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<document id="64C341578065EFC12F303FF2EB380418" ID-DOI="10.11646/zootaxa.3722.4.1" ID-GBIF-Dataset="c092e547-503a-41ef-a855-4a12b5d776c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="223434" ID-ZooBank="3A461275-7565-4F6E-9785-12C45E12992E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460692161620" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2013" docId="03DE87B0FFCDFFC7CADAB7CFFBD919C9" docLanguage="en" docName="zt03722p438.pdf" docOrigin="Zootaxa 3722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaconothrus dispela Colloff, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="413" masterDocId="FFE7FFC8FFC4FFCBCA4DB328FF871A47" masterDocTitle="Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region" masterLastPageNumber="438" masterPageNumber="401" pageNumber="410" updateTime="1698351897157" updateUser="plazi">
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<mods:title id="13D2BE1CCA6B80A8386E6A4786A9D8D6">Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region</mods:title>
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<mods:name id="036F361A56AF0EF3F9AF1FA4BBF64B36" type="personal">
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<mods:namePart id="F24E6C0C3574668A79904B5A55AF8688">Colloff, Matthew J.</mods:namePart>
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<mods:date id="B709E9F0814022FEA400B6F603A7C0AC">2013</mods:date>
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<treatment id="03DE87B0FFCDFFC7CADAB7CFFBD919C9" ID-DOI="http://doi.org/10.5281/zenodo.6157852" ID-GBIF-Taxon="119580480" ID-Zenodo-Dep="6157852" LSID="urn:lsid:plazi:treatment:03DE87B0FFCDFFC7CADAB7CFFBD919C9" httpUri="http://treatment.plazi.org/id/03DE87B0FFCDFFC7CADAB7CFFBD919C9" lastPageId="12" lastPageNumber="413" pageId="9" pageNumber="410">
<subSubSection id="C36D652DFFCDFFC2CADAB7CFFCE21FC9" pageId="9" pageNumber="410" type="nomenclature">
<paragraph id="8BC836A6FFCDFFC2CADAB7CFFD961F46" blockId="9.[151,529,1255,1314]" box="[151,529,1255,1281]" pageId="9" pageNumber="410">
<heading id="D08081CAFFCDFFC2CADAB7CFFD961F46" bold="true" box="[151,529,1255,1281]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="B903EAB4FFCDFFC2CADAB7CFFD961F46" bold="true" box="[151,529,1255,1281]" pageId="9" pageNumber="410">
<taxonomicName id="4C774D25FFCDFFC2CADAB7CFFE2A1F46" ID-CoL="7VRCN" box="[151,429,1255,1281]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="dispela" status="sp. nov.">
<emphasis id="B903EAB4FFCDFFC2CADAB7CFFE2A1F46" bold="true" box="[151,429,1255,1281]" italics="true" pageId="9" pageNumber="410">Malaconothrus dispela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFCDFFC2CBF9B7CFFD961F46" box="[436,529,1255,1281]" pageId="9" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFCDFFC2CADAB621FE951F65" blockId="9.[151,529,1255,1314]" box="[151,274,1289,1314]" pageId="9" pageNumber="410">(Figs. 6, 7)</paragraph>
<paragraph id="8BC836A6FFCDFFC2CADAB679FCE21FC9" blockId="9.[151,1437,1361,2013]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CADAB679FEA51F2D" bold="true" box="[151,290,1361,1386]" pageId="9" pageNumber="410">Dimensions</emphasis>
.
<typeStatus id="54CC8804FFCDFFC2CB7FB679FE1D1F2D" box="[306,410,1361,1386]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
female length 368 Μm, breadth 167 Μm.
<typeStatus id="54CC8804FFCDFFC2C931B67AFC591F2D" box="[892,990,1362,1386]" pageId="9" pageNumber="410" type="paratype">Paratype</typeStatus>
female lengths 381, 366 Μm, breadths 164, 170 Μm. Ratio of prodorsum to total length: 0.4 (
<typeStatus id="54CC8804FFCDFFC2C8BCB65DFCDE1FC9" box="[753,857,1397,1422]" pageId="9" pageNumber="410" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFCDFFC1CA8AB6B1FEB31DDA" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="description">
<paragraph id="8BC836A6FFCDFFC2CA8AB6B1FBFB1C59" blockId="9.[151,1437,1361,2013]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB6B1FE991FF5" bold="true" box="[199,286,1433,1458]" pageId="9" pageNumber="410">Female</emphasis>
.
