151 lines
21 KiB
XML
151 lines
21 KiB
XML
<document id="681BB617A881CF55ADDF623C022B3197" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58517FFA45991FD055115F899" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) subrubrum Silva, Louzada & Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="488" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="488" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="69C0822C6D5FF1CBB16C3222F47708AC">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="C5A874595BD01E7DCCEAAE60E224A38B" type="personal">
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<mods:namePart id="02478F377C4B2E52FD6CAB5FE4DD875E">Silva, Fernando A. B.</mods:namePart>
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<mods:roleTerm id="5373E7B9476E7ABFCFBD9C787F24A281">Author</mods:roleTerm>
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<mods:namePart id="13DA712FBE4A79E0C11284216D747BDD">Louzada, Júlio</mods:namePart>
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<mods:roleTerm id="76B0E458B8F29A8433B8A5F6EF548040">Author</mods:roleTerm>
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<mods:namePart id="F04ACFEDE52B8EBF2D04AA7A085ED8D3">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:typeOfResource id="A534F7F0141FEADCA539C2D96502B18A">text</mods:typeOfResource>
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<mods:title id="B0C6EF9CBAD9F51EE134E9C42E32A251">Zootaxa</mods:title>
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<mods:part id="8872CC255AA076EC97FAE4CFD2B1D881">
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<mods:date id="81618C39E398B11D855E5CCEEBBFE2DB">2015</mods:date>
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<mods:detail id="AEBE47048868692EB3C929C0C3FBB0E8" type="volume">
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<mods:number id="2CCC0E5B8B53BE83286340652594AE5B">3925</mods:number>
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<mods:number id="A1570DAABC33AEC21753937C42DAF7A2">4</mods:number>
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<mods:classification id="799D2C1A222294F56373380841A14401">journal article</mods:classification>
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<mods:identifier id="94E7DEB7866EE6128401B007C269C112" type="DOI">10.11646/zootaxa.3925.4.1</mods:identifier>
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<treatment id="03DE87D58517FFA45991FD055115F899" ID-DOI="http://doi.org/10.5281/zenodo.6115758" ID-GBIF-Taxon="119598957" ID-Zenodo-Dep="6115758" LSID="urn:lsid:plazi:treatment:03DE87D58517FFA45991FD055115F899" httpUri="http://treatment.plazi.org/id/03DE87D58517FFA45991FD055115F899" lastPageNumber="488" pageId="37" pageNumber="488">
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<subSubSection id="C36D65488517FFA45991FD0554FDFCEC" pageId="37" pageNumber="488" type="nomenclature">
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<paragraph id="8BC836C38517FFA45991FD055641FCC2" blockId="37.[151,782,764,824]" box="[151,782,764,790]" pageId="37" pageNumber="488">
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<heading id="D08081AF8517FFA45991FD055641FCC2" bold="true" box="[151,782,764,790]" fontSize="11" level="1" pageId="37" pageNumber="488" reason="1">
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<emphasis id="B903EAD18517FFA45991FD055641FCC2" bold="true" box="[151,782,764,790]" pageId="37" pageNumber="488">
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<taxonomicName id="4C774D408517FFA45991FD055737FCC2" ID-CoL="34KB3" box="[151,632,764,790]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="subrubrum" status="sp. nov." subGenus="Aganhyboma">
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<emphasis id="B903EAD18517FFA45991FD055462FCC2" bold="true" box="[151,301,764,790]" italics="true" pageId="37" pageNumber="488">Deltochilum</emphasis>
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(
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<emphasis id="B903EAD18517FFA4583BFD055491FCC2" bold="true" box="[317,478,764,790]" italics="true" pageId="37" pageNumber="488">Aganhyboma</emphasis>
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)
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<emphasis id="B903EAD18517FFA458F6FD055737FCC2" bold="true" box="[496,632,764,790]" italics="true" pageId="37" pageNumber="488">subrubrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057AA8517FFA45B78FD055641FCC2" box="[638,782,764,790]" pageId="37" pageNumber="488" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="8BC836C38517FFA45991FCE654FDFCEC" blockId="37.[151,782,764,824]" box="[151,434,799,824]" pageId="37" pageNumber="488">
