200 lines
24 KiB
XML
200 lines
24 KiB
XML
<document id="983A2AD3B3AFD5B4EDD20EC8BE609799" ID-CLB-Dataset="229103" ID-DOI="10.11646/zootaxa.5230.2.8" ID-GBIF-Dataset="22bc89fb-1fe5-40c5-ac5e-aa2d7c396e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="7553595" ID-ZooBank="EF8C9FC5-203A-4938-B38F-25D6DC55AE87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1674201493496" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V. & Perkovsky, Evgeny E." docDate="2023" docId="03F262413E21FFCD5EF4E90EFC07FE5A" docLanguage="en" docName="zootaxa.5230.2.8.pdf" docOrigin="Zootaxa 5230 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oracula campbelli Nabozhenko & Perkovsky 2023, sp. n." docType="treatment" docVersion="6" lastPageNumber="248" masterDocId="FFCB1A393E20FFCE5E63EF54FF99FFF7" masterDocTitle="Oracula campbelli sp. n. - the first fossil darkling beetle (Coleoptera: Tenebrionidae Alleculinae) from the late Eocene Rovno amber (Ukraine)" masterLastPageNumber="250" masterPageNumber="245" pageNumber="246" updateTime="1699474185152" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B410D0A7A654B36C03266A8E7B2CF479">
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<mods:title id="2D31507A4F1840A5B17C23E62801306F">Oracula campbelli sp. n. - the first fossil darkling beetle (Coleoptera: Tenebrionidae Alleculinae) from the late Eocene Rovno amber (Ukraine)</mods:title>
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<mods:name id="14E039907247B73EC1B80491A8B07D7C" type="personal">
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<mods:roleTerm id="842915C66BFBDBCA7CA2B6328E49B30B">Author</mods:roleTerm>
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<mods:namePart id="3E1E992821DA0F9AD5385E9D07CBF5DE">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="DCBFBFAE9CBF1E4B9080ACA0502A7369">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia.</mods:affiliation>
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<mods:name id="C1764A69153BB69E7A6D1CE459932911" type="personal">
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<mods:roleTerm id="21A9709D015B71E582DC04D01DDDE7AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BC5D69DFDD04572EA71A7CE1745B2006">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="64073DF35A39824062979D2B94A1A559" type="ORCID">0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="D79501355B8B1AB8A6803280664EFEC6">I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, 01030 Kiev, Ukraine. perkovsk @ gmail. com; https: // orcid. org / 0000 - 0002 - 7959 - 4379 & A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia</mods:affiliation>
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<mods:nameIdentifier id="65690A19991C168B91ECBCE4D7FFEDF9" type="email">perkovsk@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="AB56E4D5F8B858AD3CABDEF458C88AFF">text</mods:typeOfResource>
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<mods:title id="CC873799BB2AC4F18CAE3FE51AA086BB">Zootaxa</mods:title>
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<mods:part id="4A22B4E4F9C6362DCA69A762A4A247CD">
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<mods:date id="A24C96F897C64A52696A8928EA6F1EC1">2023</mods:date>
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<mods:number id="083DC536216D58BF2ED2D07D8548565C">2023-01-20</mods:number>
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<mods:number id="06D6DEB9C6BA424F9B55BF314B9AA214">5230</mods:number>
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<treatment id="03F262413E21FFCD5EF4E90EFC07FE5A" ID-DOI="http://doi.org/10.5281/zenodo.7553597" ID-GBIF-Taxon="209011158" ID-Zenodo-Dep="7553597" LSID="urn:lsid:plazi:treatment:03F262413E21FFCD5EF4E90EFC07FE5A" httpUri="http://treatment.plazi.org/id/03F262413E21FFCD5EF4E90EFC07FE5A" lastPageId="3" lastPageNumber="248" pageId="1" pageNumber="246">
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<subSubSection id="C34180DC3E21FFCF5EF4E90EFE58F982" box="[151,449,1626,1653]" pageId="1" pageNumber="246" type="nomenclature">