<emphasis id="B903EAB4FFCDFFC2CB60B6B1FE2D1FF5" box="[301,426,1433,1458]" italics="true" pageId="9" pageNumber="410">Prodorsum</emphasis>
. Rostrum acute, U-shaped (Fig. 6a). Rostral seta (
<emphasis id="B903EAB4FFCDFFC2C9A5B6B3FC781FF5" box="[1000,1023,1435,1458]" italics="true" pageId="9" pageNumber="410">ro</emphasis>
) thin, 4 Μm long, smooth, emerging from alveolus ventral to apex of carina. Lamellar seta (
<emphasis id="B903EAB4FFCDFFC2C967B695FCB91F91" box="[810,830,1469,1494]" italics="true" pageId="9" pageNumber="410">le</emphasis>
) smooth, sub-equal in length to
<emphasis id="B903EAB4FFCDFFC2CE8BB697FB5A1F91" box="[1222,1245,1471,1494]" italics="true" pageId="9" pageNumber="410">ro</emphasis>
, emerging from alveolus medial of carina. Interlamellar seta (
<emphasis id="B903EAB4FFCDFFC2C889B6CBFD5D1FBD" box="[708,730,1507,1530]" italics="true" pageId="9" pageNumber="410">in</emphasis>
) smooth, setiform, 22 Μm long, 4 × longer than smooth exobothridial seta
<emphasis id="B903EAB4FFCDFFC2CB28B52FFEFA1C59" box="[357,381,1543,1566]" italics="true" pageId="9" pageNumber="410">ex</emphasis>
. Prodorsum finely porose; cerotegument of small, angular foveolae.
</paragraph>
<paragraph id="8BC836A6FFCDFFC2CA8AB503FCEB1C9F" blockId="9.[151,1437,1361,2013]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB503FEC51C05" box="[199,322,1579,1602]" italics="true" pageId="9" pageNumber="410">Notogaster</emphasis>
. Anterior notogastral margin slightly convex; humeral region rounded (Fig. 6a). With 15 pairs of short, smooth, setiform setae. Setae mostly sub-equal in length (2027 Μm),
<emphasis id="B903EAB4FFCDFFC2CE48B565FB941C21" box="[1029,1043,1613,1638]" italics="true" pageId="9" pageNumber="410">d</emphasis>
1,
<emphasis id="B903EAB4FFCDFFC2CE66B567FBB01C21" box="[1067,1079,1615,1638]" italics="true" pageId="9" pageNumber="410">e</emphasis>
1,
<emphasis id="B903EAB4FFCDFFC2CE02B565FBDA1C21" box="[1103,1117,1613,1638]" italics="true" pageId="9" pageNumber="410">h</emphasis>
2 and
<emphasis id="B903EAB4FFCDFFC2CED2B565FB2A1C21" box="[1183,1197,1613,1638]" italics="true" pageId="9" pageNumber="410">h</emphasis>
3 longer (3746 Μm). Seta
<emphasis id="B903EAB4FFCDFFC2CA83B55EFF5D1CCA" box="[206,218,1654,1677]" italics="true" pageId="9" pageNumber="410">c</emphasis>
1 same distance from
<emphasis id="B903EAB4FFCDFFC2CB80B55EFE5E1CCA" box="[461,473,1654,1677]" italics="true" pageId="9" pageNumber="410">c</emphasis>
2 as from
<emphasis id="B903EAB4FFCDFFC2C808B55EFDD61CCA" box="[581,593,1654,1677]" italics="true" pageId="9" pageNumber="410">c</emphasis>
2 to
<emphasis id="B903EAB4FFCDFFC2C833B55EFD0D1CCA" box="[638,650,1654,1677]" italics="true" pageId="9" pageNumber="410">c</emphasis>
3. Notogaster shield-shaped, margins more-or-less parallel, broadest at level of
<emphasis id="B903EAB4FFCDFFC2CAB9B5B4FE891CF4" box="[244,270,1692,1715]" italics="true" pageId="9" pageNumber="410">cp</emphasis>
. Without notogastral ridges. Ratio of length to breadth 1.3. Caudal margin slightly indented at level of
<emphasis id="B903EAB4FFCDFFC2CADAB597FF221C9F" box="[151,165,1727,1752]" italics="true" pageId="9" pageNumber="410">h</emphasis>
2; V-shaped. Cerotegument of small, angular foveolae (Fig. 6d).
</paragraph>
<paragraph id="8BC836A6FFCDFFC2CA8AB5CFFE721D2D" blockId="9.[151,1437,1361,2013]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB5CFFED11CB9" box="[199,342,1767,1790]" italics="true" pageId="9" pageNumber="410">Coxisternum</emphasis>
. Epimeral setal formula 3-1-3-3 (Fig. 6b). Seta
<emphasis id="B903EAB4FFCDFFC2C93EB5CEFC081CB8" box="[883,911,1766,1791]" italics="true" pageId="9" pageNumber="410">3b</emphasis>
and
<emphasis id="B903EAB4FFCDFFC2C98AB5CFFC641CB8" box="[967,995,1767,1791]" italics="true" pageId="9" pageNumber="410">4a</emphasis>
longest (2124 Μm), much longer than other epimeral setae. Epimeral plates I and III fused anteriorly; II fused posteriorly; IV separated. Epimeral plates I with prominent spur anterolaterally. Epimeral plates II acute posterolaterally. Epimeres III and IV of equal width. Apodeme IV strongly concave.
