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;
<tableCitation id="C6D9E6A3FF9FFFF5FDE564B5FDF6FD40" box="[515,632,690,713]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="12.[97,162,1598,1621]" captionStartId-1="13.[192,257,1007,1030]" captionText-0="Table 1: Measurements (in mm) of Isometrus (I.) maculatus (Ar.-MWHU-HNLS0701 and Ar.-MWHU-HNSY0701), Isometrus (R.) hainanensis (after Lourenço, Qi &amp; Zhu 2005), Lychas mucronatus (Ar.-MWHU-HNDF0601 and Ar.-MWHU-HNDF0602) and Liocheles australasiae (Ar.-MWHU-HNSL0616 and Ar.-MWHU-HNNL 0617). A-Width = anterior width, P-Width = posterior width." captionText-1="Table 2: Pectinal teeth counts of Hainan Island scorpion species." pageId="5" pageNumber="4">Tables 12</tableCitation>
)
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</paragraph>
<paragraph id="8BE4D318FF9FFFF5FF266204FEF6FBB0" blockId="5.[192,802,751,1081]" pageId="5" pageNumber="4">
<taxonomicName id="4C5BA89BFF9FFFF5FF266204FEFFFBB0" authority=": Tikader &amp; Bastawade, 1983: 286" authorityName="Tikader &amp; Bastawade" authorityPageNumber="286" authorityYear="1983" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="europaeus" subGenus="Raddyanus">
<emphasis id="B92F0F0AFF9FFFF5FF266204FEA7FB93" box="[192,297,1027,1050]" italics="true" pageId="5" pageNumber="4">Isometrus</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FEA46204FE36FB93" box="[322,440,1027,1050]" italics="true" pageId="5" pageNumber="4">Raddyanus</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FE376204FDCEFB93" box="[465,576,1027,1050]" italics="true" pageId="5" pageNumber="4">europaeus</emphasis>
: Tikader &amp; Bastawade, 1983: 286
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3418093FF9FFFF5FF166258FE58FAC4" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BE4D318FF9FFFF5FF166258FE39FB1C" blockId="5.[192,803,1119,1909]" pageId="5" pageNumber="4">
<materialsCitation id="3B33D945FF9FFFF5FF166258FED0FB1C" country="Suriname" location="Pennsylvania" pageId="5" pageNumber="4" specimenCount="2" typeStatus="holotype">
<emphasis id="B92F0F0AFF9FFFF5FF166258FDFFFBFF" bold="true" box="[240,625,1119,1142]" pageId="5" pageNumber="4">
<typeStatus id="54E06DBAFF9FFFF5FF166258FEA4FBFF" box="[240,298,1119,1142]" pageId="5" pageNumber="4">Type</typeStatus>
locality and
<typeStatus id="54E06DBAFF9FFFF5FE246258FE7DFBFF" box="[450,499,1119,1142]" pageId="5" pageNumber="4">type</typeStatus>
repository
</emphasis>
: “
<collectingCountry id="F34C9388FF9FFFF5FD6C6258FD7CFBFF" box="[650,754,1119,1142]" name="Suriname" pageId="5" pageNumber="4">Suriname</collectingCountry>
and
<location id="8E8485C3FF9FFFF5FF246279FEDAFB1C" LSID="urn:lsid:plazi:treatment:03F2620EFF9FFFF8FF056494FD0AFB93:8E8485C3FF9FFFF5FF246279FEDAFB1C" box="[194,340,1150,1173]" country="Suriname" name="Pennsylvania" pageId="5" pageNumber="4">Pennsylvania</location>
</materialsCitation>
;
<collectionCode id="ED4A4BDDFF9FFFF5FE8E6279FE3DFB1C" box="[360,435,1150,1173]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="5" pageNumber="4">NHRS</collectionCode>
.
