatus id="444C881E5218FFE32844F959FE454AF1" box="[324,427,1689,1715]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
; PbW/PaW = 1.23 (
<quantity id="5C0F9B595218FFE32B92F959FD0F4AF1" box="[658,737,1689,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="7" pageNumber="458" unit="in" value="1.21">1.21 in</quantity>
<typeStatus id="444C881E5218FFE32BE8F959FCA14AF1" box="[744,847,1689,1715]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7F97DFA8E4B5D" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F97DFEE34A95" box="[199,269,1725,1751]" italics="true" pageId="7" pageNumber="458">Elytra</emphasis>
<materialsCitation id="2B9F3CE15218FFE32816F97DFD8C4B5D" ID-GBIF-Occurrence="4898139306" collectionCode="PW" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
widest before 2/5 from base; striae deep, punctate from base to basal half; marginal umbilicate series composed of three groups, humeral group with 5 pores, medial group with 1 pore, and apical group with 7 or 8 pores. EW/
<collectionCode id="FDE6AE795218FFE32815F8C5FED14B5D" box="[277,319,1797,1823]" country="Thailand" name="Paleontological Collections" pageId="7" pageNumber="458">PW</collectionCode>
= 1.19 (
<quantity id="5C0F9B595218FFE3289CF8C5FE024B5D" box="[412,492,1797,1823]" metricMagnitude="-2" metricUnit="m" metricValue="3.0225999999999997" pageId="7" pageNumber="458" unit="in" value="1.19">1.19 in</quantity>
<typeStatus id="444C881E5218FFE328F3F8C5FDB44B5D" box="[499,602,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2B9F3CE15218FFE32B6DF8C5FC374B5D" ID-GBIF-Occurrence="4898139316" box="[621,985,1797,1824]" collectionCode="PL" pageId="7" pageNumber="458" specimenCount="1" typeStatus="holotype">
EL/
<collectionCode id="FDE6AE795218FFE32B96F8C5FD584B5D" box="[662,694,1797,1823]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="458" type="Museum">PL</collectionCode>
= 2.02 (
<quantity id="5C0F9B595218FFE32A14F8C5FC8D4B5D" box="[788,867,1797,1823]" metricMagnitude="-2" metricUnit="m" metricValue="5.740399999999999" pageId="7" pageNumber="458" unit="in" value="2.26">2.26 in</quantity>
<typeStatus id="444C881E5218FFE32A6AF8C5FC3F4B5D" box="[874,977,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
)
</materialsCitation>
; EL/EW = 1.55 (
<quantity id="5C0F9B595218FFE32D97F8C5FB084B5D" box="[1175,1254,1797,1824]" metricMagnitude="-2" metricUnit="m" metricValue="4.0132" pageId="7" pageNumber="458" unit="in" value="1.58">1.58 in</quantity>
<typeStatus id="444C881E5218FFE32DEDF8C5FABA4B5D" box="[1261,1364,1797,1823]" pageId="7" pageNumber="458" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="9B4836BC5218FFE329C7F8E9FC464B25" blockId="7.[151,1437,825,2040]" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F8E9FEA04B01" box="[199,334,1833,1859]" italics="true" pageId="7" pageNumber="458">Ventral side</emphasis>
(
<figureCitation id="03CC2A395218FFE3285EF8E9FE014B01" box="[350,495,1833,1860]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7B, 7D</figureCitation>
). Posteromedian area of male sternite VII (
<figureCitation id="03CC2A395218FFE32ADBF8E9FBDB4B01" box="[987,1077,1833,1859]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Fig. 7D</figureCitation>
) asymmetrically depressed, not rugose, with microsculpture consisting of irregular transverse meshes.
</paragraph>
<paragraph id="9B4836BC5218FFEC29C7F8B1FB014C97" blockId="7.[151,1437,825,2040]" lastBlockId="8.[151,1437,151,249]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<emphasis id="A983EAAE5218FFE329C7F8B1FE844BC9" box="[199,362,1905,1931]" italics="true" pageId="7" pageNumber="458">Male genitalia</emphasis>
(
<figureCitation id="03CC2A395218FFE32878F8B1FE1B4BC9" box="[376,501,1905,1931]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7EK</figureCitation>
). Median lobe of aedeagus (
<figureCitation id="03CC2A395218FFE32A29F8B1FC764BC9" box="[809,920,1905,1931]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="7" pageNumber="458">Figs 7EI</figureCitation>
) stout, distinctly curved in lateral view, almost linear in dorsal view; left lateral margin below apical orifice distinctly convex; apical orifice bent to left; terminal lamella very short; apex moderately rounded in dorsal view; ventral carina well developed, widely rounded, clearly visible in right dorsolateral view; internal sac with two sclerotized areas: i) C-shaped sclerite (CS) on median area and ii) wide triangular shaped ventral sclerite (VS) on apical area; left paramere (
<figureCitation id="03CC2A395217FFEC2D1CFF57FB844CF3" box="[1052,1130,151,177]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="8" pageNumber="459">Fig. 7J</figureCitation>
) wide; right paramere (
<figureCitation id="03CC2A395217FFEC2C6FFF57FF554C97" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm AC; 0.5 mm D; 0.3 mm EI; 0.2 mm JK. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="8" pageNumber="459">Fig. 7K</figureCitation>
) slender, distinctly curved, ventral margin forming obtuse angle, dorsal margin widely rounded.
</paragraph>
</subSubSection>
<subSubSection id="D3ED65375217FFEC29C7FF1FFCC04CBB" box="[199,814,222,249]" pageId="8" pageNumber="459" type="materials_examined">
<paragraph id="9B4836BC5217FFEC29C7FF1FFCC04CBB" blockId="8.[151,1437,151,249]" box="[199,814,222,249]" pageId="8" pageNumber="459">
<materialsCitation id="2B9F3CE15217FFEC29C7FF1FFCC44CBB" ID-GBIF-Occurrence="4898139304" box="[199,810,222,249]" country="South Korea" location="Mt. Jirisan" pageId="8" pageNumber="459" specimenCount="1" stateProvince="Gyeongnam" typeStatus="holotype">
<emphasis id="A983EAAE5217FFEC29C7FF1FFE884CBA" bold="true" box="[199,358,222,249]" pageId="8" pageNumber="459">
<typeStatus id="444C881E5217FFEC29C7FF1FFEEC4CBB" box="[199,258,223,249]" pageId="8" pageNumber="459">Type</typeStatus>
locality.
</emphasis>
<location id="9E2860675217FFEC286CFF1FFE094CBB" LSID="urn:lsid:plazi:treatment:135E87AA5218FFEC2997FDA6FCC04CBB:9E2860675217FFEC286CFF1FFE094CBB" box="[364,487,223,249]" country="South Korea" name="Mt. Jirisan" pageId="8" pageNumber="459" stateProvince="Gyeongnam">Mt. Jirisan</location>
,
<collectingRegion id="5933F85E5217FFEC28F2FF1FFD384CBB" box="[498,726,223,249]" country="South Korea" name="Gyeongnam" pageId="8" pageNumber="459">Gyeongsangnam-do</collectingRegion>
,
<collectingCountry id="E3E0762C5217FFEC2BE2FF1FFCC44CBB" box="[738,810,223,249]" name="South Korea" pageId="8" pageNumber="459">Korea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>