</paragraph>
<paragraph id="8BC836A6FFCDFFC1CA8AB45EFF521ABF" blockId="9.[151,1437,1361,2013]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB45EFE1D1DC9" box="[199,410,1910,1935]" italics="true" pageId="9" pageNumber="410">Anogenital Region</emphasis>
. With six pairs of genital setae;
<emphasis id="B903EAB4FFCDFFC2C95CB45FFC981DC9" box="[785,799,1911,1934]" italics="true" pageId="9" pageNumber="410">g</emphasis>
1-5 barbed (Figs 6b,c);
<emphasis id="B903EAB4FFCDFFC2CE50B45FFBAC1DC9" box="[1053,1067,1911,1934]" italics="true" pageId="9" pageNumber="410">g</emphasis>
6 spiniform, smooth, longer than others (26 Μm), positioned three times distance from
<emphasis id="B903EAB4FFCDFFC2C8B9B4B6FC851DF2" box="[756,770,1950,1973]" italics="true" pageId="9" pageNumber="410">g</emphasis>
5 than from
<emphasis id="B903EAB4FFCDFFC2C9C7B4B6FC1F1DF2" box="[906,920,1950,1973]" italics="true" pageId="9" pageNumber="410">g</emphasis>
5 to
<emphasis id="B903EAB4FFCDFFC2C98AB4B6FC521DF2" box="[967,981,1950,1973]" italics="true" pageId="9" pageNumber="410">g</emphasis>
4. Each genital plate 77 Μm long, 29 Μm broad, ovoid, elongated; posterior margin transverse. One pair of anal setae 6 Μm long. Each adanal plate 70 Μm long, 11 Μm broad, with three pairs of smooth, setiform adanal setae, 1316 Μm long, shorter than width of adanal plate. Anterior margin of adanal plate without thickened transverse ridge, overlapped by posterior margin of genital plate.
</paragraph>
<paragraph id="8BC836A6FFCEFFC1CADAB536FE181C12" blockId="10.[151,1436,1566,1621]" pageId="10" pageNumber="411">
<emphasis id="B903EAB4FFCEFFC1CADAB536FE921C74" bold="true" box="[151,277,1566,1588]" pageId="10" pageNumber="411">FIGURE 6.</emphasis>
<taxonomicName id="4C774D25FFCEFFC1CB53B536FD8D1C73" box="[286,522,1566,1588]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="dispela" status="sp. nov.">
<emphasis id="B903EAB4FFCEFFC1CB53B536FD8D1C73" box="[286,522,1566,1588]" italics="true" pageId="10" pageNumber="411">Malaconothrus dispela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFCEFFC1C859B537FDE11C73" box="[532,614,1567,1588]" pageId="10" pageNumber="411" rank="species">
<emphasis id="B903EAB4FFCEFFC1C859B537FDE61C73" bold="true" box="[532,609,1567,1588]" pageId="10" pageNumber="411">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<typeStatus id="54CC8804FFCEFFC1C83BB536FD481C73" box="[630,719,1566,1588]" pageId="10" pageNumber="411" type="holotype">holotype</typeStatus>
female; a) dorsal; b) ventral; c) detail of genital seta
<emphasis id="B903EAB4FFCEFFC1CF49B537FA971C73" box="[1284,1296,1567,1588]" italics="true" pageId="10" pageNumber="411">g</emphasis>
5; d) detail of notogastral cerotegument.
