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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.363.6482" ID-GBIF-Dataset="36e5551d-3881-45a2-810a-386e517c86fc" ID-PMC="PMC3867181" ID-Pensoft-Pub="1313-2970-363-1" ID-PubMed="24363601" ID-ZBK="780695A9CB2E4FECA0560BA6F456185E" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-363-1" ModsDocOrigin="ZooKeys 363" ModsDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" checkinTime="1451246560986" checkinUser="pensoft" docAuthor="Mathis, Wayne N. &amp; Zatwarnicki, Tadeusz" docDate="2013" docId="0EC692C0444A33122B63207A12C9E9D1" docLanguage="en" docName="ZooKeys 363: 1-161" docOrigin="ZooKeys 363" docSource="http://dx.doi.org/10.3897/zookeys.363.6482" docTitle="Hydrochasma avanae Mathis &amp; Zatwarnicki" docType="treatment" docVersion="3" lastPageNumber="82" masterDocId="FFF2FFCA0333765855689824FFCDB839" masterDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="79" updateTime="1668157177207" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mathis, Wayne N.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>363</mods:number>
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<mods:start>1</mods:start>
<mods:end>161</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.363.6482</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.363.6482</mods:identifier>
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<treatment ID-GBIF-Taxon="152050169" LSID="urn:lsid:plazi:treatment:0EC692C0444A33122B63207A12C9E9D1" httpUri="http://treatment.plazi.org/id/0EC692C0444A33122B63207A12C9E9D1" lastPageId="81" lastPageNumber="82" pageId="78" pageNumber="79">
<subSubSection pageId="78" pageNumber="79" type="nomenclature">
<paragraph pageId="78" pageNumber="79">
<pageBreakToken pageId="78" pageNumber="79" start="start">27</pageBreakToken>
.
<taxonomicName LSID="http://species-id.net/wiki/Hydrochasma_avanae" authority="Mathis &amp; Zatwarnicki" class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae Mathis &amp; Zatwarnicki</taxonomicName>
Figs 149-153
</paragraph>
</subSubSection>
<subSubSection pageId="78" pageNumber="79" type="reference_group">
<paragraph pageId="78" pageNumber="79">
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
<bibRefCitation author="Mathis, WN" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="112" pageNumber="113" pagination="97 - 128" title="New species and other taxonomic modifications for shore flies from the Delmarva States (Diptera: Ephydridae)." url="10.4289/0013-8797-112.1.97" volume="112" year="2010">Mathis and Zatwarnicki 2010</bibRefCitation>
: 113 [United States. Utah. Grand: Swasey Beach (15.3 km N Green River;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°07'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°06.6'W</geoCoordinate>
; Green River; 1255 m); HT ♂, USNM].
</paragraph>
</subSubSection>
<subSubSection pageId="78" pageNumber="79" type="diagnosis">
<paragraph pageId="78" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="78" pageNumber="79">
This species is distinguished from congeners by the following combination of characters: Small shore flies, body length 1.30-1.85 mm. Head: Antenna mostly dark gray; parafacial silvery white, concolorous with facial coloration. Gena moderately high, height usually slightly higher than length of basal flagellomere; gena-to-eye ratio 0.71-0.73. Thorax: Wing hyaline; costal vein ratio 0.66-0.82; M vein ratio 0.45-0.50. Hindtibia bearing an apical black seta ventrally. Abdomen: Tergites lacking wedge-like marking laterally but tergites 2-4 with wide medial area extensively dark slate gray to black; tergite 5 light gray to silvery gray with undifferentiated posterior margin, uniformly colored; medial coloration on tergites 1-4 narrow, sometimes only a stripe, slightly darker than color of lateral margins. Male terminalia (Figs 149-152): Combined structures generally elongate, in posterior view (Fig. 149) height about 2.5
<normalizedToken originalValue="×">x</normalizedToken>
width; epandrium with dorsal arch above cerci attenuate, not connected, dorsal 1/3-1/2 somewhat quadrate with angles rounded, ventral portion as 2 digitiform, thick, parallel lobes that connect subapically for a short distance, ventral margin moderately narrowly rounded with deep, very narrow, apical incision medially, dorsal half bearing setulae, in lateral view (Fig. 150) mostly parallel sided with anterior shallowly sinuous, apical 1/3 tapered to sharply rounded apex; cerci short, in lateral view height about twice width (Fig. 150), narrowly semicircular, not attached with epandrium; aedeagus in lateral view (Fig. 152) elongate, with length of sclerotized portion about 5