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FF16629BFE58FAC4" blockId="5.[192,803,1119,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FF16629BFE5BFB3A" bold="true" box="[240,469,1180,1203]" pageId="5" pageNumber="4">Material examined</emphasis>
:
<materialsCitation id="3B33D945FF9FFFF5FE17629BFE41FAC4" collectingDate="2006-08" collectingDateMax="2007-04" collectingDateMin="2006-08" collectionCode="MWHU" collectorName="Yi-Bao Ma &amp; Hui Liu &amp; Liang Wu" location="Hui Liu" municipality="Lingshui District" pageId="5" pageNumber="4" specimenCode="HNLS0701-70, MWHU-HNSY0601-06" specimenCount="76" specimenCount-adult="70" specimenCount-immature="5" specimenCount-juvenile="1" stateProvince="Hainan">
<collectingRegion id="499F1DFAFF9FFFF5FE17629BFDB0FB3A" box="[497,574,1180,1203]" country="China" name="Hainan" pageId="5" pageNumber="4">Hainan</collectingRegion>
:
<collectingMunicipality id="6B804962FF9FFFF5FDBF629BFC90FB3A" box="[601,798,1180,1203]" pageId="5" pageNumber="4">Lingshui District</collectingMunicipality>
,
<date id="FFE5F5D8FF9FFFF5FF2662BCFE96FB5B" box="[192,280,1211,1234]" bridgedPair="/" pageId="5" pageNumber="4" value="2007-04">
<collectingDate id="EFA10C30FF9FFFF5FF2662BCFE96FB5B" box="[192,280,1211,1234]" bridgedPair="/" pageId="5" pageNumber="4" value="2007-04">IV/2007</collectingDate>
</date>
,
<location id="8E8485C3FF9FFFF5FECC62BCFE0CFB5B" LSID="urn:lsid:plazi:treatment:03F2620EFF9FFFF8FF056494FD0AFB93:8E8485C3FF9FFFF5FECC62BCFE0CFB5B" box="[298,386,1211,1234]" municipality="Lingshui District" name="Hui Liu" pageId="5" pageNumber="4" stateProvince="Hainan">Hui Liu</location>
and
<collectorName id="26AEB6CEFF9FFFF5FE5862BCFDB5FB5B" box="[446,571,1211,1234]" pageId="5" pageNumber="4">Yi-Bao Ma</collectorName>
leg.,
<specimenCount id="9D5D1891FF9FFFF5FD9862BCFD56FB5B" box="[638,728,1211,1234]" pageId="5" pageNumber="4" type="adult">24 adult</specimenCount>
and
<specimenCount id="9D5D1891FF9FFFF5FCF262BCFEA8FB78" pageId="5" pageNumber="4" type="immature">5 immature</specimenCount>
males,
<specimenCount id="9D5D1891FF9FFFF5FE9F62DDFE5EFB78" box="[377,464,1242,1265]" pageId="5" pageNumber="4" type="adult">41 adult</specimenCount>
females (
<collectionCode id="ED4A4BDDFF9FFFF5FDDE62DDFD1AFB78" box="[568,660,1242,1265]" pageId="5" pageNumber="4">MWHU</collectionCode>
, Ar.-
<collectionCode id="ED4A4BDDFF9FFFF5FD2D62DDFCACFB78" box="[715,802,1242,1265]" pageId="5" pageNumber="4">MWHU</collectionCode>
-
<specimenCode id="DBFD7B63FF9FFFF5FF2162FFFEE2FA86" box="[199,364,1272,1295]" collectionCode="HNLS" pageId="5" pageNumber="4">HNLS070170</specimenCode>
); Sanya,
<date id="FFE5F5D8FF9FFFF5FE3E62FFFDCBFA86" box="[472,581,1272,1295]" bridgedPair="/" pageId="5" pageNumber="4" value="2006-08">
<collectingDate id="EFA10C30FF9FFFF5FE3E62FFFDCBFA86" box="[472,581,1272,1295]" bridgedPair="/" pageId="5" pageNumber="4" value="2006-08">VIII/2006</collectingDate>
</date>
,
<collectorName id="26AEB6CEFF9FFFF5FDB562FFFD27FA86" box="[595,681,1272,1295]" pageId="5" pageNumber="4">Hui Liu</collectorName>
and Ying-
<collectorName id="26AEB6CEFF9FFFF5FF266310FEA2FAA7" box="[192,300,1303,1326]" pageId="5" pageNumber="4">Liang Wu</collectorName>
leg.,
<specimenCount id="9D5D1891FF9FFFF5FE816310FE3EFAA7" box="[359,432,1303,1326]" pageId="5" pageNumber="4" type="adult">5 adult</specimenCount>
females,
<specimenCount id="9D5D1891FF9FFFF5FDFE6310FD0CFAA7" box="[536,642,1303,1326]" pageId="5" pageNumber="4" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED4A4BDDFF9FFFF5FD766310FD62FAA7" box="[656,748,1303,1326]" pageId="5" pageNumber="4">MWHU</collectionCode>
, Ar.-
<specimenCode id="DBFD7B63FF9FFFF5FF266331FE46FAC4" box="[192,456,1334,1357]" collectionCode="MWHU-HNSY" pageId="5" pageNumber="4">MWHU-HNSY060106</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3418093FF9FFFF5FF166353FC49FE5A" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BE4D318FF9FFFF5FF166353FEFCF952" blockId="5.[192,803,1119,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FF166353FED0FAE2" bold="true" box="[240,350,1364,1387]" pageId="5" pageNumber="4">Diagnosis</emphasis>
: Moderate to large size, measuring up to
<quantity id="4CA37EFDFF9FFFF5FF246374FE83FA03" box="[194,269,1395,1418]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="mm" value="60.0">60 mm</quantity>
in adult male and about
<quantity id="4CA37EFDFF9FFFF5FDFD6374FDE9FA03" box="[539,615,1395,1418]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="4" unit="mm" value="50.0">50 mm</quantity>
in adult females. General coloration yellowish to pale yellow with symmetrical blackish-brown patterns in both adults and juveniles. Carinae and granulations moderately marked. Carapace strongly emarginate, with an open V-shaped angle. Pectines moderately long; pectinal tooth count
<quantity id="4CA37EFDFF9FFFF5FF26602CFEAEF9CB" box="[192,288,1579,1602]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="5" pageNumber="4" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">1619 in</quantity>
both sexes. Dentate margins of fixed and movable fingers of pedipalp chela with 7 almost linear rows of granules. Trichobothrium
<emphasis id="B92F0F0AFF9FFFF5FDEC606FFDABF9F6" box="[522,549,1640,1663]" italics="true" pageId="5" pageNumber="4">db</emphasis>
on chela of pedipalp situated between trichobothria
<emphasis id="B92F0F0AFF9FFFF5FDF46080FDA9F917" box="[530,551,1671,1694]" italics="true" pageId="5" pageNumber="4">dt</emphasis>
and
<emphasis id="B92F0F0AFF9FFFF5FD866080FDFDF917" box="[608,627,1671,1694]" italics="true" pageId="5" pageNumber="4">et</emphasis>
. Subaculear tubercle strongly developed and triangular, with two ventral granules.