</paragraph>
<paragraph id="8BC836A6FFCEFFC1CA8AB5ABFEB31DDA" blockId="10.[151,1436,1666,2021]" pageId="10" pageNumber="411">
<emphasis id="B903EAB4FFCEFFC1CA8AB5ABFF7B1CDD" box="[199,252,1667,1690]" italics="true" pageId="10" pageNumber="411">Legs</emphasis>
. Pre-tarsi monodactylous (
<figureCitation id="134C2A23FFCEFFC1C87DB5AAFDF11CDC" box="[560,630,1666,1691]" captionStart="FIGURE 7" captionStartId="11.[151,250,1970,1992]" captionTargetBox="[183,1403,257,1949]" captionTargetId="figure@11.[177,1413,250,1960]" captionTargetPageId="11" captionText="FIGURE 7. Malaconothrus dispela sp. nov., left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="10" pageNumber="411">Fig. 7</figureCitation>
). Legs short and broad: Leg I 104 Μm long; tarsi sub-rectangular with prominent apophysis of setae
<emphasis id="B903EAB4FFCEFFC1CBA5B58EFE7F1CF9" box="[488,504,1702,1726]" italics="true" pageId="10" pageNumber="411">ft</emphasis>
”. Ratio of length to breadth: I 1.7; II 1.7; III 2.1; IV 2.6. Setal formula: Leg
<date id="FFC91066FFCEFFC1CF11B58FFA151CF8" box="[1372,1426,1703,1727]" pageId="10" pageNumber="411">I 1-4</date>
- 2(1)-4(1)-11(3); Leg
<date id="FFC91066FFCEFFC1CBCEB5E3FE471CA4" box="[387,448,1739,1763]" pageId="10" pageNumber="411">II 1-3</date>
-1(1)-2(1)-10(1); Leg
<date id="FFC91066FFCEFFC1C8F5B5E3FC831CA4" box="[696,772,1739,1763]" pageId="10" pageNumber="411">III 2-2</date>
-1-2(1)-10; Leg
<date id="FFC91066FFCEFFC1C9F9B5E3FC791CA4" box="[948,1022,1739,1763]" pageId="10" pageNumber="411">IV 1-2</date>
-1-2-10. Leg setae smooth. Proral (
<emphasis id="B903EAB4FFCEFFC1CFC9B5E3FA151CA5" box="[1412,1426,1739,1762]" italics="true" pageId="10" pageNumber="411">p</emphasis>
) and unguinal setae (
<emphasis id="B903EAB4FFCEFFC1CBCBB5C7FE131D41" box="[390,404,1775,1798]" italics="true" pageId="10" pageNumber="411">u</emphasis>
) on all tarsi short, broad, peg-like, blunt apically (
<figureCitation id="134C2A23FFCEFFC1C9B8B5C6FBB91D40" box="[1013,1086,1774,1799]" captionStart="FIGURE 7" captionStartId="11.[151,250,1970,1992]" captionTargetBox="[183,1403,257,1949]" captionTargetId="figure@11.[177,1413,250,1960]" captionTargetPageId="11" captionText="FIGURE 7. Malaconothrus dispela sp. nov., left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="10" pageNumber="411">Fig. 7</figureCitation>
). Tarsus I with solenidion ω1 posterodorsal of ω2-3; famulus (ε) short (3 Μm), spine like; fastigial setae (
<emphasis id="B903EAB4FFCEFFC1C984B43DFC5F1D6A" box="[969,984,1813,1837]" italics="true" pageId="10" pageNumber="411">ft</emphasis>
) heteromorphic:
<emphasis id="B903EAB4FFCEFFC1CED7B43DFB2E1D6A" box="[1178,1193,1813,1837]" italics="true" pageId="10" pageNumber="411">ft</emphasis>
” short, blunt, curved, very broad, emerging from prominent conical apophysis (
<figureCitation id="134C2A23FFCEFFC1C950B414FCE61D13" box="[797,865,1852,1877]" captionStart="FIGURE 7" captionStartId="11.[151,250,1970,1992]" captionTargetBox="[183,1403,257,1949]" captionTargetId="figure@11.[177,1413,250,1960]" captionTargetPageId="11" captionText="FIGURE 7. Malaconothrus dispela sp. nov., left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="10" pageNumber="411">Fig. 7</figureCitation>
a);
<emphasis id="B903EAB4FFCEFFC1C9C8B414FC131D13" box="[901,916,1852,1876]" italics="true" pageId="10" pageNumber="411">ft</emphasis>
long, broad, curved, pointed. Tarsi IIIV and II with
<emphasis id="B903EAB4FFCEFFC1CAA6B448FF7D1D3F" box="[235,250,1888,1912]" italics="true" pageId="10" pageNumber="411">ft</emphasis>
similarly heteromorphic (
<figureCitation id="134C2A23FFCEFFC1C86FB477FDE11D3F" box="[546,614,1887,1912]" captionStart="FIGURE 7" captionStartId="11.[151,250,1970,1992]" captionTargetBox="[183,1403,257,1949]" captionTargetId="figure@11.[177,1413,250,1960]" captionTargetPageId="11" captionText="FIGURE 7. Malaconothrus dispela sp. nov., left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="10" pageNumber="411">Fig. 7</figureCitation>
b,c,d). Tarsus IV with seta
<emphasis id="B903EAB4FFCEFFC1C9DAB449FC251D3F" box="[919,930,1889,1912]" italics="true" pageId="10" pageNumber="411">s</emphasis>
same length and shape as primiventral setae (
<emphasis id="B903EAB4FFCEFFC1CAEDB4ADFF3C1DDB" box="[160,187,1925,1948]" italics="true" pageId="10" pageNumber="411">pv</emphasis>
) (
<figureCitation id="134C2A23FFCEFFC1CA9EB4ACFE901DDA" box="[211,279,1924,1949]" captionStart="FIGURE 7" captionStartId="11.[151,250,1970,1992]" captionTargetBox="[183,1403,257,1949]" captionTargetId="figure@11.[177,1413,250,1960]" captionTargetPageId="11" captionText="FIGURE 7. Malaconothrus dispela sp. nov., left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV." httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="10" pageNumber="411">Fig. 7</figureCitation>
d).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFCEFFC0CA8AB48FFEF21A92" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" type="materials_examined">
<paragraph id="8BC836A6FFCEFFC0CA8AB48FFEF21A92" blockId="10.[151,1436,1666,2021]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
<emphasis id="B903EAB4FFCEFFC1CA8AB48FFE331D87" bold="true" box="[199,436,1959,1984]" pageId="10" pageNumber="411">Material Examined</emphasis>
.