<normalizedToken originalValue="×">x</normalizedToken>
width, with greatly enlarged apical and ventral, membranous portion, shape in lateral view (Fig. 152) triangular, with apex broad and shallowly arched, thereafter toward base tapered to narrow, bluntly rounded margin, membrane bearing sub-basally numerous scale-like spicules, in ventral view (Fig. 151) also showing expanded apex, basal 1/3 narrowed, thereafter with middle portion ballooning out to more than twice basal width, apical 1/3 greatly expanded, broadly obovate; phallapodeme in lateral view (Fig. 152) narrow, elongate, evenly shallowly arched, keel weakly developed, barely evident at end toward attachment with hypandrium, in ventral view (Fig. 151) with hypandrial end narrowly T-shaped, thereafter toward base of aedeagus gradually expanded to blunt apex; gonite in lateral view (Fig. 152) narrow, elongate, bar-like with slightly curvature, in ventral view (Fig. 151) moderately broad but with apices narrowed, especially end toward hypandrium; hypandrium in lateral view (Fig. 152) elongate, moderately shallow, width slightly more than 1/4 length, very slightly angulate, in ventral view (Fig. 151) generally V-shaped with posterior margin deeply U-shaped, anterior margin moderately narrowly rounded.
</paragraph>
<caption pageId="78" pageNumber="79">
<paragraph pageId="78" pageNumber="79">
Figures 149-152.
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
Mathis and Zatwarnicki (USA. Virginia. Fairfax: Turkey Run) 149 epandrium and cerci, posterior view 150 same, lateral view 151 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 152 same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="type material">
<paragraph pageId="79" pageNumber="80">
<pageBreakToken pageId="79" pageNumber="80" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="79" pageNumber="80">
The holotype male is labeled &quot;USA. UT[AH]. Grand: Green River (15.3 km N;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°6.6'W</geoCoordinate>
; 1255 m), 15 Aug 2008,D.&amp;W.N.Mathis/USNM ENT 00118101 [plastic bar code label]/HOLOTYPE ♂
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
W. Mathis &amp; T. Zatwarnicki USNM [red].&quot; The holotype is double mounted (minuten in a block of plastic), is in excellent condition, and is deposited in the USNM. Forty-one paratypes (32♂, 9♀; USNM) bear the same label data as the holotype. Other paratypes are as follows: UTAH. Emery: Green River (3 km N;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.983334">38°59'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-110.15">110°09'W</geoCoordinate>
), 5 Aug 1992, D. and W. N. Mathis (7♂, 4♀; USNM); Green River (Green River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.993332">38°59.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.14167">110°08.5'W</geoCoordinate>
; 1240 m), 31 Jul 2007, D. and W. N. Mathis (6♂, 1♀; USNM); San Rafael River (22.5 km SW Green River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.928333">38°55.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.40833">110°24.5'W</geoCoordinate>
; 1270 m), 31 Jul 2007, D. and W. N. Mathis (1♀; USNM). Grand: Crystal Geyser (14.5 km SE Green River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.93833">38°56.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.135">110°08.1'W</geoCoordinate>
), 15 Aug 2008, D. and W. N. Mathis (27♂, 1♀; USNM); Swasey Beach (15.3 km N Green River;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°07'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°06.6'W</geoCoordinate>
; Green River; 1255 m), 29 May-30 Jul 2007, 2008, D. and W. N. Mathis (7♂, 1♀; USNM); Thompson Spring (8.9 km N Thompson Springs;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.038334">39°02.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.723335">109°43.4'W</geoCoordinate>
; 1740 m), 1-16 Aug 2007, 2008, D. and W. N. Mathis (9♂, 4♀; USNM).
</paragraph>
</subSubSection>
<subSubSection pageId="79" pageNumber="80" type="type locality">
<paragraph pageId="79" pageNumber="80">Type locality.</paragraph>
<paragraph pageId="79" pageNumber="80">
United States. Utah. Grand: Swasey Beach (15.3 km N Green River;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°07'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°06.6'W</geoCoordinate>
; Green River; 1255 m).