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FF1660E4FC49FE5A" blockId="5.[192,803,1119,1909]" lastBlockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FF1660E4FE04F973" bold="true" box="[240,394,1763,1786]" pageId="5" pageNumber="4">Relationships</emphasis>
: In its general morphology and coloration,
<taxonomicName id="4C5BA89BFF9FFFF5FED96105FD25F890" authorityName=": Vachon" authorityYear="1972" box="[319,683,1794,1817]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="maculatus" subGenus="Isometrus">
<emphasis id="B92F0F0AFF9FFFF5FED96105FE26F890" box="[319,424,1794,1817]" italics="true" pageId="5" pageNumber="4">Isometrus</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FE586105FDA9F890" box="[446,551,1794,1817]" italics="true" pageId="5" pageNumber="4">Isometrus</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FDDB6105FD25F890" box="[573,683,1794,1817]" italics="true" pageId="5" pageNumber="4">maculatus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C5BA89BFF9FFFF5FF266127FDD6F8BE" authorityName="Lourenco, Qi &amp; Zhu" authorityYear="2005" box="[192,600,1824,1847]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hainanensis" subGenus="Reddyanus">
<emphasis id="B92F0F0AFF9FFFF5FF266127FEA7F8BE" box="[192,297,1824,1847]" italics="true" pageId="5" pageNumber="4">Isometrus</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FEA16127FE32F8BE" box="[327,444,1824,1847]" italics="true" pageId="5" pageNumber="4">Reddyanus</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FE3F6127FDD6F8BE" box="[473,600,1824,1847]" italics="true" pageId="5" pageNumber="4">hainanensis</emphasis>
</taxonomicName>
, described from Southeast region in
<collectingRegion id="499F1DFAFF9FFFF5FE4F6138FE78F8DF" box="[425,502,1855,1878]" country="China" name="Hainan" pageId="5" pageNumber="4">Hainan</collectingRegion>
Island. It can be distinguished from the latter species by the following char- acters: (i) trichobothrium
<emphasis id="B92F0F0AFF9FFFF5FB9366C0FB1EFF57" box="[1141,1168,199,222]" italics="true" pageId="5" pageNumber="4">db</emphasis>
on chela of pedipalp situated between trichobothria
<emphasis id="B92F0F0AFF9FFFF5FB7666E1FB2BFF74" box="[1168,1189,230,253]" italics="true" pageId="5" pageNumber="4">dt</emphasis>
and
<emphasis id="B92F0F0AFF9FFFF5FB3D66E1FB60FF74" box="[1243,1262,230,253]" italics="true" pageId="5" pageNumber="4">et</emphasis>
, while in
<taxonomicName id="4C5BA89BFF9FFFF5FA8766E1FC30FE92" authorityName="Lourenco, Qi &amp; Zhu" authorityYear="2005" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hainanensis" subGenus="Reddyanus">
<emphasis id="B92F0F0AFF9FFFF5FA8766E1FAFFFF74" box="[1377,1393,230,253]" italics="true" pageId="5" pageNumber="4">I.</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FA6666E1FA16FF74" box="[1408,1432,230,253]" italics="true" pageId="5" pageNumber="4">R.</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FCD86703FC30FE92" box="[830,958,260,283]" italics="true" pageId="5" pageNumber="4">hainanensis</emphasis>
</taxonomicName>
it is situated between trichobothria e
<emphasis id="B92F0F0AFF9FFFF5FA806703FAE3FE92" box="[1382,1389,260,283]" italics="true" pageId="5" pageNumber="4">t</emphasis>
and
<emphasis id="B92F0F0AFF9FFFF5FCD86724FCD2FEB3" box="[830,860,291,314]" italics="true" pageId="5" pageNumber="4">est</emphasis>
; (ii) subaculear tubercle with two ventral granules, while in
<taxonomicName id="4C5BA89BFF9FFFF5FC446745FBE7FED0" authorityName="Lourenco, Qi &amp; Zhu" authorityYear="2005" box="[930,1129,322,345]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hainanensis" subGenus="Reddyanus">
<emphasis id="B92F0F0AFF9FFFF5FC446745FC3CFED0" box="[930,946,322,345]" italics="true" pageId="5" pageNumber="4">I.</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FC276745FC56FED0" box="[961,984,322,345]" italics="true" pageId="5" pageNumber="4">R.</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FC0F6745FBE7FED0" box="[1001,1129,322,345]" italics="true" pageId="5" pageNumber="4">hainanensis</emphasis>
</taxonomicName>
subaculear tubercle has five ventral granules; and (iii) the total length of adult males about