<typeStatus id="54CC8804FFCEFFC1CB8AB48FFDA81D87" box="[455,559,1959,1984]" pageId="10" pageNumber="411" type="holotype">Holotype</typeStatus>
and two
<typeStatus id="54CC8804FFCEFFC1C8E8B480FC941D87" box="[677,787,1960,1984]" pageId="10" pageNumber="411" type="paratype">paratypes</typeStatus>
, ANIC 418, litter, rainforest, Highlands Highway, near Komum (ca.
<quantity id="4C8F9B43FFCEFFC1CB7CB4E4FEFB1DA2" box="[305,380,1996,2021]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="411" unit="km" value="15.0">15 km</quantity>
east of Mt. Hagen),
<collectingCountry id="F3607636FFCEFFC1C838B4E5FCDF1DA2" box="[629,856,1996,2021]" name="Papua New Guinea" pageId="10" pageNumber="411">Papua New Guinea</collectingCountry>
,
<geoCoordinate id="EE435061FFCEFFC1C92AB4E4FC5F1DA2" box="[871,984,1996,2021]" direction="south" orientation="latitude" pageId="10" pageNumber="411" precision="15" value="-5.828611">5°49'43&quot;S</geoCoordinate>
<geoCoordinate id="EE435061FFCEFFC1C9AEB4E4FBF31DA3" box="[995,1140,1996,2020]" direction="east" orientation="longitude" pageId="10" pageNumber="411" precision="15" value="144.37195">144°22'19&quot;E</geoCoordinate>
,
<quantity id="4C8F9B43FFCEFFC1CECEB4E4FB661DA3" box="[1155,1249,1996,2020]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="10" pageNumber="411" unit="m" value="1830.0">1830 m</quantity>
., coll. G.
<collectingCountry id="F3607636FFCEFFC1CF1EB4E4FA1F1DA2" box="[1363,1432,1996,2021]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="411">Baker</collectingCountry>
,
<date id="FFC91066FFCFFFC0CADAB3BFFE9D1AF7" box="[151,282,151,176]" pageId="11" pageNumber="412" value="1972-07-16">16.vii.1972</date>
.
<typeStatus id="54CC8804FFCFFFC0CB65B3BFFE171AF7" box="[296,400,151,176]" pageId="11" pageNumber="412" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54CC8804FFCFFFC0CB83B3B0FDBE1AF7" box="[462,569,152,176]" pageId="11" pageNumber="412" type="paratype">paratypes</typeStatus>
deposited in the Australian National Insect Collection, CSIRO Ecosystem Sciences, Canberra.
</paragraph>
</subSubSection>
<caption id="DF08662EFFCFFFC0CADAB49AFB781D8F" box="[151,1279,1970,1992]" httpUri="https://zenodo.org/record/223440/files/figure.png" pageId="11" pageNumber="412" targetBox="[183,1403,257,1949]" targetPageId="11">
<paragraph id="8BC836A6FFCFFFC0CADAB49AFB781D8F" blockId="11.[151,1279,1970,1992]" box="[151,1279,1970,1992]" pageId="11" pageNumber="412">
<emphasis id="B903EAB4FFCFFFC0CADAB49AFE941D8F" bold="true" box="[151,275,1970,1992]" pageId="11" pageNumber="412">FIGURE 7.</emphasis>
<taxonomicName id="4C774D25FFCFFFC0CB54B49AFD841D8F" box="[281,515,1970,1992]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="dispela" status="sp. nov.">
<emphasis id="B903EAB4FFCFFFC0CB54B49AFD841D8F" box="[281,515,1970,1992]" italics="true" pageId="11" pageNumber="412">Malaconothrus dispela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFCFFFC0C844B49BFDDD1D8F" box="[521,602,1971,1992]" pageId="11" pageNumber="412" rank="species">
<emphasis id="B903EAB4FFCFFFC0C844B49BFDD31D8F" bold="true" box="[521,596,1971,1992]" pageId="11" pageNumber="412">sp. nov</emphasis>
.
</taxonomicNameLabel>
, left legs, left-hand view; a) Leg I; b) Leg II; c) Leg III; d) Leg IV.