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<paragraph id="8BE4D3573E21FFCF5EF4E90EFE58F982" blockId="1.[151,449,1626,1689]" box="[151,449,1626,1653]" pageId="1" pageNumber="246">
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<heading id="D0AC643B3E21FFCF5EF4E90EFE58F982" bold="true" box="[151,449,1626,1653]" fontSize="11" level="1" pageId="1" pageNumber="246" reason="1">
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<emphasis id="B92F0F453E21FFCF5EF4E90EFE58F982" bold="true" box="[151,449,1626,1653]" pageId="1" pageNumber="246">
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<taxonomicName id="4C5BA8D43E21FFCF5EF4E90EFEEEF982" authority="Nabozhenko & Perkovsky, 2023" authorityName="Nabozhenko & Perkovsky" authorityYear="2023" box="[151,375,1626,1653]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="campbelli" status="sp. nov.">
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<emphasis id="B92F0F453E21FFCF5EF4E90EFEEEF982" bold="true" box="[151,375,1626,1653]" italics="true" pageId="1" pageNumber="246">Oracula campbelli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21CB23E3E21FFCF5F1CE90FFE58F982" box="[383,449,1627,1653]" pageId="1" pageNumber="246" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34180DC3E21FFCF5EF4E92BFF72F96E" box="[151,235,1663,1689]" pageId="1" pageNumber="246" type="description">
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<paragraph id="8BE4D3573E21FFCF5EF4E92BFF72F96E" blockId="1.[151,449,1626,1689]" box="[151,235,1663,1689]" pageId="1" pageNumber="246">
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(
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<figureCitation id="1360CFD23E21FFCF5EFCE92BFF79F96E" box="[159,224,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[152,255,1784,1808]" captionTargetBox="[151,1436,705,1761]" captionTargetId="figure-240@2.[151,1436,705,1761]" captionTargetPageId="2" captionText="FIGURE 1. Oracula campbelli sp. n. from Rovno amber, habitus: A—dorsal view; B—lateral view (arrow in the inset shows the dark area of hollow claw); C—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7553599" httpUri="https://zenodo.org/record/7553599/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34180DC3E21FFCF5EF4E993FE29F8DE" pageId="1" pageNumber="246" type="materials_examined">
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<paragraph id="8BE4D3573E21FFCF5EF4E993FBD6F916" blockId="1.[151,1437,1734,2013]" box="[151,1103,1734,1761]" pageId="1" pageNumber="246">
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<emphasis id="B92F0F453E21FFCF5EF4E993FEDCF917" bold="true" box="[151,325,1734,1761]" pageId="1" pageNumber="246">Type material.</emphasis>
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<materialsCitation id="3B33D90A3E21FFCF5F2FE993FBD6F916" ID-GBIF-Occurrence="4016388301" box="[332,1103,1734,1761]" collectionCode="SIZK" location="Klesov" pageId="1" pageNumber="246" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
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<emphasis id="B92F0F453E21FFCF5F2FE993FD8CF917" bold="true" box="[332,533,1734,1761]" pageId="1" pageNumber="246">
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<typeStatus id="54E06DF53E21FFCF5F2FE993FE20F916" box="[332,441,1735,1761]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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<collectionCode id="ED4A4B923E21FFCF5C7FE993FDC3F916" box="[540,602,1735,1761]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="246">SIZK</collectionCode>
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K-3503,
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<location id="8E84858C3E21FFCF5CA6E993FC8CF916" LSID="urn:lsid:plazi:treatment:03F262413E21FFCD5EF4E90EFC07FE5A:8E84858C3E21FFCF5CA6E993FC8CF916" box="[709,789,1735,1761]" name="Klesov" pageId="1" pageNumber="246" stateProvince="Rivne">Klesov</location>
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,
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<collectingRegion id="499F1DB53E21FFCF5D43E993FCF2F916" box="[800,875,1735,1761]" country="Ukraine" name="Rivne" pageId="1" pageNumber="246">Rovno</collectingRegion>