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(
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<figureCitation id="134C2A468517FFA45999FCE655BAFCEC" box="[159,245,799,824]" captionStart="FIGURES 20 – 23" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[169,1418,225,1833]" captionTargetId="figure@40.[151,1436,193,1836]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Aedeagus in Deltochilum (Aganhyboma). 20. D. paresi new species. 21. D. icariforme. 22. D. ritamourae new species. 23. D. subrubrum new species. Figure 24. Aedeagus illustration showing the angle between the phallobase and the parameres. Figures 25 – 27. Genital segment in Deltochilum (Aganhyboma). 25. D. cupreicolle. 26. D. viridescens new status. 27. D. trisignatum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244799/files/figure.png" pageId="37" pageNumber="488">Figs 23</figureCitation>
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,
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<figureCitation id="134C2A468517FFA45807FCD95450FCEC" box="[257,287,800,824]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="37" pageNumber="488">41</figureCitation>
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, 86, 94, 123)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D65488517FFA45991FC9E510EFC54" box="[151,1089,871,896]" pageId="37" pageNumber="488" type="etymology">
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<paragraph id="8BC836C38517FFA45991FC9E510EFC54" blockId="37.[151,1437,871,1869]" box="[151,1089,871,896]" pageId="37" pageNumber="488">
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<emphasis id="B903EAD18517FFA45991FC9E5451FC54" bold="true" box="[151,286,871,896]" pageId="37" pageNumber="488">Etymology.</emphasis>
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<taxonomicName id="4C774D408517FFA45822FC9154EDFC54" box="[292,418,872,896]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="502" phylum="Arthropoda" rank="species" species="subrubrum" status="sp. nov.">
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<emphasis id="B903EAD18517FFA45822FC9154EDFC54" box="[292,418,872,896]" italics="true" pageId="37" pageNumber="488">Subrubrum</emphasis>
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</taxonomicName>
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(reddish), Latin adjective, refers to the color of individuals.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D65488517FFA459C1FC7551BCFA31" pageId="37" pageNumber="488" type="description">
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<paragraph id="8BC836C38517FFA459C1FC7551BCFA31" blockId="37.[151,1437,871,1869]" pageId="37" pageNumber="488">
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<emphasis id="B903EAD18517FFA459C1FC7554EBFC71" bold="true" box="[199,420,908,933]" pageId="37" pageNumber="488">Description. Body.</emphasis>
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Head, pronotum, elytra and ventral surface dark brown or black, with greenish or reddish metallic sheen. Length 14.8 to 15.8 mm.
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<emphasis id="B903EAD18517FFA45B67FC5657E7FC1C" bold="true" box="[609,680,943,968]" pageId="37" pageNumber="488">Head.</emphasis>
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Foveiform punctures dense, touching on dorsal interocular surface. Punctures around clypeus shallower, smaller, and more dispersed than those on the interocular surface. Dorsal interocular distance approximately four times eye width. Lateral margin of gena strongly curved outward, with row of setae around eyes.
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<emphasis id="B903EAD18517FFA45892FBE554BBFBE1" bold="true" box="[404,500,1052,1077]" pageId="37" pageNumber="488">Thorax.</emphasis>
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Surface of pronotum with dense foveiform punctures, touching on the sides and anterior angles (however not as deep as in
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<taxonomicName id="4C774D408517FFA45B9CFBB85679FB8C" box="[666,822,1088,1112]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="icariforme">
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<emphasis id="B903EAD18517FFA45B9CFBB85679FB8C" box="[666,822,1088,1112]" italics="true" pageId="37" pageNumber="488">D. icariforme</emphasis>
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</taxonomicName>
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). Posteromedial punctures separated by at least one diameter. Margin between lateral and posterior angles straight.