</paragraph>
</caption>
<subSubSection id="C36D652DFFC8FFC7CA8AB3BFFDCA1A92" pageId="12" pageNumber="413" type="etymology">
<paragraph id="8BC836A6FFC8FFC7CA8AB3BFFDCA1A92" blockId="12.[151,1436,151,910]" pageId="12" pageNumber="413">
<emphasis id="B903EAB4FFC8FFC7CA8AB3BFFEC91AF7" bold="true" box="[199,334,151,176]" pageId="12" pageNumber="413">Etymology.</emphasis>
The specific name
<taxonomicName id="4C774D25FFC8FFC7C86AB3BFFDFF1AF7" box="[551,632,151,176]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7C86AB3BFFDFF1AF7" box="[551,632,151,176]" italics="true" pageId="12" pageNumber="413">dispela</emphasis>
</taxonomicName>
is Tok Pisin (the official language of
<collectingCountry id="F3607636FFC8FFC7CE51B3B0FB721AF7" box="[1052,1269,151,176]" name="Papua New Guinea" pageId="12" pageNumber="413">Papua New Guinea</collectingCountry>
) for this one, derived from the English, this fellow.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC8FFC7CA8AB3F7FBD919C9" pageId="12" pageNumber="413" type="discussion">
<paragraph id="8BC836A6FFC8FFC7CA8AB3F7FDF51BF6" blockId="12.[151,1436,151,910]" pageId="12" pageNumber="413">
<emphasis id="B903EAB4FFC8FFC7CA8AB3F7FEB31ABF" bold="true" box="[199,308,223,248]" pageId="12" pageNumber="413">Remarks</emphasis>
.
<taxonomicName id="4C774D25FFC8FFC7CB04B3F7FDD31ABF" box="[329,596,223,248]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela" status="sp. nov.">
<emphasis id="B903EAB4FFC8FFC7CB04B3F7FDD31ABF" box="[329,596,223,248]" italics="true" pageId="12" pageNumber="413">Malaconothrus dispela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC8FFC7C82EB3F7FD441AB0" box="[611,707,223,248]" pageId="12" pageNumber="413" rank="species">
<emphasis id="B903EAB4FFC8FFC7C82EB3F7FD3A1ABF" bold="true" box="[611,701,223,248]" pageId="12" pageNumber="413">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from other members of the genus by the following combination of characters: 1) the shield-shaped notogaster with the caudal region V shaped; 2) with notogastral setae
<emphasis id="B903EAB4FFC8FFC7CA9AB20FFF621B07" box="[215,229,295,320]" italics="true" pageId="12" pageNumber="413">d</emphasis>
1,
<emphasis id="B903EAB4FFC8FFC7CAB1B201FE8F1B07" box="[252,264,297,320]" italics="true" pageId="12" pageNumber="413">e</emphasis>
1 and
<emphasis id="B903EAB4FFC8FFC7CB07B20FFEDF1B07" box="[330,344,295,320]" italics="true" pageId="12" pageNumber="413">h</emphasis>
2 longer than others; 3) notogaster without posterior or lateral ridges; 4) epimeral plates II acute posterolaterally; 5) the epimeral setae
<emphasis id="B903EAB4FFC8FFC7C805B266FDE31B20" box="[584,612,334,359]" italics="true" pageId="12" pageNumber="413">3b</emphasis>
and
<emphasis id="B903EAB4FFC8FFC7C8D1B267FD311B20" box="[668,694,335,359]" italics="true" pageId="12" pageNumber="413">4c</emphasis>
longer than others; 6) the six pairs of genital setae; 7) the genital setae are heteromorphic:
<emphasis id="B903EAB4FFC8FFC7CBF5B25CFE411BCC" box="[440,454,372,395]" italics="true" pageId="12" pageNumber="413">g</emphasis>
5 is spiniform and smooth, the others are setiform and barbed; 8) seta
<emphasis id="B903EAB4FFC8FFC7CEA1B25CFB7D1BCC" box="[1260,1274,372,395]" italics="true" pageId="12" pageNumber="413">g</emphasis>
5 spaced some distance from the next most posterior seta.
</paragraph>
<paragraph id="8BC836A6FFC8FFC7CA8AB295FBD919C9" blockId="12.[151,1436,151,910]" pageId="12" pageNumber="413">
<taxonomicName id="4C774D25FFC8FFC7CA8AB295FE4E1B91" box="[199,457,445,470]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela" status="sp. nov.">
<emphasis id="B903EAB4FFC8FFC7CA8AB295FE4E1B91" box="[199,457,445,470]" italics="true" pageId="12" pageNumber="413">Malaconothrus dispela</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC8FFC7CB82B295FDAF1B92" box="[463,552,445,470]" pageId="12" pageNumber="413" rank="species">
<emphasis id="B903EAB4FFC8FFC7CB82B295FDA61B91" bold="true" box="[463,545,445,470]" pageId="12" pageNumber="413">sp. nov</emphasis>
.