</paragraph>
</subSubSection>
<subSubSection lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80" type="other specimens examined">
<paragraph pageId="79" pageNumber="80">Other specimens examined.</paragraph>
<paragraph pageId="79" pageNumber="80">
Nearctic. UNITED STATES. ARIZONA. Mojave: Littlefield (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.885">36°53.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-113.93">113°55.8'W</geoCoordinate>
), 1 May 2002, D. and W. N. Mathis (4♂, 4♀; USNM).
</paragraph>
<paragraph pageId="79" pageNumber="80">
CALIFORNIA. San Bernardino: Afton (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.038334">35°02.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-116.38167">116°22.9'W</geoCoordinate>
), 28 Apr 2002, D. and W. N. Mathis (1♀; USNM).
</paragraph>
<paragraph pageId="79" pageNumber="80">
GEORGIA. Floyd: Lavender (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="34.291668">34°17.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-85.34167">85°20.5'W</geoCoordinate>
), 23 Aug 1910, J. C. Bradley (1♂; ANSP). Tift: Tifton (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="31.45">31°27'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.50833">83°30.5'W</geoCoordinate>
), 6 Sep-Oct 1896 (1♂, 2♀; ANSP).
</paragraph>
<paragraph pageId="79" pageNumber="80">
ILLINOIS. Cook: Chicago (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.883335">41°53'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-87.63167">87°37.9'W</geoCoordinate>
) (1♂, 1♀; ANSP).
</paragraph>
<paragraph pageId="79" pageNumber="80">
INDIANA. Tippecanoe: Lafayette (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="40.416668">40°25'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-86.875">86°52.5'W</geoCoordinate>
), 24 May-5 Sep 1916, E. W. Stafford (2♂; ANSP).
</paragraph>
<paragraph pageId="79" pageNumber="80">
MICHIGAN. Kalamazoo: Kalamazoo (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.291668">42°17.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-85.58667">85°35.2'W</geoCoordinate>
), 10 May 1936, C. W. Sabrosky (1♂; ANSP).
</paragraph>
<paragraph pageId="79" pageNumber="80">
NEVADA. Clark: Las Vegas Wash (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.09">36°05.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-114.97833">114°58.7'W</geoCoordinate>
), 11 Apr 2005, D. and W. N. Mathis (2♂; USNM).
</paragraph>
<paragraph pageId="79" pageNumber="80">
NEW MEXICO. Catron: Gila River (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="33.226665">33°13.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.25166">106°15.1'W</geoCoordinate>
; 1750 m), 15 Aug 2007, D. and W. N. Mathis (4♂; USNM). Chaves: Roswell (9 km E; Pecos River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="33.396667">33°23.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-104.4">104°24'W</geoCoordinate>
; 1060 m), 10 Aug 2007, D. and W. N. Mathis (7♂, 3♀; USNM). Grant: Bill Evans Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="32.868332">32°52.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-108.575">108°34.5'W</geoCoordinate>
; 1416 m), 14 Aug 2007, D. and W. N. Mathis (1♂; USNM); Mimbres River (NM Hwy. 61 &amp; Royal John Mine Road;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="32.73">32°43.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-107.86667">107°52'W</geoCoordinate>
; 1665 m), 13-22 Aug 2007, 2009, D. and W. N. Mathis, T. Zatwarnicki (4♂, 4♀; USNM). Lincoln: Capitan (3.2 km E; Salado Creek;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="33.543335">33°32.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-105.53833">105°32.3'W</geoCoordinate>
; 1890 m), 10 Aug 2007, D. and W. N. Mathis (11♂; USNM). Otero: Alamagordo (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="32.9">32°54'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-105.96">105°57.6'W</geoCoordinate>
), 6-12 May 1902 (1♂, 1♀; ANSP). Valencia:
<normalizedToken originalValue="Río">Rio</normalizedToken>
Puerco (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="34.796665">34°47.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.99167">106°59.5'W</geoCoordinate>
; 1575 m), 9 Aug 2007, D. and W. N. Mathis (4♂, 3♀; USNM).
</paragraph>
<paragraph pageId="80" pageNumber="81">
<pageBreakToken pageId="80" pageNumber="81" start="start">OKLAHOMA</pageBreakToken>
. Pontotoc: Ada (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="34.775">34°46.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-96.67834">96°40.7'W</geoCoordinate>
), 18 Jul 1937, Standish-Kaiser (1♂; ANSP).