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amber, late Eocene.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE4D3573E21FFCF5EA4E9BEFE29F8DE" blockId="1.[151,1437,1734,2013]" pageId="1" pageNumber="246">
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<emphasis id="B92F0F453E21FFCF5EA4E9BEFEEBF8F3" bold="true" box="[199,370,1770,1796]" pageId="1" pageNumber="246">Syninclusions.</emphasis>
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<collectionCode id="ED4A4B923E21FFCF5FE7E9BFFE5BF8F2" box="[388,450,1771,1797]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="246">SIZK</collectionCode>
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K-3502
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<taxonomicName id="4C5BA8D43E21FFCF5C5DE9BFFD5DF8F2" box="[574,708,1771,1797]" class="Insecta" family="Ortheziidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Ortheziidae</taxonomicName>
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,
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<taxonomicName id="4C5BA8D43E21FFCF5CB9E9BFFCE7F8F2" box="[730,894,1771,1797]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
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;
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<collectionCode id="ED4A4B923E21FFCF5DF6E9BFFC4AF8F2" box="[917,979,1771,1797]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="246">SIZK</collectionCode>
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K-3503
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<taxonomicName id="4C5BA8D43E21FFCF5A2DE9BFFB45F8F2" box="[1102,1244,1771,1797]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
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,
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<taxonomicName id="4C5BA8D43E21FFCF5A96E9BFFA0CF8F2" box="[1269,1429,1771,1797]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
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,
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<taxonomicName id="4C5BA8D43E21FFCF5EF4E85BFE84F8DE" authorityName="Haliday" authorityYear="1839" box="[151,285,1807,1833]" class="Insecta" family="Scelionidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Scelionidae</taxonomicName>
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,
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<taxonomicName id="4C5BA8D43E21FFCF5F44E85BFE35F8DE" authorityName="Sundevall" authorityYear="1833" box="[295,428,1807,1833]" class="Arachnida" family="Theridiidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="246" phylum="Arthropoda" rank="family">Theridiidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C34180DC3E21FFCC5EA4E866FBE7FD5E" lastPageId="2" lastPageNumber="247" pageId="1" pageNumber="246" type="description">
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<paragraph id="8BE4D3573E21FFCF5EA4E866FAB5F82A" blockId="1.[151,1437,1734,2013]" pageId="1" pageNumber="246">
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<emphasis id="B92F0F453E21FFCF5EA4E866FEC8F8BB" bold="true" box="[199,337,1842,1868]" pageId="1" pageNumber="246">Description</emphasis>
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. Body light, slender, elongate, widest at elytral midlength. Legs probably yellow, because of femoral apices are clearly dark. Length
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<quantity id="4CA37EB23E21FFCF5CE9E803FD7FF886" box="[650,742,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="246" unit="mm" value="8.8">8.8 mm</quantity>
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. Head wider, than anterior margin of pronotum. Eyes large, transverse diameter of one eye 1.3 times as long as interocular distance. Ocular index 26.Apical maxillary palpomere strongly securiform, with moderately oblique apical margin. Antennae not serrate, with strongly elongate and thin antennomeres (comparative length of antennae and the majority of antennomeres cannot be determined).