</taxonomicNameLabel>
is morphologically most similar to
<taxonomicName id="4C774D25FFC8FFC7C9F9B296FB571B91" authority="Aoki, 1969" authorityName="Aoki" authorityYear="1969" box="[948,1232,445,470]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B903EAB4FFC8FFC7C9F9B296FBCE1B92" box="[948,1097,446,469]" italics="true" pageId="12" pageNumber="413">M. pygmaeus</emphasis>
Aoki, 1969
</taxonomicName>
,
<taxonomicName id="4C774D25FFC8FFC7CE97B296FEAD1BBD" authority="Subias, 2004" authorityName="Subias" authorityYear="2004" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="neonominatus">
<emphasis id="B903EAB4FFC8FFC7CE97B296FA1B1B92" box="[1242,1436,446,469]" italics="true" pageId="12" pageNumber="413">M. neonominatus</emphasis>
Subías, 2004
</taxonomicName>
(=
<taxonomicName id="4C774D25FFC8FFC7CB00B2CAFD061BBD" authority="Hammer, 1961" authorityName="Hammer" authorityYear="1961" box="[333,641,481,506]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B903EAB4FFC8FFC7CB00B2CAFE4C1BBD" box="[333,459,481,506]" italics="true" pageId="12" pageNumber="413">M. pulcher</emphasis>
Hammer, 1961
</taxonomicName>
),
<taxonomicName id="4C774D25FFC8FFC7C8D5B2CAFC541BBD" authority="Fujikawa, 2005" authorityName="Fujikawa" authorityYear="2005" box="[664,979,481,506]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B903EAB4FFC8FFC7C8D5B2CAFC9E1BBE" box="[664,793,482,505]" italics="true" pageId="12" pageNumber="413">M. minutus</emphasis>
Fujikawa, 2005
</taxonomicName>
and
<taxonomicName id="4C774D25FFC8FFC7CE42B2CAFACB1BBD" authority="Fujikawa, 2005" authorityName="Fujikawa" authorityYear="2005" box="[1039,1356,481,506]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="setoumi">
<emphasis id="B903EAB4FFC8FFC7CE42B2CAFB091BBE" box="[1039,1166,482,505]" italics="true" pageId="12" pageNumber="413">M. setoumi</emphasis>
Fujikawa, 2005
</taxonomicName>
. These species are members of the
<taxonomicName id="4C774D25FFC8FFC7CB83B12DFDE91859" box="[462,622,517,542]" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="430" phylum="Arthropoda" rank="species" species="monodactylus">Monodactylus</taxonomicName>
species group (cf. below). All the species mentioned above have barbed genital setae, but the latter two species have five pairs all of which are barbed;
<taxonomicName id="4C774D25FFC8FFC7CE54B102FB5B1806" box="[1049,1244,554,577]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="neonominatus">
<emphasis id="B903EAB4FFC8FFC7CE54B102FB5B1806" box="[1049,1244,554,577]" italics="true" pageId="12" pageNumber="413">M. neonominatus</emphasis>
</taxonomicName>
has six pairs, all barbed, but much longer marginal notogastral setae, especially
<emphasis id="B903EAB4FFC8FFC7C912B165FCEA1821" box="[863,877,589,614]" italics="true" pageId="12" pageNumber="413">h</emphasis>
2, than
<taxonomicName id="4C774D25FFC8FFC7C9F6B166FBB51821" box="[955,1074,589,614]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7C9F6B166FBB51821" box="[955,1074,589,614]" italics="true" pageId="12" pageNumber="413">M. dispela</emphasis>
</taxonomicName>
.
<taxonomicName id="4C774D25FFC8FFC7CE0CB165FAE41822" box="[1089,1379,589,614]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B903EAB4FFC8FFC7CE0CB165FAE41822" box="[1089,1379,589,614]" italics="true" pageId="12" pageNumber="413">Malaconothrus pygmaeus</emphasis>
</taxonomicName>
, like
<taxonomicName id="4C774D25FFC8FFC7CADAB15DFE9718CA" box="[151,272,628,653]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7CADAB15DFE9718CA" box="[151,272,628,653]" italics="true" pageId="12" pageNumber="413">M. dispela</emphasis>
</taxonomicName>
, has six pairs, but
<emphasis id="B903EAB4FFC8FFC7CBBDB15DFE7918CB" box="[496,510,629,652]" italics="true" pageId="12" pageNumber="413">g</emphasis>
1-4 are barbed and
<emphasis id="B903EAB4FFC8FFC7C89CB15DFD5818CB" box="[721,735,629,652]" italics="true" pageId="12" pageNumber="413">g</emphasis>
5-6 are smooth according to the supplementary description by Yamamoto (1998), whereas in
<emphasis id="B903EAB4FFC8FFC7CBA0B1B4FDF018F4" box="[493,631,667,692]" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="4C774D25FFC8FFC7CBA0B1B4FDE518F3" box="[493,610,667,692]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">M. dispela</taxonomicName>
g
</emphasis>
1-5 are barbed and
<emphasis id="B903EAB4FFC8FFC7C976B1B4FCCE18F4" box="[827,841,668,691]" italics="true" pageId="12" pageNumber="413">g</emphasis>
6 is smooth.