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<emphasis id="B903EAD18517FFA45A63FB9D56F6FBA9" bold="true" box="[869,953,1124,1149]" pageId="37" pageNumber="488">Elytra.</emphasis>
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Lateral margins almost parallel (
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<figureCitation id="134C2A468517FFA45C35FB9D50DBFBA8" box="[1331,1428,1124,1149]" captionStart="FIGURES 119 – 123" captionStartId="51.[151,264,1596,1619]" captionTargetBox="[178,1436,193,1570]" captionTargetId="figure@51.[150,1436,193,1576]" captionTargetPageId="51" captionText="FIGURES 119 – 123. Species of D. (Aganhyboma). 119. D. ritamourae new species. 120. D. icariforme. 121. D. paresi new species. 122. D. kolleri new species. 123. D. subrubrum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244810/files/figure.png" pageId="37" pageNumber="488">Fig. 123</figureCitation>
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) and surface slightly irregular (rugose). Elytral striae wider, carinate margins clearly separated (as in Fig. 75). Interstriae clearly convex, with its central portion nearly flat. Interstriae usually more punctate on the sides, close to striae. Foveiform punctures of third interstria denser laterally (punctures can be so scattered at center of interstria to produce smooth midlongitudinal strip) (Fig. 86). Microtubercles of interstriae almost inconspicuous. Basal carina of seventh interstria elongate, length about four or more times the width of the seventh interestria. Basal carina of ninth interstria present in three-quarters of elytra (not reaching anterior limit of apical carina of seventh interstria). Apical carina of third interstria rounded, better defined as tubercle, more prominent than apical carinae of second and fourth interstriae. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria less than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D65488517FFA459C1FA165013F9F8" pageId="37" pageNumber="488" type="materials_examined">
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<paragraph id="8BC836C38517FFA459C1FA165058F9DC" blockId="37.[151,1437,871,1869]" box="[199,1303,1519,1544]" pageId="37" pageNumber="488">
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<emphasis id="B903EAD18517FFA459C1FA165782F9DC" bold="true" box="[199,717,1519,1544]" pageId="37" pageNumber="488">
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Material studied.
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<typeStatus id="54CC88618517FFA4589DFA165743F9DC" box="[411,524,1519,1544]" pageId="37" pageNumber="488" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F36076538517FFA45B1EFA165787F9DC" box="[536,712,1519,1544]" name="Argentina" pageId="37" pageNumber="488">ARGENTINA</collectingCountry>
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:
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</emphasis>
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MISIONES: Loreto, 1955, F. H. Walz (
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<specimenCount id="9D71FD4A8517FFA45D8BFA0951F9F9DC" box="[1165,1206,1520,1544]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
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<collectionCode id="ED66AE068517FFA45DBBFA095044F9DC" box="[1213,1291,1520,1544]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="37" pageNumber="488">DZUP</collectionCode>
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).
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</paragraph>
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<paragraph id="8BC836C38517FFA459C1F9ED5013F9F8" blockId="37.[151,1437,871,1869]" box="[199,1372,1556,1581]" pageId="37" pageNumber="488">
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<emphasis id="B903EAD18517FFA459C1F9ED5764F9F9" bold="true" box="[199,555,1556,1581]" pageId="37" pageNumber="488">
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<typeStatus id="54CC88618517FFA459C1F9ED5470F9F9" box="[199,319,1556,1581]" pageId="37" pageNumber="488" type="paratype">Paratypes</typeStatus>
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[3]:
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<collectingCountry id="F36076538517FFA45870F9ED5769F9F9" box="[374,550,1556,1581]" name="Argentina" pageId="37" pageNumber="488">ARGENTINA</collectingCountry>
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:
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</emphasis>
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MISIONES: Loreto (1 ♀
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<collectionCode id="ED66AE068517FFA45A57F9ED56EBF9F8" box="[849,932,1556,1580]" pageId="37" pageNumber="488">CEMT</collectionCode>
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); same, San Ignacio (
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<specimenCount id="9D71FD4A8517FFA45D93F9ED51F2F9F8" box="[1173,1213,1556,1580]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
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1 ♀