</paragraph>
<paragraph pageId="80" pageNumber="81">
TENNESSEE. Shelby: Meeman Shelby State Park (Mississippi River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.37">35°22.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-90.075">90°04.5'W</geoCoordinate>
; 73 m), 10 Jun 2004, W. N. Mathis (8♂, 4♀; USNM).
</paragraph>
<paragraph pageId="80" pageNumber="81">
TEXAS. Augustine: Rayburn Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="31.066668">31°04'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-95.083336">95°05'W</geoCoordinate>
), 15 May 1993, D. and W. N. Mathis (1♀; USNM). Fort Bend: Sugar Land (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="29.62">29°37.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-95.635">95°38.1'W</geoCoordinate>
), 11 Jul 1933, R. Nabors, C. W. Sabrosky (1♂; ANSP). Hidalgo: Bentsen, Rio Grande Valley State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="26.173334">26°10.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-98.38333">98°23'W</geoCoordinate>
), 30- Nov-2 Dec 1978, E. E. Gris sell, A. S. Menke (1♀; USNM). Jim Wells: Mathis (7.5 km S; Nueces River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="28.036667">28°02.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-97.87">97°52.2'W</geoCoordinate>
; 15 m), 6 Jun 2004, W. N. Mathis (4♂, 1♀; USNM). Kimble: Junction (South Llano River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="30.493334">30°29.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-99.75166">99°45.1'W</geoCoordinate>
; 510 m), 4 Jun 2004, W. N. Mathis (1♀; USNM). Travis: Austin (Zilker Park;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="30.263334">30°15.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-97.77167">97°46.3'W</geoCoordinate>
), 2 Jun 2004, W. N. Mathis (2♂, 1♀; USNM).
</paragraph>
<paragraph lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
UTAH. Garfield: Willow Tank, Hurricane Wash (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.386665">37°23.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-111.13333">111°08'W</geoCoordinate>
), 22 May 2001, D. and W. N. Mathis (5♂, 1♀; USNM). Kane: Drip Tank Canyon (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.323334">37°19.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-111.53">111°31.8'W</geoCoordinate>
), 15 May 2001, D. and W. N. Mathis (2♂; USNM); Kanab (6.5 km N;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.145">37°08.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-112.54">112°32.4'W</geoCoordinate>
), 14 May 2001, D. and W. N. Mathis (1♀; USNM). San Juan: Blanding (Reservoir 4;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.665">37°39.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.495">109°29.7'W</geoCoordinate>
; 1975 m), 20 Aug 2013, D. and W. N. Mathis (7♂, 5♀; USNM); Blanding, Recapture Reservoir (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.665">37°39.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.441666">109°26.5'W</geoCoordinate>
; 1840 m), 23 Jul 2013, D. and W. N. Mathis (7♂, 4♀; USNM); Johnson
<pageBreakToken pageId="81" pageNumber="82" start="start">Creek</pageBreakToken>
meadow (22 km N Blanding:
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.791668">37°47.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.50833">109°30.5'W</geoCoordinate>
; 2384 m), 20 Aug 2013, D. and W. N. Mathis (1♀; USNM).
</paragraph>
<paragraph pageId="81" pageNumber="82">
VIRGINIA. Essex: Tappahannock (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.93">37°55.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.85667">76°51.4'W</geoCoordinate>
; Rappahannock River), 18 Sep 2004, D. and W. N. Mathis (1♂; USNM). Fairfax: Great Falls (Patowmack Canal;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.001667">39°00.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.253334">77°15.2'W</geoCoordinate>
), 3 Oct 2006, D. and W. N. Mathis (1♂; USNM); Turkey Run (mouth;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.15667">77°09.4'W</geoCoordinate>
), 10 Jul-7 Sep 2006, 2007, 2008, D. and W. N. Mathis, H. B. Williams (9♂, 3♀; USNM). Patrick: Meadows of Dan (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.736668">36°44.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.38167">80°22.9'W</geoCoordinate>
), 18 May 2005, D. and W. N. Mathis (2♀; USNM). Stafford: Aquia Harbour, Lions Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.45">38°27'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
), 21 Jul-4 Sep 2006, 2008, D. and W. N. Mathis (5♂; USNM); Aquia Landing (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.386665">38°23.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-77.316666">77°19'W</geoCoordinate>
), 7 May 2005, D. and W. N. Mathis (2♂, 1♀; USNM); Falmouth (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.32">38°19.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.46833">77°28.1'W</geoCoordinate>
; Rappahannock River; 9 m), 26 May-30 Jun 2007, D. and W. N. Mathis (4♂; USNM).
</paragraph>
<paragraph pageId="81" pageNumber="82">
WASHINGTON. Franklin: Columbia River N of Pasco (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.358334">46°21.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.25833">119°15.5'W</geoCoordinate>
), 29 Jul 1998, W. N. Mathis (6♂; USNM); Lake Kahlotus (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.65">46°39'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-118.53333">118°32'W</geoCoordinate>
), 25 Jul 1992, D. and W. N. Mathis (1♀; USNM); Levey (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.083332">46°05'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-118.86667">118°52'W</geoCoordinate>
; Snake River), 27 Jul 1992, D. and W. N. Mathis (27♂, 8♀; USNM); Ringold (NW Pasco;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.506668">46°30.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.255">119°15.3'W</geoCoordinate>
; Columbia River), 30 Jul 1998, D. and W. N. Mathis (2♂; USNM); Ringold (4.8 km N;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.515">46°30.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.255">119°15.3'W</geoCoordinate>
), Columbia River, 2 Jul 1988, D. and W. N. Mathis (1♂, 1♀; USNM); Snake River, Levey Landing (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.083332">46°05'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-118.86667">118°52'W</geoCoordinate>
), 27 Jul 1992, D. and W. N. Mathis (29♂, 7♀; USNM). Klickitat: Glenn Wood, Klickitat River (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="45.696667">45°41.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-121.291664">121°17.5'W</geoCoordinate>
), 17 Jun 1917, A. L. Melander (1♀; ANSP). Whitman: Pullman (3.2 km NW;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.733334">46°44'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-117.18166">117°10.9'W</geoCoordinate>
), 11 Jul 1988, D. and W. N. Mathis (1♂; USNM).
</paragraph>
<paragraph pageId="81" pageNumber="82">
WEST VIRGINIA. Mercer: Ceres (Kee Reservoir;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.306667">37°18.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.17333">81°10.4'W</geoCoordinate>
; 757 m), 24 Sep 2007, D. and W. N. Mathis (2♂; USNM). Summers: Hinton (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.696667">37°41.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-80.88333">80°53'W</geoCoordinate>
; New River; 427 m), 26 Sep 2007, D. and W. N. Mathis (2♂, 2♀; USNM). Wyoming: R. D. Bailey Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.595">37°35.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.78">81°46.8'W</geoCoordinate>
; 324 m), 25 Sep 2007, D. and W. N. Mathis (1♂, 1♀; USNM).
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="distribution">
<paragraph pageId="81" pageNumber="82">Distribution</paragraph>
<paragraph pageId="81" pageNumber="82">(Fig. 153). Nearctic: United States (Arizona, California, Georgia, Illinois, Indiana, Michigan, Nevada, New Mexico, Oklahoma, Tennessee, Texas, Utah, Virginia, Washington, West Virginia).</paragraph>
<caption pageId="81" pageNumber="82">
<paragraph pageId="81" pageNumber="82">
Figure 153. Distribution of
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
Mathis and Zatwarnicki.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="etymology">
<paragraph pageId="81" pageNumber="82">Etymology.</paragraph>
<paragraph pageId="81" pageNumber="82">The species epithet, avanae, is a feminine genitive patronym to recognize Ms. Avan Thayn who guided us to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="remarks">
<paragraph pageId="81" pageNumber="82">Remarks.</paragraph>
<paragraph pageId="81" pageNumber="82">
This is a widespread species that is relatively common in the Nearctic Region. Externally, this species is very similar and evidently closely related to
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma andeum" order="Diptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="andeum">Hydrochasma andeum</taxonomicName>
and
<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
but can be distinguished from either of these two species by having a hindtibia that bears a black setula ventrally near the apex, a nearly parallel-sided ventral epandrial process, and by having a more robustly developed hypandrium, especially the anterior two-thirds (Fig. 151). In addition, the apical portion of the aedeagus is frequently greatly inflated (best seen in lateral view; Fig. 152)
</paragraph>
</subSubSection>
</treatment>
</document>