<taxonomicName id="4C774D25FFC8FFC7C99DB1B3FB7618F4" box="[976,1265,667,692]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B903EAB4FFC8FFC7C99DB1B3FB7618F4" box="[976,1265,667,692]" italics="true" pageId="12" pageNumber="413">Malaconothrus pygmaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D25FFC8FFC7CF6BB1B4FA1B18F3" box="[1318,1436,667,692]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7CF6BB1B4FA1B18F3" box="[1318,1436,667,692]" italics="true" pageId="12" pageNumber="413">M. dispela</emphasis>
</taxonomicName>
have the epimeral setal formula 3-1-3-3 (seta
<emphasis id="B903EAB4FFC8FFC7C8EDB1EAFD3B189C" box="[672,700,706,731]" italics="true" pageId="12" pageNumber="413">3b</emphasis>
is present), whereas in
<taxonomicName id="4C774D25FFC8FFC7C981B1EBFB08189D" box="[972,1167,707,730]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="neonominatus">
<emphasis id="B903EAB4FFC8FFC7C981B1EBFB08189D" box="[972,1167,707,730]" italics="true" pageId="12" pageNumber="413">M. neonominatus</emphasis>
</taxonomicName>
it is 3-1-2-3 (seta
<emphasis id="B903EAB4FFC8FFC7CF28B1EAFA06189C" box="[1381,1409,706,731]" italics="true" pageId="12" pageNumber="413">3b</emphasis>
is absent). In
<taxonomicName id="4C774D25FFC8FFC7CB5BB1CFFE2B18B9" box="[278,428,743,766]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B903EAB4FFC8FFC7CB5BB1CFFE2B18B9" box="[278,428,743,766]" italics="true" pageId="12" pageNumber="413">M. pygmaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D25FFC8FFC7CBA8B1CFFD2F18B9" box="[485,680,743,766]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="neonominatus">
<emphasis id="B903EAB4FFC8FFC7CBA8B1CFFD2F18B9" box="[485,680,743,766]" italics="true" pageId="12" pageNumber="413">M. neonominatus</emphasis>
</taxonomicName>
setae
<emphasis id="B903EAB4FFC8FFC7C8BDB1CEFC8C18B9" box="[752,779,742,766]" italics="true" pageId="12" pageNumber="413">4a</emphasis>
,
<emphasis id="B903EAB4FFC8FFC7C951B1CEFCB118B9" box="[796,822,742,766]" italics="true" pageId="12" pageNumber="413">4c</emphasis>
and
<emphasis id="B903EAB4FFC8FFC7C923B1CEFC0E18B9" box="[878,905,742,766]" italics="true" pageId="12" pageNumber="413">3c</emphasis>
are sub-equal in length, whereas in
<taxonomicName id="4C774D25FFC8FFC7CF69B1CFFA1B18B9" box="[1316,1436,741,766]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7CF69B1CFFA1B18B9" box="[1316,1436,741,766]" italics="true" pageId="12" pageNumber="413">M. dispela</emphasis>
</taxonomicName>
setae
<emphasis id="B903EAB4FFC8FFC7CA9BB022FF751964" box="[214,242,778,803]" italics="true" pageId="12" pageNumber="413">3b</emphasis>
and
<emphasis id="B903EAB4FFC8FFC7CB67B022FEC31965" box="[298,324,778,802]" italics="true" pageId="12" pageNumber="413">4c</emphasis>
are each about twice the length of the other two setae. Finally, the notogastral cerotegument of
<taxonomicName id="4C774D25FFC8FFC7CF30B023FEB31901" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="neonominatus">
<emphasis id="B903EAB4FFC8FFC7CF30B023FEB31901" italics="true" pageId="12" pageNumber="413">M. neonominatus</emphasis>
</taxonomicName>
is of closely-spaced ovoid foveolae of similar size, whereas in
<taxonomicName id="4C774D25FFC8FFC7CE7BB007FB561901" box="[1078,1233,815,838]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B903EAB4FFC8FFC7CE7BB007FB561901" box="[1078,1233,815,838]" italics="true" pageId="12" pageNumber="413">M. pygmaeus</emphasis>
</taxonomicName>
the foveolae are broadly-separated ovoids greatly varying in size and in
<taxonomicName id="4C774D25FFC8FFC7C962B07BFC2C192C" box="[815,939,850,875]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="dispela">
<emphasis id="B903EAB4FFC8FFC7C962B07BFC2C192C" box="[815,939,850,875]" italics="true" pageId="12" pageNumber="413">M. dispela</emphasis>
</taxonomicName>
they are of equal size, sub-pentagonal or hexagonal and regularly and closely spaced with clearly-defined ridges between them.
</paragraph>
</subSubSection>
</treatment>
</document>