<quantity id="4CA37EFDFF9FFFF5FC626778FC5AFE1F" box="[900,980,383,406]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="mm" value="60.0">60 mm</quantity>
and
<quantity id="4CA37EFDFF9FFFF5FBE96778FBD0FE1F" box="[1039,1118,383,406]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="4" unit="mm" value="50.0">50 mm</quantity>
in adult females, while in
<taxonomicName id="4C5BA89BFF9FFFF5FA776778FC7EFE3C" authorityName="Lourenco, Qi &amp; Zhu" authorityYear="2005" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="hainanensis" subGenus="Reddyanus">
<emphasis id="B92F0F0AFF9FFFF5FA776778FA2FFE1F" box="[1425,1441,383,406]" italics="true" pageId="5" pageNumber="4">I.</emphasis>
(
<emphasis id="B92F0F0AFF9FFFF5FCA16799FCD1FE3C" box="[839,863,414,437]" italics="true" pageId="5" pageNumber="4">R.</emphasis>
)
<emphasis id="B92F0F0AFF9FFFF5FC976799FC7EFE3C" box="[881,1008,414,437]" italics="true" pageId="5" pageNumber="4">hainanensis</emphasis>
</taxonomicName>
it is about
<quantity id="4CA37EFDFF9FFFF5FB956799FB67FE3C" box="[1139,1257,414,437]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="5" pageNumber="4" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">5055 mm</quantity>
and
<quantity id="4CA37EFDFF9FFFF5FAC56799FA17FE3C" box="[1315,1433,414,437]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="5" pageNumber="4" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">3035 mm</quantity>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C3418093FF9FFFF8FC8867DCFD0AFB93" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4" type="description">
<paragraph id="8BE4D318FF9FFFF5FC8867DCFB26FD98" blockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FC8867DCFC7CFE7B" bold="true" box="[878,1010,475,498]" pageId="5" pageNumber="4">Description</emphasis>
: Based on male specimens from Lingshui District (
<collectingRegion id="499F1DFAFF9FFFF5FBE467FDFBDFFD98" box="[1026,1105,506,529]" country="China" name="Hainan" pageId="5" pageNumber="4">Hainan</collectingRegion>
Island).
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FC88641FFAB0FC54" blockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FC88641FFC6DFDA6" box="[878,995,536,559]" italics="true" pageId="5" pageNumber="4">Coloration</emphasis>
: Generally yellowish to pale yellow. Prosoma: Carapace yellowish with blackish-brown patterns; eyes surrounded by black pigment (
<figureCitation id="1360CF9DFF9FFFF5FAED6451FAD8FDE4" box="[1291,1366,598,621]" captionStart="Figures 14" captionStartId="3.[192,278,1870,1893]" captionTargetBox="[191,1430,191,1841]" captionTargetId="figure-6@3.[192,1430,192,1840]" captionTargetPageId="3" captionText="Figures 14: Habitus of Isometrus maculatus. 12. Male (Ar.-MWHU-HNLS0701), dorsal and ventral views. 34. Female (Ar.-MWHU-HNSY0701), dorsal and ventral views. Scale bars: 10.0 mm." pageId="5" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="1360CF9DFF9FFFF5FA806451FA1EFDE4" box="[1382,1424,598,621]" captionStart="Figures 511" captionStartId="4.[192,278,1851,1874]" captionTargetBox="[218,1422,176,1804]" captionTargetId="figure-10@4.[204,1450,168,1824]" captionTargetPageId="4" captionText="Figures 511: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701). 5. Carapace. 6. Lateral eyes. 7. Femur dorsal aspect. 89. Patella dorsal and external aspects. 1011. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="5" pageNumber="4">5, 6</figureCitation>
).
<taxonomicName id="4C5BA89BFF9FFFF5FCD86473FBA3FD02" authority=": Tergites" authorityName="Tergites" box="[830,1069,628,651]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Mesosoma: Tergites</taxonomicName>
yellowish with symmetrical blackish-brown stripes; Sternum, genital operculum, sternites IIIVI, pectines yellowish, sternites IIIVII yellowish with symmetrical brown patterns. Metasomal segments pale yellow, with some diffuse, brownish spots. Vesicle pale yellow; aculeus yellowish at base and reddish-brown at tip. Chelicerae pale yellow with brownish variegated spots; base of fingers pale yellow, rest of fingers blackish-brown, teeth reddish. Pedipalps pale yellow with brownish spots; chela fingers reddishbrown; rows of granules on dentate margins of fingers dark reddish. Legs yellowish with diffuse spots.