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</paragraph>
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<paragraph id="8BE4D3573E22FFCC5EA4EFC3FBE4FEEA" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="247">Prothorax. Pronotum weakly transverse (1.375 as wide as long), widest at base, looks bare, 1.3 times as wide as head at eye level. Lateral margins straight at least at basal third and weakly rounded and narrowed to apical margin at apical 2/3. Base of pronotum weakly bisinuate; posterior angles look weakly acute. Puncturation of pronotum dense and moderately coarse, puncture diameter subequal to distance between punctures.</paragraph>
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<paragraph id="8BE4D3573E22FFCC5EA4EE73FED2FE26" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="247">Pterothorax. Scutellar shield with slightly rounded margins. Elytral base wider than pronotal base. Elytra 1.5 as wide as head and 1.13 times as wide (at maximal level of elytral basal part) as pronotum; surface looks bare (pubescence may not be visible under the white coating), at least without dense long erect pubescence; with deep striae, consisting of round deep punctures; puncture diameter slightly longer than interpuncture distance. Interstriae slightly convex.</paragraph>
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<paragraph id="8BE4D3573E22FFCC5EA4EE8FFDB1FDEE" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="247">
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Abdomen (
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<figureCitation id="1360CFD23E22FFCC5F26EE8FFE04FE02" box="[325,413,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1784,1808]" captionTargetBox="[151,1436,705,1761]" captionTargetId="figure-240@2.[151,1436,705,1761]" captionTargetPageId="2" captionText="FIGURE 1. Oracula campbelli sp. n. from Rovno amber, habitus: A—dorsal view; B—lateral view (arrow in the inset shows the dark area of hollow claw); C—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7553599" httpUri="https://zenodo.org/record/7553599/files/figure.png" pageId="2" pageNumber="247">Fig. 1C</figureCitation>
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). Abdominal ventrites 1–3 beaded, 4 and 5 ones not beaded; ventrite 3 with oblique laterial margins; ventrite 5 widely rounded.
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</paragraph>
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<paragraph id="8BE4D3573E22FFCC5EA4ED77FBE7FD5E" blockId="2.[151,1437,150,681]" pageId="2" pageNumber="247">
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Legs (
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<figureCitation id="1360CFD23E22FFCC5F69ED77FE18FDCA" box="[266,385,547,573]" captionStart="FIGURE 1" captionStartId="2.[152,255,1784,1808]" captionTargetBox="[151,1436,705,1761]" captionTargetId="figure-240@2.[151,1436,705,1761]" captionTargetPageId="2" captionText="FIGURE 1. Oracula campbelli sp. n. from Rovno amber, habitus: A—dorsal view; B—lateral view (arrow in the inset shows the dark area of hollow claw); C—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7553599" httpUri="https://zenodo.org/record/7553599/files/figure.png" pageId="2" pageNumber="247">Figs1B,C</figureCitation>
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). Femora thickened, strong, light, with dark apices.Tibiae without dense suberected pubescence. Pro- and mesotibiae clearly curved, metatibiae slightly curved. Protarsomeres 1–3 strongly widened, wider than mesotarsomeres 1–3. Preapical tarsomeres strongly widened, wider than other tarsomeres, strongly lobed. Tarsal claws large and hollow (
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<figureCitation id="1360CFD23E22FFCC5FCAEDDBFD99FD5E" box="[425,512,655,681]" captionStart="FIGURE 1" captionStartId="2.[152,255,1784,1808]" captionTargetBox="[151,1436,705,1761]" captionTargetId="figure-240@2.[151,1436,705,1761]" captionTargetPageId="2" captionText="FIGURE 1. Oracula campbelli sp. n. from Rovno amber, habitus: A—dorsal view; B—lateral view (arrow in the inset shows the dark area of hollow claw); C—ventral view." figureDoi="http://doi.org/10.5281/zenodo.7553599" httpUri="https://zenodo.org/record/7553599/files/figure.png" pageId="2" pageNumber="247">Fig. 1B</figureCitation>
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, inset); anterior claws with 19 (9+10) teeth on both sides
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</paragraph>
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</subSubSection>
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||
<caption id="DF2483DF3E22FFCC5EFBE9ACFDE5F8C3" ID-DOI="http://doi.org/10.5281/zenodo.7553599" ID-Zenodo-Dep="7553599" httpUri="https://zenodo.org/record/7553599/files/figure.png" pageId="2" pageNumber="247" startId="2.[152,255,1784,1808]" targetBox="[151,1436,705,1761]" targetPageId="2">
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||
<paragraph id="8BE4D3573E22FFCC5EFBE9ACFDE5F8C3" blockId="2.[151,1436,1784,1844]" pageId="2" pageNumber="247">
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||
<emphasis id="B92F0F453E22FFCC5EFBE9ACFE8AF8E7" bold="true" box="[152,275,1784,1808]" pageId="2" pageNumber="247">FIGURE 1</emphasis>
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.
