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<document id="B2603AB161016CABEF7732E2E2324EA8" ID-DOI="10.11646/zootaxa.3722.4.1" ID-GBIF-Dataset="c092e547-503a-41ef-a855-4a12b5d776c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="223434" ID-ZooBank="3A461275-7565-4F6E-9785-12C45E12992E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460692161620" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2013" docId="03DE87B0FFC6FFC9CADAB251FDE61DF4" docLanguage="en" docName="zt03722p438.pdf" docOrigin="Zootaxa 3722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus papuensis Colloff, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="403" masterDocId="FFE7FFC8FFC4FFCBCA4DB328FF871A47" masterDocTitle="Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region" masterLastPageNumber="438" masterPageNumber="401" pageNumber="403" updateTime="1698351897157" updateUser="plazi">
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<mods:title id="929E233BBF6E7498C6B7DC85D14D610A">Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region</mods:title>
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<mods:name id="210A8956FFBC9A15687DCBC4CD728F5C" type="personal">
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<mods:namePart id="302C210C7ACBF8419DCAB7EDF59C262C">Colloff, Matthew J.</mods:namePart>
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<mods:date id="1E9B1AF72F000AE82A456736BE4C1BE0">2013</mods:date>
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<treatment id="03DE87B0FFC6FFC9CADAB251FDE61DF4" ID-DOI="http://doi.org/10.5281/zenodo.6157842" ID-GBIF-Taxon="119580494" ID-Zenodo-Dep="6157842" LSID="urn:lsid:plazi:treatment:03DE87B0FFC6FFC9CADAB251FDE61DF4" httpUri="http://treatment.plazi.org/id/03DE87B0FFC6FFC9CADAB251FDE61DF4" lastPageNumber="403" pageId="2" pageNumber="403">
<subSubSection id="C36D652DFFC6FFC9CADAB251FD4D1867" pageId="2" pageNumber="403" type="nomenclature">
<paragraph id="8BC836A6FFC6FFC9CADAB251FDB61BD4" blockId="2.[151,561,377,437]" box="[151,561,377,403]" pageId="2" pageNumber="403">
<heading id="D08081CAFFC6FFC9CADAB251FDB61BD4" bold="true" box="[151,561,377,403]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B903EAB4FFC6FFC9CADAB251FDB61BD4" bold="true" box="[151,561,377,403]" pageId="2" pageNumber="403">
<taxonomicName id="4C774D25FFC6FFC9CADAB251FE4C1BD4" ID-CoL="7X639" box="[151,459,377,403]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="papuensis" status="sp. nov.">
<emphasis id="B903EAB4FFC6FFC9CADAB251FE4C1BD4" bold="true" box="[151,459,377,403]" italics="true" pageId="2" pageNumber="403">Tyrphonothrus papuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC6FFC9CB9EB251FDB61BD4" box="[467,561,377,403]" pageId="2" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CADAB2B4FF6C1BF3" blockId="2.[151,561,377,437]" box="[151,235,412,437]" pageId="2" pageNumber="403">
(
<figureCitation id="134C2A23FFC6FFC9CAD2B2B4FF671BF3" box="[159,224,412,437]" captionStart="FIGURE 1" captionStartId="3.[151,250,1321,1343]" captionTargetBox="[162,1424,205,1294]" captionTargetId="figure@3.[150,1435,195,1296]" captionTargetPageId="3" captionText="FIGURE 1. Tyrphonothrus papuensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223435/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
)
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CADAB2CCFD4D1867" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CADAB2CCFEA51BBA" bold="true" box="[151,290,484,509]" pageId="2" pageNumber="403">Dimensions</emphasis>
.
<typeStatus id="54CC8804FFC6FFC9CB7CB2CCFE1E1BBA" box="[305,409,484,509]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
female length 602 Μm, breadth 328 Μm.
<typeStatus id="54CC8804FFC6FFC9C920B2CDFC481BBA" box="[877,975,485,509]" pageId="2" pageNumber="403" type="paratype">Paratype</typeStatus>
female length 583 Μm, breadth 317 Μm. Ratio of prodorsum to total length: 0.2 (
<typeStatus id="54CC8804FFC6FFC9C818B12FFD3A1867" box="[597,701,519,544]" pageId="2" pageNumber="403" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC6FFC9CA8AB104FDC61F2A" pageId="2" pageNumber="403" type="description">
<paragraph id="8BC836A6FFC6FFC9CA8AB104FD231893" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB104FE991802" bold="true" box="[199,286,556,581]" pageId="2" pageNumber="403">Female</emphasis>
.
<emphasis id="B903EAB4FFC6FFC9CB60B104FE2D1802" box="[301,426,556,581]" italics="true" pageId="2" pageNumber="403">Prodorsum</emphasis>
. Rostrum acute (
<figureCitation id="134C2A23FFC6FFC9C821B104FD2A1803" box="[620,685,556,581]" captionStart="FIGURE 1" captionStartId="3.[151,250,1321,1343]" captionTargetBox="[162,1424,205,1294]" captionTargetId="figure@3.[150,1435,195,1296]" captionTargetPageId="3" captionText="FIGURE 1. Tyrphonothrus papuensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223435/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
a). Rostral seta (
<emphasis id="B903EAB4FFC6FFC9C926B105FC051803" box="[875,898,557,580]" italics="true" pageId="2" pageNumber="403">ro</emphasis>
) smooth, curved, setiform, 28 Μm long, shorter than mutual distance. Lamellar seta (
<emphasis id="B903EAB4FFC6FFC9C87FB167FDC1182F" box="[562,582,591,616]" italics="true" pageId="2" pageNumber="403">le</emphasis>
) smooth, setiform, 40 Μm long, shorter than mutual distance, emerging from alveolus on median apex of incurved S-shaped lateral carina. Anterior portion of ridge projecting laterally. Interlamellar seta (
<emphasis id="B903EAB4FFC6FFC9CB3CB1B1FE0018F7" box="[369,391,665,688]" italics="true" pageId="2" pageNumber="403">in</emphasis>
) smooth, flagelliform apically, 116 Μm long, slightly longer than mutual distance, 810 × longer than smooth seta
<emphasis id="B903EAB4FFC6FFC9CBE5B195FE471893" box="[424,448,701,724]" italics="true" pageId="2" pageNumber="403">ex</emphasis>
. Prodorsum porose.
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CA8AB1C9FD7E1E47" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB1C9FEC518BF" box="[199,322,737,760]" italics="true" pageId="2" pageNumber="403">Notogaster</emphasis>
. Anterior notogastral margin slightly convex (
<figureCitation id="134C2A23FFC6FFC9C901B1F7FC0C18BF" box="[844,907,735,760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1321,1343]" captionTargetBox="[162,1424,205,1294]" captionTargetId="figure@3.[150,1435,195,1296]" captionTargetPageId="3" captionText="FIGURE 1. Tyrphonothrus papuensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223435/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
a). Humeral region obtusely angled; lyrifissure
<emphasis id="B903EAB4FFC6FFC9CADAB02DFF2A195B" box="[151,173,773,796]" italics="true" pageId="2" pageNumber="403">ia</emphasis>
19 Μm long, not associated with lateral longitudinal ridge. With 15 pairs of mostly setiform notogastral setae. Setae
<emphasis id="B903EAB4FFC6FFC9CA91B001FF6F1907" box="[220,232,809,832]" italics="true" pageId="2" pageNumber="403">e</emphasis>
2,
<emphasis id="B903EAB4FFC6FFC9CAB1B00FFE8D1907" box="[252,266,807,832]" italics="true" pageId="2" pageNumber="403">h</emphasis>
1-2 and
<emphasis id="B903EAB4FFC6FFC9CB1AB001FEE21907" box="[343,357,809,832]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2 smooth, much longer (71122 Μm long) than others (1764 Μm long), all of which are barbed except
<emphasis id="B903EAB4FFC6FFC9CAA7B066FF7F1920" box="[234,248,846,871]" italics="true" pageId="2" pageNumber="403">d</emphasis>
1. Setae
<emphasis id="B903EAB4FFC6FFC9CB1BB078FEE31920" box="[342,356,848,871]" italics="true" pageId="2" pageNumber="403">p</emphasis>
3 markedly anterior of
<emphasis id="B903EAB4FFC6FFC9C826B078FDFE1920" box="[619,633,848,871]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2;
<emphasis id="B903EAB4FFC6FFC9C8DFB078FD271920" box="[658,672,848,871]" italics="true" pageId="2" pageNumber="403">p</emphasis>
3 39 Μm long, extending almost as far as base of
<emphasis id="B903EAB4FFC6FFC9CE90B078FB6C1920" box="[1245,1259,848,871]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2 (83 Μm long). Posterior ends of median and lateral longitudinal ridges joined with transverse, shallow M-shaped ridge. Ventrally with curved ridges between
<emphasis id="B903EAB4FFC6FFC9CBABB0B2FE7319F6" box="[486,500,922,945]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2 and
<emphasis id="B903EAB4FFC6FFC9C871B0B1FDCD19F5" box="[572,586,921,946]" italics="true" pageId="2" pageNumber="403">h</emphasis>
3 and between
<emphasis id="B903EAB4FFC6FFC9C8B7B0B2FC8F19F6" box="[762,776,922,945]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2 and the caudal margin. Notogastral margins narrowing anteriorly, caudal region broadest between
<emphasis id="B903EAB4FFC6FFC9C83BB0E9FDF9199E" box="[630,638,961,985]" italics="true" pageId="2" pageNumber="403">f</emphasis>
2 and
<emphasis id="B903EAB4FFC6FFC9C8F6B0E9FD40199F" box="[699,711,961,984]" italics="true" pageId="2" pageNumber="403">e</emphasis>
2. Ratio of length to breadth 1.2. Caudal margin broadly rounded. Lyrifissurae
<emphasis id="B903EAB4FFC6FFC9CB69B0C0FEBD19B8" box="[292,314,1000,1023]" italics="true" pageId="2" pageNumber="403">ip</emphasis>
obtuse. Notogaster with large foveolae.
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CA8AB724FA1A1E2B" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB724FED11E64" box="[199,342,1036,1059]" italics="true" pageId="2" pageNumber="403">Coxisternum</emphasis>
. Epimeral setal formula 3-1-2-3; setae smooth, setiform (
<figureCitation id="134C2A23FFC6FFC9C993B723FB991E63" box="[990,1054,1035,1060]" captionStart="FIGURE 1" captionStartId="3.[151,250,1321,1343]" captionTargetBox="[162,1424,205,1294]" captionTargetId="figure@3.[150,1435,195,1296]" captionTargetPageId="3" captionText="FIGURE 1. Tyrphonothrus papuensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223435/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
b). Setae
<emphasis id="B903EAB4FFC6FFC9CEC6B723FB201E63" box="[1163,1191,1035,1060]" italics="true" pageId="2" pageNumber="403">1b</emphasis>
,
<emphasis id="B903EAB4FFC6FFC9CEFBB723FB571E64" box="[1206,1232,1035,1059]" italics="true" pageId="2" pageNumber="403">1c</emphasis>
and
<emphasis id="B903EAB4FFC6FFC9CF45B723FAA51E64" box="[1288,1314,1035,1059]" italics="true" pageId="2" pageNumber="403">3c</emphasis>
1825 Μm long; 23 × length of others. Setae
<emphasis id="B903EAB4FFC6FFC9C864B707FDC21E0F" box="[553,581,1071,1096]" italics="true" pageId="2" pageNumber="403">3b</emphasis>
absent;
<emphasis id="B903EAB4FFC6FFC9C8E9B707FD471E0F" box="[676,704,1071,1096]" italics="true" pageId="2" pageNumber="403">4b</emphasis>
in lateral position. Epimeral plates IV not fully separated; fused posteromedially; plates I with transverse median indentation. Apodeme IV straight, transverse; without blunt spine.
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CA8AB75FFD6D1F0E" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB75FFE1F1EC8" box="[199,408,1143,1168]" italics="true" pageId="2" pageNumber="403">Anogenital Region</emphasis>
. Genital plates with six pairs of setae, smooth, thin, flagelliform, unequal in length (
<emphasis id="B903EAB4FFC6FFC9CF01B750FADD1EC8" box="[1356,1370,1144,1167]" italics="true" pageId="2" pageNumber="403">g</emphasis>
<date id="FFC91066FFC6FFC9CF14B7AEFF341EF1" pageId="2" pageNumber="403" value="1917-01-24" valueMax="1917-02-24" valueMin="1917-01-24">1-2 17 24</date>
Μm long;
<emphasis id="B903EAB4FFC6FFC9CB68B7B7FEB41EF1" box="[293,307,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
<date id="FFC91066FFC6FFC9CB7EB785FEEE1EF1" box="[307,361,1182,1211]" pageId="2" pageNumber="403" value="1933-06-03">3-6 33</date>
47 Μm long;),
<emphasis id="B903EAB4FFC6FFC9C85AB7B7FDA21EF1" box="[535,549,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
1-5 spaced equidistantly,
<emphasis id="B903EAB4FFC6FFC9C962B7B7FCBA1EF1" box="[815,829,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
6 2 × distance from
<emphasis id="B903EAB4FFC6FFC9CE58B7B7FBA41EF1" box="[1045,1059,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
4 as that from
<emphasis id="B903EAB4FFC6FFC9CEF0B7B7FB4C1EF1" box="[1213,1227,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
4 from
<emphasis id="B903EAB4FFC6FFC9CF5AB7B7FAA21EF1" box="[1303,1317,1183,1206]" italics="true" pageId="2" pageNumber="403">g</emphasis>
3 (
<figureCitation id="134C2A23FFC6FFC9CF71B7B5FAFC1EF1" box="[1340,1403,1181,1206]" captionStart="FIGURE 1" captionStartId="3.[151,250,1321,1343]" captionTargetBox="[162,1424,205,1294]" captionTargetId="figure@3.[150,1435,195,1296]" captionTargetPageId="3" captionText="FIGURE 1. Tyrphonothrus papuensis sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223435/files/figure.png" pageId="2" pageNumber="403">Fig. 1</figureCitation>
b). Each plate 127 Μm long, 55 Μm broad. Posterior margin of genital plate transverse; anterior margin rounded. One pair of anal setae 22 Μm long, longer than width of anal plate. Each adanal plate 116 Μm long, 33 Μm broad; three pairs of smooth, setiform adanal setae, 3142 Μm long, longer than width of adanal plate. Anterior margin of adanal plate overlapped by posterior margin of genital plate.
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CA8AB67EFDC61F2A" blockId="2.[151,1437,484,1971]" box="[199,577,1364,1389]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB67EFF7B1F2A" box="[199,252,1366,1389]" italics="true" pageId="2" pageNumber="403">Legs</emphasis>
. Pre-tarsi heterotridactylous.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC6FFC9CA8AB650FDC81F9E" pageId="2" pageNumber="403" type="materials_examined">
<paragraph id="8BC836A6FFC6FFC9CA8AB650FDC81F9E" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB650FE361FD6" bold="true" box="[199,433,1400,1425]" pageId="2" pageNumber="403">Material Examined</emphasis>
<typeStatus id="54CC8804FFC6FFC9CBF6B650FDA41FD6" box="[443,547,1400,1425]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54CC8804FFC6FFC9C813B651FD461FD6" box="[606,705,1401,1425]" pageId="2" pageNumber="403" type="paratype">paratype</typeStatus>
, ANIC 411, litter, rainforest, Yawasora, nr Wewak,
<collectingCountry id="F3607636FFC6FFC9CF54B651FF6C1FF2" name="Papua New Guinea" pageId="2" pageNumber="403">Papua New Guinea</collectingCountry>
, ca.
<quantity id="4C8F9B43FFC6FFC9CB68B6B4FEE31FF3" box="[293,356,1436,1461]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="403" unit="m" value="50.0">50 m</quantity>
, coll. R.W. Taylor,
<date id="FFC91066FFC6FFC9C802B6B4FD431FF2" box="[591,708,1436,1461]" pageId="2" pageNumber="403" value="1972-07-08">8.vii.1972</date>
.
<typeStatus id="54CC8804FFC6FFC9C89EB6B5FC911FF2" box="[723,790,1437,1461]" pageId="2" pageNumber="403">Types</typeStatus>
deposited in the Australian National Insect Collection, CSIRO Ecosystem Sciences, Canberra.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC6FFC9CA8AB6CCFBC21FBA" box="[199,1093,1508,1533]" pageId="2" pageNumber="403" type="etymology">
<paragraph id="8BC836A6FFC6FFC9CA8AB6CCFBC21FBA" blockId="2.[151,1437,484,1971]" box="[199,1093,1508,1533]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB6CCFEC91FBA" bold="true" box="[199,334,1508,1533]" pageId="2" pageNumber="403">Etymology.</emphasis>
This species is named for its country of origin,
<collectingCountry id="F3607636FFC6FFC9C928B6CDFBC61FBA" box="[869,1089,1508,1533]" name="Papua New Guinea" pageId="2" pageNumber="403">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC6FFC9CA8AB520FDE61DF4" pageId="2" pageNumber="403" type="discussion">
<paragraph id="8BC836A6FFC6FFC9CA8AB520FADB1CF4" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<emphasis id="B903EAB4FFC6FFC9CA8AB520FEB31C66" bold="true" box="[199,308,1544,1569]" pageId="2" pageNumber="403">Remarks</emphasis>
.
<taxonomicName id="4C774D25FFC6FFC9CB05B520FDEE1C66" box="[328,617,1544,1569]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="papuensis" status="sp. nov.">
<emphasis id="B903EAB4FFC6FFC9CB05B520FDEE1C66" box="[328,617,1544,1569]" italics="true" pageId="2" pageNumber="403">Tyrphonothrus papuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC6FFC9C83BB520FD521C67" box="[630,725,1544,1569]" pageId="2" pageNumber="403" rank="species">
<emphasis id="B903EAB4FFC6FFC9C83BB520FD491C66" bold="true" box="[630,718,1544,1569]" pageId="2" pageNumber="403">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from other members of the genus by the following combination of characters: 1) the prominent anterior protruding S-shaped carinae; 2) the combination of barbed and smooth notogastral setae, with
<emphasis id="B903EAB4FFC6FFC9C86EB578FDB61C2E" box="[547,561,1616,1641]" italics="true" pageId="2" pageNumber="403">d</emphasis>
1,
<emphasis id="B903EAB4FFC6FFC9C80AB57AFDD41C2E" box="[583,595,1618,1641]" italics="true" pageId="2" pageNumber="403">e</emphasis>
2 and
<emphasis id="B903EAB4FFC6FFC9C8DFB578FD271C2E" box="[658,672,1616,1641]" italics="true" pageId="2" pageNumber="403">h</emphasis>
1-2 smooth; 3) the very short setae
<emphasis id="B903EAB4FFC6FFC9CE50B57AFBAE1C2E" box="[1053,1065,1618,1641]" italics="true" pageId="2" pageNumber="403">c</emphasis>
1-3; 4) the well-developed median and lateral longitudinal ridges joining a complete M-shaped posterior ridge; 5) the well-developed foveae on the notogaster; 6) epimeral setal formula 3-1-2-3; 7) six pairs of flagelliform genital setae; 8) the long anal setae.
</paragraph>
<paragraph id="8BC836A6FFC6FFC9CA8AB597FDE61DF4" blockId="2.[151,1437,484,1971]" pageId="2" pageNumber="403">
<taxonomicName id="4C774D25FFC6FFC9CA8AB597FE641C90" box="[199,483,1727,1752]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="papuensis" status="sp. nov.">
<emphasis id="B903EAB4FFC6FFC9CA8AB597FE641C90" box="[199,483,1727,1752]" italics="true" pageId="2" pageNumber="403">Tyrphonothrus papuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC6FFC9CBA6B597FDC21C90" box="[491,581,1727,1752]" pageId="2" pageNumber="403" rank="species">
<emphasis id="B903EAB4FFC6FFC9CBA6B597FDB91C9F" bold="true" box="[491,574,1727,1752]" pageId="2" pageNumber="403">sp. nov</emphasis>
.
</taxonomicNameLabel>
is a member of the
<taxonomicName id="4C774D25FFC6FFC9C965B597FC381C9F" box="[808,959,1727,1752]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="428" phylum="Arthropoda" rank="species" species="crassisetosus">Crassisetosus</taxonomicName>
species group (cf. below). Like
<taxonomicName id="4C774D25FFC6FFC9CF62B5E8FA1B1C9F" box="[1327,1436,1727,1752]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="kanaka" status="sp. nov.">
<emphasis id="B903EAB4FFC6FFC9CF62B5E8FA1B1C9F" box="[1327,1436,1727,1752]" italics="true" pageId="2" pageNumber="403">T. kanaka</emphasis>
</taxonomicName>
<emphasis id="B903EAB4FFC6FFC9CADAB5CBFF761CBB" bold="true" box="[151,241,1763,1788]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A23057CFFFC6FFC9CADAB5CBFF761CBB" box="[151,241,1763,1788]" pageId="2" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C774D25FFC6FFC9CB65B5CDFD2E1CBB" authority="Wallwork, 1973" authorityName="Wallwork" authorityYear="1973" box="[296,681,1763,1788]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="heterotrichus">
<emphasis id="B903EAB4FFC6FFC9CB65B5CDFE5F1CBB" box="[296,472,1763,1788]" italics="true" pageId="2" pageNumber="403">T. heterotrichus</emphasis>
(Wallwork, 1973)
</taxonomicName>
, it has some of the notogastral setae smooth and the others barbed. In
<taxonomicName id="4C774D25FFC6FFC9CAF9B420FEA71D67" box="[180,288,1799,1824]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="kanaka">
<emphasis id="B903EAB4FFC6FFC9CAF9B420FEA71D67" box="[180,288,1799,1824]" italics="true" pageId="2" pageNumber="403">T. kanaka</emphasis>
</taxonomicName>
it is the
<emphasis id="B903EAB4FFC6FFC9CB33B420FE0B1D58" box="[382,396,1800,1823]" italics="true" pageId="2" pageNumber="403">p</emphasis>
series and
<emphasis id="B903EAB4FFC6FFC9C84AB42FFD921D67" box="[519,533,1799,1824]" italics="true" pageId="2" pageNumber="403">h</emphasis>
3 that are barbed, whereas in
<taxonomicName id="4C774D25FFC6FFC9C919B420FC581D58" box="[852,991,1800,1823]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B903EAB4FFC6FFC9C919B420FC581D58" box="[852,991,1800,1823]" italics="true" pageId="2" pageNumber="403">T. papuensis</emphasis>
</taxonomicName>
it is the longer setae (
<emphasis id="B903EAB4FFC6FFC9CE9EB420FB581D58" box="[1235,1247,1800,1823]" italics="true" pageId="2" pageNumber="403">e</emphasis>
2,
<emphasis id="B903EAB4FFC6FFC9CEB9B42FFA851D67" box="[1268,1282,1799,1824]" italics="true" pageId="2" pageNumber="403">h</emphasis>
1-2,
<emphasis id="B903EAB4FFC6FFC9CF69B420FAB51D58" box="[1316,1330,1800,1823]" italics="true" pageId="2" pageNumber="403">p</emphasis>
2) that are smooth and the others are barbed; a character state it shares with
<taxonomicName id="4C774D25FFC6FFC9C920B407FB9A1D00" box="[877,1053,1838,1863]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="heterotrichus">
<emphasis id="B903EAB4FFC6FFC9C920B407FB9A1D00" box="[877,1053,1838,1863]" italics="true" pageId="2" pageNumber="403">T. heterotrichus</emphasis>
</taxonomicName>
with which it also shares six pairs of genital setae, the relative lengths of the notogastral setae and the foveolate notogaster. They differ in that
<taxonomicName id="4C774D25FFC6FFC9CFCAB47BFEAB1DC8" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="heterotrichus">
<emphasis id="B903EAB4FFC6FFC9CFCAB47BFEAB1DC8" italics="true" pageId="2" pageNumber="403">T. heterotrichus</emphasis>
</taxonomicName>
has the alveoli of a second pair of anal setae, lacks the lateral longitudinal ridges and the M-shaped posterior ridge appears to be incomplete.
</paragraph>
</subSubSection>
</treatment>
</document>