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FC8865E3FA0FFA86" blockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FC8865E3FC40FC72" box="[878,974,996,1019]" italics="true" pageId="5" pageNumber="4">Prosoma</emphasis>
: Tegument coarse. Surface of carapace coarsely granular with a few smooth patches. Anterior margin of carapace strongly emarginate, with an open Vshaped angle. Carapace carinae weakly developed. Median ocular tubercle anterior to the centre of the carapace with granules; median eyes separated by one ocular diameter. Three pairs of lateral eyes present on anteriorlateral portion with granular lateral ocular tubercles (some specimens with two eyespot pairs dorso-posteriorly and posteriorly respectively, see in
<figureCitation id="1360CF9DFF9FFFF5FAEB62FFFAFAFA86" box="[1293,1396,1272,1295]" captionStart="Figures 511" captionStartId="4.[192,278,1851,1874]" captionTargetBox="[218,1422,176,1804]" captionTargetId="figure-10@4.[204,1450,168,1824]" captionTargetPageId="4" captionText="Figures 511: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701). 5. Carapace. 6. Lateral eyes. 7. Femur dorsal aspect. 89. Patella dorsal and external aspects. 1011. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="5" pageNumber="4">Figs. 56</figureCitation>
).
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FC886310FBA8F9CB" blockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<taxonomicName id="4C5BA89BFF9FFFF5FC886310FBD0FAA7" authority=": Tegument" authorityName="Tegument" box="[878,1118,1303,1326]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B92F0F0AFF9FFFF5FC886310FC6EFAA7" box="[878,992,1303,1326]" italics="true" pageId="5" pageNumber="4">Mesosoma</emphasis>
: Tegument
</taxonomicName>
coarse. Tergites IVI with a median carina; weak on I, weak to moderate on IIVI. Tergite VII with two lateral pairs of carinae moderate to strong; median carinae present in proximal half, moderately developed. Intercarinal spaces moderately granular. Sternum pentagonal with a genital operculum subtriangular. Sternites smooth and shiny. Pectines moderately long, tooth count
<quantity id="4CA37EFDFF9FFFF5FB8963E9FB59F98C" box="[1135,1239,1518,1541]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="5" pageNumber="4" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">1617 in</quantity>
the male, fulcra marked (
<figureCitation id="1360CF9DFF9FFFF5FC47600BFC7CF9AA" box="[929,1010,1548,1571]" captionStart="Figures 1221" captionStartId="6.[192,278,1810,1833]" captionTargetBox="[212,1409,210,1746]" captionTargetId="figure-10@6.[192,1440,192,1789]" captionTargetPageId="6" captionText="Figures 1221: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 1213. Chelicera, dorsal and ventral aspects. 16. Sternum, genital operculum and pectines. 18. Telson, lateral aspect. Female (Ar.-MWHU-HNSY0701): 1415. Chelicera, dorsal and ventral aspects. 17. Sternum, genital operculum and pectines. 19. Telson, lateral aspect. 2021. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="5" pageNumber="4">Fig. 16</figureCitation>
). Sternites III to VI smooth; VII with four granular carinae.
</paragraph>
<paragraph id="8BE4D318FF9FFFF5FC88604DFABBF8BE" blockId="5.[830,1441,199,1909]" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FC88604DFC53F9E8" box="[878,989,1610,1633]" italics="true" pageId="5" pageNumber="4">Metasoma</emphasis>
: Tegument coarse with few granules. Segments I and II with 10 carinae, crenulate; IIIIV with 8 carinae, crenulate. Segment V with five carinae. Dorsal furrows of all segments developed, smooth; intercarinal spaces very weakly granular to smooth. Telson very weakly granular, almost smooth with one vestigial ventral carina; subaculear tubercle marked and triangular, with 2 granules on the ventral surface.
</paragraph>
<paragraph id="8BE4D318FF9FFFF8FC886138FDB7FE3C" blockId="5.[830,1441,199,1909]" lastBlockId="8.[192,804,199,1050]" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4">
<emphasis id="B92F0F0AFF9FFFF5FC886138FC57F8DF" box="[878,985,1855,1878]" italics="true" pageId="5" pageNumber="4">Pedipalps</emphasis>
: Tegument almost smooth. Femora with all carinae marked crenulate, intercarinal spaces without granules in ventral aspect and other aspects with few granules. Patella with seven crenulate carinae. Chela with obsolete carinae. Dentate margins on fixed and movable fingers composed of six linear rows of granules (
<figureCitation id="1360CF9DFF92FFF8FF2E6745FE91FED0" box="[200,287,322,345]" captionStart="Figures 2229" captionStartId="7.[192,278,1843,1866]" captionTargetBox="[223,1418,224,1767]" captionTargetId="figure-6@7.[192,1441,192,1813]" captionTargetPageId="7" captionText="Figures 2229: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 25. Dentate margin of movable finger, showing rows of granules. 2627. Chela, dorsal and ventral aspects. Female (Ar.-MWHU-HNSY0701): 22. Femur dorsal aspect. 2324. Patella external and dorsal aspects. 2829. Chela dorsal and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">Fig. 25</figureCitation>
). Trichobothrial pattern
<typeStatus id="54E06DBAFF92FFF8FDA06745FDFAFED0" box="[582,628,322,345]" pageId="8" pageNumber="7">type</typeStatus>
A, orthobothriotaxic (
<figureCitation id="1360CF9DFF92FFF8FEC46767FE29FEFE" box="[290,423,352,375]" captionStart="Figures 2229" captionStartId="7.[192,278,1843,1866]" captionTargetBox="[223,1418,224,1767]" captionTargetId="figure-6@7.[192,1441,192,1813]" captionTargetPageId="7" captionText="Figures 2229: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 25. Dentate margin of movable finger, showing rows of granules. 2627. Chela, dorsal and ventral aspects. Female (Ar.-MWHU-HNSY0701): 22. Femur dorsal aspect. 2324. Patella external and dorsal aspects. 2829. Chela dorsal and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">Figs. 26, 27</figureCitation>
). For the position and distribution of trichobothria of the femur, patella and chela of pedipalps see
<figureCitation id="1360CF9DFF92FFF8FEC96799FE1AFE3C" box="[303,404,414,437]" captionStart="Figures 511" captionStartId="4.[192,278,1851,1874]" captionTargetBox="[218,1422,176,1804]" captionTargetId="figure-10@4.[204,1450,168,1824]" captionTargetPageId="4" captionText="Figures 511: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701). 5. Carapace. 6. Lateral eyes. 7. Femur dorsal aspect. 89. Patella dorsal and external aspects. 1011. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">Figs. 79</figureCitation>
,
<figureCitation id="1360CF9DFF92FFF8FE466799FDBCFE3C" box="[416,562,414,437]" captionStart="Figures 2229" captionStartId="7.[192,278,1843,1866]" captionTargetBox="[223,1418,224,1767]" captionTargetId="figure-6@7.[192,1441,192,1813]" captionTargetPageId="7" captionText="Figures 2229: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 25. Dentate margin of movable finger, showing rows of granules. 2627. Chela, dorsal and ventral aspects. Female (Ar.-MWHU-HNSY0701): 22. Femur dorsal aspect. 2324. Patella external and dorsal aspects. 2829. Chela dorsal and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">2224, 2629</figureCitation>
.
</paragraph>
<caption id="DF248390FF9CFFF6FF266115FDCCF8FD" pageId="6" pageNumber="5" startId="6.[192,278,1810,1833]" targetBox="[212,1409,210,1746]" targetPageId="6">
<paragraph id="8BE4D318FF9CFFF6FF266115FDCCF8FD" blockId="6.[192,1441,1810,1908]" pageId="6" pageNumber="5">
<emphasis id="B92F0F0AFF9CFFF6FF266115FEFEF8A0" bold="true" box="[192,368,1810,1833]" pageId="6" pageNumber="5">Figures 1221:</emphasis>
<taxonomicName id="4C5BA89BFF9CFFF6FE9A6113FDC7F8A0" authorityName=": Vachon" authorityYear="1972" box="[380,585,1812,1833]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B92F0F0AFF9CFFF6FE9A6113FDC7F8A0" box="[380,585,1812,1833]" italics="true" pageId="6" pageNumber="5">Isometrus maculatus</emphasis>
</taxonomicName>
. Male (Ar.-MWHU-HNLS0701):
<emphasis id="B92F0F0AFF9CFFF6FC4F6113FC65F8A0" bold="true" box="[937,1003,1812,1833]" pageId="6" pageNumber="5">1213.</emphasis>
Chelicera, dorsal and ventral aspects.
<emphasis id="B92F0F0AFF9CFFF6FA646113FA2EF8A0" bold="true" box="[1410,1440,1812,1833]" pageId="6" pageNumber="5">16.</emphasis>
Sternum, genital operculum and pectines.
<emphasis id="B92F0F0AFF9CFFF6FDBD612AFDF6F8CB" bold="true" box="[603,632,1837,1858]" pageId="6" pageNumber="5">18.</emphasis>
Telson, lateral aspect. Female (Ar.-MWHU-HNSY0701):
<emphasis id="B92F0F0AFF9CFFF6FB53612AFB78F8CB" bold="true" box="[1205,1270,1837,1858]" pageId="6" pageNumber="5">1415.</emphasis>
Chelicera, dorsal and ventral aspects.
<emphasis id="B92F0F0AFF9CFFF6FE656141FE2CF8D2" bold="true" box="[387,418,1862,1883]" pageId="6" pageNumber="5">17.</emphasis>
Sternum, genital operculum and pectines.
<emphasis id="B92F0F0AFF9CFFF6FCDA6141FCD5F8D2" bold="true" box="[828,859,1862,1883]" pageId="6" pageNumber="5">19.</emphasis>
Telson, lateral aspect.
<emphasis id="B92F0F0AFF9CFFF6FBDE6141FBF4F8D2" bold="true" box="[1080,1146,1862,1883]" pageId="6" pageNumber="5">2021.</emphasis>
Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="DF248390FF9DFFF7FF266134FB98F8F2" pageId="7" pageNumber="6" startId="7.[192,278,1843,1866]" targetBox="[223,1418,224,1767]" targetPageId="7">
<paragraph id="8BE4D318FF9DFFF7FF266134FB98F8F2" blockId="7.[192,1440,1843,1915]" pageId="7" pageNumber="6">
<emphasis id="B92F0F0AFF9DFFF7FF266134FEE7F8C3" bold="true" box="[192,361,1843,1866]" pageId="7" pageNumber="6">Figures 2229:</emphasis>
<taxonomicName id="4C5BA89BFF9DFFF7FE966132FDB6F8C3" authorityName=": Vachon" authorityYear="1972" box="[368,568,1845,1866]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B92F0F0AFF9DFFF7FE966132FDB6F8C3" box="[368,568,1845,1866]" italics="true" pageId="7" pageNumber="6">Isometrus maculatus</emphasis>
</taxonomicName>
. Male (Ar.-MWHU-HNLS0701):
<emphasis id="B92F0F0AFF9DFFF7FC616132FC2AF8C3" bold="true" box="[903,932,1845,1866]" pageId="7" pageNumber="6">25.</emphasis>
Dentate margin of movable finger, showing rows of granules.
<emphasis id="B92F0F0AFF9DFFF7FEC66149FEEFF8EA" bold="true" box="[288,353,1870,1891]" pageId="7" pageNumber="6">2627.</emphasis>
Chela, dorsal and ventral aspects. Female (Ar.-MWHU-HNSY0701):
<emphasis id="B92F0F0AFF9DFFF7FBFC6149FBB9F8EA" bold="true" box="[1050,1079,1870,1891]" pageId="7" pageNumber="6">22.</emphasis>
Femur dorsal aspect.
<emphasis id="B92F0F0AFF9DFFF7FAF26149FAD8F8EA" bold="true" box="[1300,1366,1870,1891]" pageId="7" pageNumber="6">2324.</emphasis>
Patella external and dorsal aspects.
<emphasis id="B92F0F0AFF9DFFF7FE286161FD9EF8F2" bold="true" box="[462,528,1894,1915]" pageId="7" pageNumber="6">2829.</emphasis>
Chela dorsal and ventral aspects. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BE4D318FF92FFF8FF1667BBFEE7FDA6" blockId="8.[192,804,199,1050]" pageId="8" pageNumber="7">
<emphasis id="B92F0F0AFF92FFF8FF1667BBFEEBFE5A" box="[240,357,444,467]" italics="true" pageId="8" pageNumber="7">Chelicerae</emphasis>
: Tegument smooth. Tibiae with reticulated pattern, with dentition characteristic of buthids (
<figureCitation id="1360CF9DFF92FFF8FF2167FDFEC1FD98" box="[199,335,506,529]" captionStart="Figures 1221" captionStartId="6.[192,278,1810,1833]" captionTargetBox="[212,1409,210,1746]" captionTargetId="figure-10@6.[192,1440,192,1789]" captionTargetPageId="6" captionText="Figures 1221: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 1213. Chelicera, dorsal and ventral aspects. 16. Sternum, genital operculum and pectines. 18. Telson, lateral aspect. Female (Ar.-MWHU-HNSY0701): 1415. Chelicera, dorsal and ventral aspects. 17. Sternum, genital operculum and pectines. 19. Telson, lateral aspect. 2021. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">Figs. 12, 13</figureCitation>
); two small basal teeth on dorsal aspect of movable finger.
</paragraph>
<paragraph id="8BE4D318FF92FFF8FF166430FD4BFCCA" blockId="8.[192,804,199,1050]" pageId="8" pageNumber="7">
<emphasis id="B92F0F0AFF92FFF8FF166430FEADFDC7" box="[240,291,567,590]" italics="true" pageId="8" pageNumber="7">Legs</emphasis>
: Tegument smooth except dorsal aspect of femora with granules. Femora with 2 granular carinae in internal aspect, 2 granular carinae in external aspect and 1 granular carina in dorsal aspect. Patellae with 1 dentate carina in internal, 1 granular carina in dorsal aspect and 3 granular carinae in dorsal aspect. Tibiae with few setae, without spurs. Basitarsi with some setae and two lateral pedal spurs. Tarsi ventrally with two rows of short setae. Tarsal ungues curved and hook-like.
</paragraph>
<paragraph id="8BE4D318FF92FFF8FF16654CFDA3FC54" blockId="8.[192,804,199,1050]" pageId="8" pageNumber="7">
<emphasis id="B92F0F0AFF92FFF8FF16654CFED8FCEB" box="[240,342,843,866]" italics="true" pageId="8" pageNumber="7">Variation</emphasis>
: Females coloration very similar to males. Sexual dimorphism: females differ from males (
<figureCitation id="1360CF9DFF92FFF8FD2F656DFF43FC16" captionStart="Figures 14" captionStartId="3.[192,278,1870,1893]" captionTargetBox="[191,1430,191,1841]" captionTargetId="figure-6@3.[192,1430,192,1840]" captionTargetPageId="3" captionText="Figures 14: Habitus of Isometrus maculatus. 12. Male (Ar.-MWHU-HNLS0701), dorsal and ventral views. 34. Female (Ar.-MWHU-HNSY0701), dorsal and ventral views. Scale bars: 10.0 mm." pageId="8" pageNumber="7">Figs. 3 4</figureCitation>
,
<figureCitation id="1360CF9DFF92FFF8FF06658FFEADFC16" box="[224,291,904,927]" captionStart="Figures 1221" captionStartId="6.[192,278,1810,1833]" captionTargetBox="[212,1409,210,1746]" captionTargetId="figure-10@6.[192,1440,192,1789]" captionTargetPageId="6" captionText="Figures 1221: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 1213. Chelicera, dorsal and ventral aspects. 16. Sternum, genital operculum and pectines. 18. Telson, lateral aspect. Female (Ar.-MWHU-HNSY0701): 1415. Chelicera, dorsal and ventral aspects. 17. Sternum, genital operculum and pectines. 19. Telson, lateral aspect. 2021. Metasomal segment V, lateral and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">1921</figureCitation>
,
<figureCitation id="1360CF9DFF92FFF8FED0658FFEF4FC16" box="[310,378,904,927]" captionStart="Figures 2229" captionStartId="7.[192,278,1843,1866]" captionTargetBox="[223,1418,224,1767]" captionTargetId="figure-6@7.[192,1441,192,1813]" captionTargetPageId="7" captionText="Figures 2229: Isometrus maculatus. Male (Ar.-MWHU-HNLS0701): 25. Dentate margin of movable finger, showing rows of granules. 2627. Chela, dorsal and ventral aspects. Female (Ar.-MWHU-HNSY0701): 22. Femur dorsal aspect. 2324. Patella external and dorsal aspects. 2829. Chela dorsal and ventral aspects. Scale bars: 1.0 mm." pageId="8" pageNumber="7">2226</figureCitation>
) in having bigger mesosoma, shorter pedipalps and metasoma. Measurements, see
<tableCitation id="C6D9E6A3FF92FFF8FD2265A0FC92FC37" box="[708,796,935,958]" captionStart="Table 1" captionStartId="12.[97,162,1598,1621]" captionText="Table 1: Measurements (in mm) of Isometrus (I.) maculatus (Ar.-MWHU-HNLS0701 and Ar.-MWHU-HNSY0701), Isometrus (R.) hainanensis (after Lourenço, Qi &amp; Zhu 2005), Lychas mucronatus (Ar.-MWHU-HNDF0601 and Ar.-MWHU-HNDF0602) and Liocheles australasiae (Ar.-MWHU-HNSL0616 and Ar.-MWHU-HNNL 0617). A-Width = anterior width, P-Width = posterior width." pageId="8" pageNumber="7">Table 1</tableCitation>
. Pectinal tooth counts, see
<tableCitation id="C6D9E6A3FF92FFF8FE3065C1FDA7FC54" box="[470,553,966,989]" captionStart="Table 2" captionStartId="13.[192,257,1007,1030]" captionText="Table 2: Pectinal teeth counts of Hainan Island scorpion species." pageId="8" pageNumber="7">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE4D318FF92FFF8FF1665E3FDB8FC72" blockId="8.[192,804,199,1050]" box="[240,566,996,1019]" pageId="8" pageNumber="7">
<emphasis id="B92F0F0AFF92FFF8FF1665E3FECCFC72" box="[240,322,996,1019]" italics="true" pageId="8" pageNumber="7">Habitat</emphasis>
: Under stones or bark.
</paragraph>
<paragraph id="8BE4D318FF92FFF8FF166204FD0AFB93" blockId="8.[192,804,199,1050]" box="[240,644,1027,1050]" pageId="8" pageNumber="7">
<emphasis id="B92F0F0AFF92FFF8FF166204FEFCFB93" box="[240,370,1027,1050]" italics="true" pageId="8" pageNumber="7">Distribution</emphasis>
: See Fet &amp; Lowe (2000).
</paragraph>
</subSubSection>
</treatment>
</document>