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||
<taxonomicName id="4C5BA8D43E22FFCC5F7CE9ACFE47F8E7" authority="Nabozhenko & Perkovsky, 2023" authorityName="Nabozhenko & Perkovsky" authorityYear="2023" box="[287,478,1784,1808]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="campbelli" status="sp. nov.">
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<emphasis id="B92F0F453E22FFCC5F7CE9ACFE47F8E7" box="[287,478,1784,1808]" italics="true" pageId="2" pageNumber="247">Oracula campbelli</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B92F0F453E22FFCC5F87E9ACFD87F8E7" bold="true" box="[484,542,1784,1808]" pageId="2" pageNumber="247">
|
||
<taxonomicNameLabel id="A21CB23E3E22FFCC5F87E9ACFD87F8E7" box="[484,542,1784,1808]" pageId="2" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Rovno amber, habitus: A—dorsal view; B—lateral view (arrow in the inset shows the dark area of hollow claw); C—ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34180DC3E22FFCD5EA4E837FC07FE5A" lastPageId="3" lastPageNumber="248" pageId="2" pageNumber="247" type="etymology">
|
||
<paragraph id="8BE4D3573E22FFCC5EA4E837FCF6F857" blockId="2.[151,1436,1890,2025]" pageId="2" pageNumber="247">
|
||
<emphasis id="B92F0F453E22FFCC5EA4E837FED7F88A" bold="true" box="[199,334,1891,1917]" pageId="2" pageNumber="247">Etymology.</emphasis>
|
||
The species is named in memory of Dr. Milton J. Campbell (
|
||
<date id="FFE5F5973E22FFCC5D88E836FB5CF88B" box="[1003,1221,1890,1917]" pageId="2" pageNumber="247" value="1936-12-19">December 19, 1936</date>
|
||
—
|
||
<date id="FFE5F5973E22FFCC5A81E836FA09F88A" box="[1250,1424,1890,1917]" pageId="2" pageNumber="247" value="2022-08-06">August 6, 2022</date>
|
||
), the famous specialist on comb-clawed beetles of the New World.
|
||
</paragraph>
|
||
<paragraph id="8BE4D3573E22FFCC5EA4E8FEFD84F81F" blockId="2.[151,1436,1890,2025]" pageId="2" pageNumber="247">
|
||
<emphasis id="B92F0F453E22FFCC5EA4E8FEFE7AF833" bold="true" box="[199,483,1962,1988]" pageId="2" pageNumber="247">Comparative diagnosis.</emphasis>
|
||
The new species belongs to the subgenus
|
||
<emphasis id="B92F0F453E22FFCC5DAFE8FEFBB5F833" box="[972,1068,1962,1988]" italics="true" pageId="2" pageNumber="247">Duocula</emphasis>
|
||
Novák,
|
||
<quantity id="4CA37EB23E22FFCC5AECE8FFFB7FF833" box="[1167,1254,1962,1989]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="2" pageNumber="247" unit="in" value="2019.0">2019 in</quantity>
|
||
the presence of teeth on both sides of hollow claw.