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<collectionCode id="ED66AE068517FFA45DF3F9ED501FF9F8" LSID="urn:lsid:biocol.org:col:34988" box="[1269,1360,1556,1580]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="37" pageNumber="488">MNHN</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D65488517FFA459C1F9CE5115F899" pageId="37" pageNumber="488" type="diagnosis">
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||
<paragraph id="8BC836C38517FFA459C1F9CE55F5F8FC" blockId="37.[151,1437,871,1869]" pageId="37" pageNumber="488">
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<emphasis id="B903EAD18517FFA459C1F9CE5496F984" bold="true" box="[199,473,1591,1616]" pageId="37" pageNumber="488">Diagnosis and remarks</emphasis>
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(within the
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<taxonomicName id="4C774D408517FFA45B58F9C1579AF984" box="[606,725,1592,1616]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="icariforme">
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||
<emphasis id="B903EAD18517FFA45B58F9C1579AF984" box="[606,725,1592,1616]" italics="true" pageId="37" pageNumber="488">icariforme</emphasis>
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</taxonomicName>
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subcomplex): length 14.8 to 15.8 mm; posteromedial punctures of pronotum separated by at least one diameter; length of apical carina of seventh interstria less than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae; microtubercles of elytral interstriae absent or almost inconspicuous. The following aedeagus characters are also diagnostic: parameres symmetric and elongate, directed downward, almost at an angle of 60° with phallobase; apex weakly acuminate (
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<figureCitation id="134C2A468517FFA45D1EF93E5122F934" box="[1048,1133,1735,1760]" captionStart="FIGURES 20 – 23" captionStartId="40.[151,264,1884,1907]" captionTargetBox="[169,1418,225,1833]" captionTargetId="figure@40.[151,1436,193,1836]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Aedeagus in Deltochilum (Aganhyboma). 20. D. paresi new species. 21. D. icariforme. 22. D. ritamourae new species. 23. D. subrubrum new species. Figure 24. Aedeagus illustration showing the angle between the phallobase and the parameres. Figures 25 – 27. Genital segment in Deltochilum (Aganhyboma). 25. D. cupreicolle. 26. D. viridescens new status. 27. D. trisignatum. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244799/files/figure.png" pageId="37" pageNumber="488">Fig. 23</figureCitation>
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). Lamella copulatrix (LC): left lobe subrectangular (
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<figureCitation id="134C2A468517FFA458A8F915574EF8D0" box="[430,513,1772,1797]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="37" pageNumber="488">Fig. 41</figureCitation>
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); right lobe "b"-shaped (
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<figureCitation id="134C2A468517FFA45A12F9155628F8D0" box="[788,871,1772,1797]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="37" pageNumber="488">Fig. 41</figureCitation>
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). Fronto-lateral peripheral sclerite (FLP) as in
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<figureCitation id="134C2A468517FFA45C76F91555F9F8FC" captionStart="FIGURES 55 – 58" captionStartId="43.[151,264,1783,1806]" captionTargetBox="[205,1383,207,1725]" captionTargetId="figure@43.[174,1383,193,1728]" captionTargetPageId="43" captionText="FIGURES 55 – 58. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 55. D. valgum. 56. D. kolleri new species. 57. D. icariforme. 58. D. ritamourae new species. Figure 59. Complex of axial and subaxial sclerites (A + AS) in D. viridescens new status. Figures 60 – 61. Head in D. (Aganhyboma). 60. D. viridescens new status. 61. D. longiceps new status. Figures 62 – 63. Lateral view of the elytra in D. (Aganhyboma). 62. D. viridescens new status (arrow points the basal carina of ninth interstria). 63. D. finestriatum new species. (arrow points the basal carina of ninth interstria). Figures 64 – 69. Legs. 64. Metafemur in D. viridescens new status. 65. Metafemur in D. longiceps new status. 66. Protibia in D. trisignatum. 67. Metatibia in D. trisignatum. 68. Metatibia in D. viridescens new status. 69. Profemur in D. trisignatum (arrow indicates the anterior margin discontinuous). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244802/files/figure.png" pageId="37" pageNumber="488">Fig. 56</figureCitation>
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.
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</paragraph>
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<paragraph id="8BC836C38517FFA459C1F8CC5115F899" blockId="37.[151,1437,871,1869]" box="[199,1114,1844,1869]" pageId="37" pageNumber="488">We noted a single female with lateral margins of elytra distinctly curved outward.</paragraph>
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</subSubSection>
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</treatment>
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</document> |