|
||
</paragraph>
|
||
<paragraph id="8BE4D3573E23FFCD5EA4EFC3FC07FE5A" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="248">
|
||
The new species is most similar externally to
|
||
<taxonomicName id="4C5BA8D43E23FFCD5CDDEFC3FC20FF46" authority="Novak, 2019" authorityName="Novak" authorityYear="2019" box="[702,953,151,177]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="amica">
|
||
<emphasis id="B92F0F453E23FFCD5CDDEFC3FCBAFF47" box="[702,803,151,177]" italics="true" pageId="3" pageNumber="248">O. amica</emphasis>
|
||
Novák, 2019
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F34C93C73E23FFCD5D9FEFC3FBA6FF46" box="[1020,1087,151,177]" name="Nepal" pageId="3" pageNumber="248">Nepal</collectingCountry>
|
||
in the presence of dark femoral apices with more or less distinct colour borders, straight lateral margins of pronotum at least at basal third and the absence of erected or suberected long dense pubescence on elytra and tibiae. The new species differs from
|
||
<taxonomicName id="4C5BA8D43E23FFCD5B55EF8BFA05FF0F" authorityName="Novak" authorityYear="2019" box="[1334,1436,223,249]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="amica">
|
||
<emphasis id="B92F0F453E23FFCD5B55EF8BFA05FF0F" box="[1334,1436,223,249]" italics="true" pageId="3" pageNumber="248">O. amica</emphasis>
|
||
</taxonomicName>
|
||
by the less dense puncturation of the pronotum (puncture diameter longer than interpuncture distance in
|
||
<taxonomicName id="4C5BA8D43E23FFCD5B4FEE57FA0BFEEB" authorityName="Novak" authorityYear="2019" box="[1324,1426,259,285]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="amica">
|
||
<emphasis id="B92F0F453E23FFCD5B4FEE57FA0BFEEB" box="[1324,1426,259,285]" italics="true" pageId="3" pageNumber="248">O. amica</emphasis>
|
||
</taxonomicName>
|
||
) and the shape of the pronotum with pointed at apex basal angles (narrowly rounded apices in
|
||
<taxonomicName id="4C5BA8D43E23FFCD5AD8EE73FABBFEB7" authorityName="Novak" authorityYear="2019" box="[1211,1314,295,321]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="amica">
|
||
<emphasis id="B92F0F453E23FFCD5AD8EE73FABBFEB7" box="[1211,1314,295,321]" italics="true" pageId="3" pageNumber="248">O. amica</emphasis>
|
||
</taxonomicName>
|
||
). The new species has darkened part only at the most distal part of femora, while femora of
|
||
<taxonomicName id="4C5BA8D43E23FFCD5A79EE1FFB19FE93" authorityName="Novak" authorityYear="2019" box="[1050,1152,331,357]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="amica">
|
||
<emphasis id="B92F0F453E23FFCD5A79EE1FFB19FE93" box="[1050,1152,331,357]" italics="true" pageId="3" pageNumber="248">O. amica</emphasis>
|
||
</taxonomicName>
|
||
are darkened at the distal quarter.
|
||
<taxonomicName id="4C5BA8D43E23FFCD5E97EE3BFE51FE7E" authority="Nabozhenko & Perkovsky, 2023" authorityName="Nabozhenko & Perkovsky" authorityYear="2023" box="[244,456,367,393]" class="Insecta" family="Tenebrionidae" genus="Oracula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="campbelli" status="sp. nov.">
|
||
<emphasis id="B92F0F453E23FFCD5E97EE3BFE51FE7E" box="[244,456,367,393]" italics="true" pageId="3" pageNumber="248">Oracula campbelli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92F0F453E23FFCD5FB2EE3BFD8BFE7E" bold="true" box="[465,530,367,393]" pageId="3" pageNumber="248">
|
||
<taxonomicNameLabel id="A21CB23E3E23FFCD5FB2EE3BFD8BFE7E" box="[465,530,367,393]" pageId="3" pageNumber="248" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all species of the genus by the presence of 19 teeth on male claws (more than
|
||
<quantity id="4CA37EB23E23FFCD5F7BEEC7FEC8FE5A" box="[280,337,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="248" unit="in" value="30.0">30 in</quantity>
|
||
all extant representatives; more than
|
||
<quantity id="4CA37EB23E23FFCD5C96EEC7FCB7FE5A" box="[757,814,403,429]" metricMagnitude="0" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="248" unit="in" value="40.0">40 in</quantity>
|
||
<emphasis id="B92F0F453E23FFCD5D56EEC7FC0CFE5A" box="[821,917,403,429]" italics="true" pageId="3" pageNumber="248">Duocula</emphasis>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |