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<document id="F08A319EFB6E7BA2F84632CD4A821B05" ID-CLB-Dataset="46384" ID-DOI="10.5281/zenodo.278635" ID-GBIF-Dataset="42311ac7-9128-48bf-9a2d-00da0f36148b" ID-ISSN="1175-5326" ID-Zenodo-Dep="278635" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460387208906" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2011" docId="03DE8797FFDD955367AEFA18FC9AC588" docLanguage="en" docName="zt03018p026.pdf" docOrigin="Zootaxa 3018" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Zehntnerobolus hoffmani Wesener, new species" docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FFE7FFEFFFDF95576739FFE9FF6DC206" masterDocTitle="Re-discovery after more than a century: a redefinition of the Malagasy endemic millipede genus Zehntnerobolus, with a description of a new species (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="26" masterPageNumber="21" pageNumber="23" updateTime="1698297763292" updateUser="ExternalLinkService">
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<mods:title id="306AE87230069280B78D2F20FCD91996">Re-discovery after more than a century: a redefinition of the Malagasy endemic millipede genus Zehntnerobolus, with a description of a new species (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="17341962E690604C2CF0BBF2BF2736AC">Wesener, Thomas</mods:namePart>
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<mods:title id="DF2E0600F76D5A86B6A63F71F8227E45">Zootaxa</mods:title>
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<mods:date id="D88A18F3CFD881D9460383242770F314">2011</mods:date>
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<mods:identifier id="F9A62295B11990D1DC0A534B19C6A48B" type="DOI">10.5281/zenodo.278635</mods:identifier>
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<treatment id="03DE8797FFDD955367AEFA18FC9AC588" ID-DOI="http://doi.org/10.5281/zenodo.6192913" ID-GBIF-Taxon="119415850" ID-Zenodo-Dep="6192913" LSID="urn:lsid:plazi:treatment:03DE8797FFDD955367AEFA18FC9AC588" httpUri="http://treatment.plazi.org/id/03DE8797FFDD955367AEFA18FC9AC588" lastPageId="4" lastPageNumber="25" pageId="2" pageNumber="23">
<subSubSection id="C36D650AFFDD955567AEFA18FDB5C40D" box="[151,728,1521,1547]" pageId="2" pageNumber="23" type="nomenclature">
<paragraph id="8BC83681FFDD955567AEFA18FDB5C40D" blockId="2.[151,728,1521,1547]" box="[151,728,1521,1547]" pageId="2" pageNumber="23">
<heading id="D08081EDFFDD955567AEFA18FDB5C40D" bold="true" box="[151,728,1521,1547]" fontSize="11" level="1" pageId="2" pageNumber="23" reason="1">
<emphasis id="B903EA93FFDD955567AEFA18FDB5C40D" bold="true" box="[151,728,1521,1547]" pageId="2" pageNumber="23">
<taxonomicName id="4C774D02FFDD955567AEFA18FD2DC40D" authority="Wesener" authorityName="Wesener" box="[151,576,1521,1547]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="hoffmani" status="sp. nov.">
<emphasis id="B903EA93FFDD955567AEFA18FEA3C40D" bold="true" box="[151,462,1521,1547]" italics="true" pageId="2" pageNumber="23">Zehntnerobolus hoffmani</emphasis>
Wesener
</taxonomicName>
<taxonomicNameLabel id="A23057E8FFDD95556571FA18FDB5C40D" box="[584,728,1521,1547]" pageId="2" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36D650AFFDD955567AEF9D1FA60C456" box="[151,1293,1592,1616]" pageId="2" pageNumber="23" type="etymology">
<paragraph id="8BC83681FFDD955567AEF9D1FA60C456" blockId="2.[151,1436,1592,2012]" box="[151,1293,1592,1616]" pageId="2" pageNumber="23">
<emphasis id="B903EA93FFDD955567AEF9D1FE73C456" bold="true" box="[151,286,1592,1616]" pageId="2" pageNumber="23">Etymology.</emphasis>
<taxonomicName id="4C774D02FFDD9555661DF9D1FEEAC456" box="[292,391,1592,1616]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="B903EA93FFDD9555661DF9D1FEEAC456" box="[292,391,1592,1616]" italics="true" pageId="2" pageNumber="23">hoffmani</emphasis>
</taxonomicName>
, adjective, to honour the 'Altmeister' of the Diplopoda, Dr. Richard L. Hoffman.
</paragraph>
</subSubSection>
<subSubSection id="C36D650AFFDD955567FEF9B5FC06C4E6" pageId="2" pageNumber="23" type="materials_examined">
<paragraph id="8BC83681FFDD955567FEF9B5FCEEC4BA" blockId="2.[151,1436,1592,2012]" pageId="2" pageNumber="23">
<emphasis id="B903EA93FFDD955567FEF9B5FEDDC473" bold="true" box="[199,432,1628,1653]" pageId="2" pageNumber="23">Material examined.</emphasis>
<emphasis id="B903EA93FFDD9555668EF9B5FD70C472" box="[439,541,1628,1652]" italics="true" pageId="2" pageNumber="23">
<typeStatus id="54CC8823FFDD9555668EF9B5FD70C472" box="[439,541,1628,1652]" pageId="2" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<emphasis id="B903EA93FFDD95556553F9B5FC6EC473" bold="true" box="[618,771,1628,1653]" pageId="2" pageNumber="23">
<collectionCode id="ED66AE44FFDD95556553F9B5FDA9C473" box="[618,708,1628,1653]" pageId="2" pageNumber="23">FMMC</collectionCode>
4014
</emphasis>
),
<collectingCountry id="F3607611FFDD95556423F9B5FCCEC472" box="[794,931,1628,1652]" name="Madagascar" pageId="2" pageNumber="23">Madagascar</collectingCountry>
, Province Fianarantsoa, Ihorombe, R. S. Ivohibe, 6.5 km ESE Ivohibe, camp III,
<geoCoordinate id="EE435046FFDD9555657FF969FDB4C49E" box="[582,729,1664,1688]" direction="south" orientation="latitude" pageId="2" pageNumber="23" precision="15" value="-22.496668">22°29' 48&quot; S</geoCoordinate>
,
<geoCoordinate id="EE435046FFDD955565D1F969FC18C49E" box="[744,885,1664,1688]" direction="east" orientation="longitude" pageId="2" pageNumber="23" precision="15" value="46.955">46°57'18&quot; E</geoCoordinate>
,
<quantity id="4C8F9B64FFDD955564BAF969FCB1C49E" box="[899,988,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="2" pageNumber="23" unit="m" value="1575.0">1575 m</quantity>
, montane rainforest, coll. B. L. Fisher, mini-Winkler, sifted litter (leaf mold, rotten wood),
<date id="FFC91041FFDD955565E5F94DFCEDC4BA" box="[732,896,1700,1724]" pageId="2" pageNumber="23" value="1997-10-24" valueMax="1997-10-30" valueMin="1997-10-24">2430.X.1997</date>
.
</paragraph>
<paragraph id="8BC83681FFDD955567AEF920FC06C4E6" blockId="2.[151,1436,1592,2012]" box="[151,875,1735,1760]" pageId="2" pageNumber="23">
<typeStatus id="54CC8823FFDD955567AEF920FF93C4E6" box="[151,254,1737,1760]" pageId="2" pageNumber="23" type="paratype">
<emphasis id="B903EA93FFDD955567AEF920FF93C4E6" box="[151,254,1737,1760]" italics="true" pageId="2" pageNumber="23">Paratype</emphasis>
</typeStatus>
: 1 immature M (
<emphasis id="B903EA93FFDD95556684F92EFD3BC4E6" bold="true" box="[445,598,1735,1760]" pageId="2" pageNumber="23">
<collectionCode id="ED66AE44FFDD95556684F92EFD7AC4E6" box="[445,535,1735,1760]" pageId="2" pageNumber="23">FMMC</collectionCode>
4014
</emphasis>
), same data as
<typeStatus id="54CC8823FFDD9555643BF921FC05C4E6" box="[770,872,1736,1760]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D650AFFDD955567FEF905FB23C592" pageId="2" pageNumber="23" type="diagnosis">
<paragraph id="8BC83681FFDD955567FEF905FB23C592" blockId="2.[151,1436,1592,2012]" pageId="2" pageNumber="23">
<emphasis id="B903EA93FFDD955567FEF905FE57C503" bold="true" box="[199,314,1772,1797]" pageId="2" pageNumber="23">Diagnosis</emphasis>
: species of
<taxonomicName id="4C774D02FFDD955566FCF905FD19C502" box="[453,628,1772,1796]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA93FFDD955566FCF905FD19C502" box="[453,628,1772,1796]" italics="true" pageId="2" pageNumber="23">Zehntnerobolus</emphasis>
</taxonomicName>
differing from its congener in shape of posterior gonopods: (1) coxite separated by articulation into basis and process; (2) telopodite with apical membranous lamellae resembling a 'sail'.
<taxonomicName id="4C774D02FFDD955567AEF8DCFE77C54A" box="[151,282,1844,1868]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="hoffmani" status="sp. nov.">
<emphasis id="B903EA93FFDD955567AEF8DCFE77C54A" box="[151,282,1844,1868]" italics="true" pageId="2" pageNumber="23">Z. hoffmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057E8FFDD9555661DF8DCFE0BC54B" box="[292,358,1845,1869]" pageId="2" pageNumber="23" rank="species">
<emphasis id="B903EA93FFDD9555661DF8DCFE0BC54B" bold="true" box="[292,358,1845,1869]" pageId="2" pageNumber="23">n. sp.</emphasis>
</taxonomicNameLabel>
differs also from
<taxonomicName id="4C774D02FFDD95556503F8DCFDD4C54A" box="[570,697,1844,1868]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B903EA93FFDD95556503F8DCFDD4C54A" box="[570,697,1844,1868]" italics="true" pageId="2" pageNumber="23">Z. rubripes</emphasis>
</taxonomicName>
in the lack of sclerotized nodules on the telopodite of posterior gonopods, presence of a slender sternite of the anterior gonopods (massive in
<taxonomicName id="4C774D02FFDD95556323F8B0FBF4C576" box="[1050,1177,1880,1904]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B903EA93FFDD95556323F8B0FBF4C576" box="[1050,1177,1880,1904]" italics="true" pageId="2" pageNumber="23">Z. rubripes</emphasis>
</taxonomicName>
), and the only weakly developed coxal processes on male leg pairs 35 (strongly developed in
<taxonomicName id="4C774D02FFDD955564FBF894FB53C592" box="[962,1086,1916,1940]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="rubripes">
<emphasis id="B903EA93FFDD955564FBF894FB53C592" box="[962,1086,1916,1940]" italics="true" pageId="2" pageNumber="23">Z. rubripes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D650AFFDD955367FEF876FB3AC524" lastPageId="4" lastPageNumber="25" pageId="2" pageNumber="23" type="description">
<paragraph id="8BC83681FFDD955567FEF876FDC9C5DA" blockId="2.[151,1436,1592,2012]" pageId="2" pageNumber="23">
<emphasis id="B903EA93FFDD955567FEF876FE34C5BE" bold="true" box="[199,345,1951,1976]" pageId="2" pageNumber="23">Description.</emphasis>
<emphasis id="B903EA93FFDD9555665BF848FD69C5BE" box="[354,516,1953,1976]" italics="true" pageId="2" pageNumber="23">Measurements</emphasis>
: male
<typeStatus id="54CC8823FFDD9555656EF849FDD4C5BE" box="[599,697,1952,1976]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
with 40 rings, circa
<quantity id="4C8F9B64FFDD95556496F849FB6CC5BE" box="[943,1025,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="23" unit="mm" value="36.0">36 mm</quantity>
long, 3.2 mm wide. Male
<typeStatus id="54CC8823FFDD95556205F848FAF1C5BF" box="[1340,1436,1953,1977]" pageId="2" pageNumber="23" type="paratype">paratype</typeStatus>
(immature) with 38+2 rings. Female unknown.
</paragraph>
<paragraph id="8BC83681FFDC955467AEFF71FD64C2F1" blockId="3.[151,1437,151,247]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467AEFF71FE7FC2B6" box="[151,274,152,176]" italics="true" pageId="3" pageNumber="24">Coloration</emphasis>
: decolorized after 13 years in ethanol. Head and antennae greyish-red (
<figureCitation id="134C2A04FFDC95546376FF7EFBF8C2A9" box="[1103,1173,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
A), legs brown-yellow. Body rings laterally grey, except for red posterior margin (
<figureCitation id="134C2A04FFDC95546419FF55FC0EC2D2" box="[800,867,188,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
B), dorsally with thin red stripe. Preanal scale grey, anal valves orange-red (
<figureCitation id="134C2A04FFDC9554669AFF36FE8AC2F1" box="[419,487,223,247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
C).
</paragraph>
<caption id="DF086609FFDC955467AEFBFDFD0FC6C2" httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24" targetBox="[151,1436,301,1023]" targetPageId="3">
<paragraph id="8BC83681FFDC955467AEFBFDFD0FC6C2" blockId="3.[151,1437,1044,1220]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467AEFBFDFE7FC62C" bold="true" box="[151,274,1044,1067]" pageId="3" pageNumber="24">FIGURE 2.</emphasis>
<taxonomicName id="4C774D02FFDC95546621FBFCFD7BC62D" box="[280,534,1045,1067]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="hoffmani" status="sp. nov.">
<emphasis id="B903EA93FFDC95546621FBFCFD7BC62D" box="[280,534,1045,1067]" italics="true" pageId="3" pageNumber="24">Zehntnerobolus hoffmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057E8FFDC95546525FBFCFD38C62C" box="[540,597,1045,1066]" pageId="3" pageNumber="24" rank="species">
<emphasis id="B903EA93FFDC95546525FBFCFD38C62C" bold="true" box="[540,597,1045,1066]" pageId="3" pageNumber="24">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype (FMMC 4014), multi-layer photographs.
<emphasis id="B903EA93FFDC95546358FBFCFB1EC62C" bold="true" box="[1121,1139,1045,1066]" pageId="3" pageNumber="24">A</emphasis>
: head and anterior five rings, lateral view;
<emphasis id="B903EA93FFDC95546624FBDDFE43C64F" bold="true" box="[285,302,1076,1097]" pageId="3" pageNumber="24">B</emphasis>
: midbody with ozopores, lateral view;
<emphasis id="B903EA93FFDC955465F9FBDAFDBFC64C" bold="true" box="[704,722,1075,1098]" pageId="3" pageNumber="24">C</emphasis>
: posterior body end;
<emphasis id="B903EA93FFDC95546496FBDDFCACC64F" bold="true" box="[943,961,1076,1097]" pageId="3" pageNumber="24">D</emphasis>
: head and anterior five rings, ventral view;
<emphasis id="B903EA93FFDC955462BDFBDDFAF8C64F" bold="true" box="[1412,1429,1076,1097]" pageId="3" pageNumber="24">E</emphasis>
: anterior gonopods, posterior view;
<emphasis id="B903EA93FFDC955466C0FBBAFD65C66E" bold="true" box="[505,520,1107,1128]" pageId="3" pageNumber="24">F</emphasis>
: posterior gonopods, posterior view;
<emphasis id="B903EA93FFDC955464BAFBBBFCF2C66F" bold="true" box="[899,927,1106,1129]" pageId="3" pageNumber="24">G:</emphasis>
posterior gonopods, anterior view. Abbreviations:
<emphasis id="B903EA93FFDC955467AEFB9BFFDCC681" bold="true" box="[151,177,1138,1159]" pageId="3" pageNumber="24">av</emphasis>
= paraproct;
<emphasis id="B903EA93FFDC95546603FB98FE34C68E" bold="true" box="[314,345,1137,1160]" pageId="3" pageNumber="24">Co</emphasis>
= collum;
<emphasis id="B903EA93FFDC955466F3FB98FE84C68E" bold="true" box="[458,489,1137,1160]" pageId="3" pageNumber="24">Cx</emphasis>
= coxite;
<emphasis id="B903EA93FFDC95546568FB98FD1FC68E" bold="true" box="[593,626,1137,1160]" pageId="3" pageNumber="24">Gn</emphasis>
= gnathochilarium;
<emphasis id="B903EA93FFDC9554647DFB9BFC32C681" bold="true" box="[836,863,1138,1159]" pageId="3" pageNumber="24">IL</emphasis>
= incisura lateralis;
<emphasis id="B903EA93FFDC95546310FB9BFB22C681" bold="true" box="[1065,1103,1138,1159]" pageId="3" pageNumber="24">Md</emphasis>
= mandible;
<emphasis id="B903EA93FFDC955463E1FB9BFA68C681" bold="true" box="[1240,1285,1138,1159]" pageId="3" pageNumber="24">Mes</emphasis>
= mesozonite;
<emphasis id="B903EA93FFDC955467AEFB79FFAEC6A3" bold="true" box="[151,195,1168,1189]" pageId="3" pageNumber="24">Met</emphasis>
= metazonite;
<emphasis id="B903EA93FFDC95546659FB66FE19C6A0" bold="true" box="[352,372,1167,1190]" pageId="3" pageNumber="24">O</emphasis>
= ozopore;
<emphasis id="B903EA93FFDC955466CAFB79FD74C6A3" bold="true" box="[499,537,1168,1189]" pageId="3" pageNumber="24">Pre</emphasis>
= epiproct;
<emphasis id="B903EA93FFDC955465A3FB79FDC2C6A3" bold="true" box="[666,687,1168,1189]" pageId="3" pageNumber="24">sc</emphasis>
= efferent duct (sperm canal);
<emphasis id="B903EA93FFDC955464CDFB66FB65C6A0" bold="true" box="[1012,1032,1167,1190]" pageId="3" pageNumber="24">St</emphasis>
= sternite;
<emphasis id="B903EA93FFDC955463BAFB79FBD2C6A3" bold="true" box="[1155,1215,1168,1189]" pageId="3" pageNumber="24">sub =</emphasis>
hypoproct;
<emphasis id="B903EA93FFDC95546204FB79FA23C6A3" bold="true" box="[1341,1358,1168,1189]" pageId="3" pageNumber="24">T</emphasis>
= telopodite;
<emphasis id="B903EA93FFDC955467DDFB46FE6FC6C2" bold="true" box="[228,258,1199,1220]" pageId="3" pageNumber="24">Tp</emphasis>
= telopodite process. Not to scale.
</paragraph>
</caption>
<paragraph id="8BC83681FFDC955467FCFB05FB73C792" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCFB05FE6CC702" box="[197,257,1260,1284]" italics="true" pageId="3" pageNumber="24">Head</emphasis>
: each eye with 2224 ocelli arranged in 6 vertical rows (
<figureCitation id="134C2A04FFDC95546490FB05FC9CC702" box="[937,1009,1260,1284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
A). Incisura lateralis open (
<figureCitation id="134C2A04FFDC95546208FB05FA15C702" box="[1329,1400,1260,1284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
A). Labrum with standard three irregular teeth and a single row of 1012 stout marginal setae. Clypeus with two setiferous foveolae on each side (
<figureCitation id="134C2A04FFDC955466D9FADDFD48C74A" box="[480,549,1332,1356]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
D). Antennae protruding back to ring
<quantity id="4C8F9B64FFDC955464E8FADDFC90C74A" box="[977,1021,1332,1356]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="24" unit="in" value="5.0">5 in</quantity>
males (
<figureCitation id="134C2A04FFDC9554636FFADDFBF6C74A" box="[1110,1179,1332,1356]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
D). Relative lengths of antennomeres: 1&lt;&lt;2=3=4=5=6. Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 and 6 latero-apically with sensilla basiconica.
</paragraph>
<paragraph id="8BC83681FFDC955467FCFA49FBCDC7DA" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCFA49FEEBC7BE" box="[197,390,1440,1464]" italics="true" pageId="3" pageNumber="24">Gnathochilarium</emphasis>
: lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Basal part on mentum with transverse ridge.
<emphasis id="B903EA93FFDC955464ADFA2DFC92C7DA" box="[916,1023,1476,1500]" italics="true" pageId="3" pageNumber="24">Mandible</emphasis>
not dissected.
</paragraph>
<paragraph id="8BC83681FFDC955467FCFA01FCB9C7F9" blockId="3.[151,1437,1260,1893]" box="[197,980,1511,1536]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCFA01FE4DC406" box="[197,288,1512,1536]" italics="true" pageId="3" pageNumber="24">Collum:</emphasis>
smooth, laterally not protruding as far as ring 2 (
<figureCitation id="134C2A04FFDC9554647FFA0EFCE1C7F9" box="[838,908,1511,1535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Figs 2</figureCitation>
A, D).
</paragraph>
<paragraph id="8BC83681FFDC955467FCF9E5FEF9C46A" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCF9E5FE27C422" box="[197,330,1548,1572]" italics="true" pageId="3" pageNumber="24">Body rings:</emphasis>
ozopores marked by reddish-brown color (
<figureCitation id="134C2A04FFDC9554640EF9E5FC15C422" box="[823,888,1548,1572]" captionStart="FIGURE 1" captionStartId="2.[151,250,1199,1222]" captionTargetBox="[476,1117,196,1174]" captionTargetId="figure@2.[474,1120,193,1178]" captionTargetPageId="2" captionText="FIGURE 1. Distribution map of Zehntnerobolus Wesener, 2009. Elevation in meters." httpUri="https://zenodo.org/record/278636/files/figure.png" pageId="3" pageNumber="24">Fig. 1</figureCitation>
B), starting at ring 6, located on suture between mesozona and metazona (
<figureCitation id="134C2A04FFDC95546680F9C6FD6FC441" box="[441,514,1583,1607]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Figs 2</figureCitation>
B, C). Rings with smooth, but irregular coriaecous surface, ventrally on metazona with transverse ridges.
</paragraph>
<paragraph id="8BC83681FFDC955467FCF991FCDEC4B2" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCF991FE66C496" box="[197,267,1656,1680]" italics="true" pageId="3" pageNumber="24">Telson</emphasis>
: paraprocts with weak lips, little micropunctation towards edges (
<figureCitation id="134C2A04FFDC955464CCF991FB55C496" box="[1013,1080,1656,1680]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
A). Epiproct sharp, but not protruding above paraproct (
<figureCitation id="134C2A04FFDC9554668BF975FE98C4B2" box="[434,501,1692,1716]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
C). Hypoproct inconspicuous (
<figureCitation id="134C2A04FFDC95546475F975FCE3C4B2" box="[844,910,1692,1716]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8BC83681FFDC955467FCF928FEC3C526" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467FCF928FF97C4DE" box="[197,250,1729,1752]" italics="true" pageId="3" pageNumber="24">Legs</emphasis>
: length of midbody legs circa 1.0 times body diameter in males (
<figureCitation id="134C2A04FFDC955464F4F929FB63C4DE" box="[973,1038,1728,1752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
B). Coxa long and elongated, cylindrical (
<figureCitation id="134C2A04FFDC955467DEF90DFE47C4FA" box="[231,298,1764,1788]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
B). Each podomere ventrally with an apical seta, tarsus with a stout dorso-apical seta and three pairs of regular ventral spines.
</paragraph>
<paragraph id="8BC83681FFDC955467AEF8C5FDCAC563" blockId="3.[151,1437,1260,1893]" pageId="3" pageNumber="24">
<emphasis id="B903EA93FFDC955467AEF8C5FEF3C542" box="[151,414,1836,1860]" italics="true" pageId="3" pageNumber="24">Male sexual characters</emphasis>
: tarsal pads absent (
<figureCitation id="134C2A04FFDC955465BBF8C5FDA9C542" box="[642,708,1836,1860]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
B). Coxae 3 and 4 with short, rod-shaped coxal process, protruding posteriorly only to adjacent coxa (
<figureCitation id="134C2A04FFDC95546579F8A4FDEEC563" box="[576,643,1869,1893]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
D).
</paragraph>
<caption id="DF086609FFDB955367AEFBB0FFB5C70F" httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25" targetBox="[409,1179,194,1093]" targetPageId="4">
<paragraph id="8BC83681FFDB955367AEFBB0FFB5C70F" blockId="4.[151,1436,1113,1289]" pageId="4" pageNumber="25">
<emphasis id="B903EA93FFDB955367AEFBB0FE7FC669" bold="true" box="[151,274,1113,1136]" pageId="4" pageNumber="25">FIGURE 3.</emphasis>
<taxonomicName id="4C774D02FFDB95536620FBB3FD7AC676" box="[281,535,1114,1136]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="hoffmani" status="sp. nov.">
<emphasis id="B903EA93FFDB95536620FBB3FD7AC676" box="[281,535,1114,1136]" italics="true" pageId="4" pageNumber="25">Zehntnerobolus hoffmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057E8FFDB95536527FBB3FD3AC669" box="[542,599,1114,1135]" pageId="4" pageNumber="25" rank="species">
<emphasis id="B903EA93FFDB95536527FBB3FD3AC669" bold="true" box="[542,599,1114,1135]" pageId="4" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype (FMMC 4014), schematic drawings.
<emphasis id="B903EA93FFDB95536303FBB3FB21C669" bold="true" box="[1082,1100,1114,1135]" pageId="4" pageNumber="25">A</emphasis>
: posterior body end;
<emphasis id="B903EA93FFDB9553621CFBB3FA5BC669" bold="true" box="[1317,1334,1114,1135]" pageId="4" pageNumber="25">B</emphasis>
: left midbody leg, posterior view;
<emphasis id="B903EA93FFDB95536699FB91FEDFC689" bold="true" box="[416,434,1144,1167]" pageId="4" pageNumber="25">C</emphasis>
: anterior gonopods, anterior view;
<emphasis id="B903EA93FFDB95536427FB90FC5DC688" bold="true" box="[798,816,1145,1166]" pageId="4" pageNumber="25">D</emphasis>
: anterior gonopods, posterior view;
<emphasis id="B903EA93FFDB95536391FB90FBD4C688" bold="true" box="[1192,1209,1145,1166]" pageId="4" pageNumber="25">E</emphasis>
: right posterior gonopod, overview, posterior view;
<emphasis id="B903EA93FFDB955366E3FB71FE84C6AB" bold="true" box="[474,489,1176,1197]" pageId="4" pageNumber="25">F</emphasis>
: right posterior gonopod, apical part, mesal view;
<emphasis id="B903EA93FFDB955364C0FB7EFB78C6A8" bold="true" box="[1017,1045,1175,1198]" pageId="4" pageNumber="25">G:</emphasis>
right posterior gonopods, apical part, anterior view. Abbreviations:
<emphasis id="B903EA93FFDB955366F6FB5FFE85C6CD" bold="true" box="[463,488,1206,1227]" pageId="4" pageNumber="25">av</emphasis>
= paraproct;
<emphasis id="B903EA93FFDB95536542FB5CFDF7C6CA" bold="true" box="[635,666,1205,1228]" pageId="4" pageNumber="25">Cx</emphasis>
= coxite;
<emphasis id="B903EA93FFDB95536433FB5CFC52C6CA" bold="true" box="[778,831,1205,1228]" pageId="4" pageNumber="25">Cx-p</emphasis>
= coxite process;
<emphasis id="B903EA93FFDB9553633FFB5CFB35C6CA" bold="true" box="[1030,1112,1205,1228]" pageId="4" pageNumber="25">mem-fr</emphasis>
= membraneous fringe;
<emphasis id="B903EA93FFDB95536267FB5FFAEEC6CD" bold="true" box="[1374,1411,1206,1227]" pageId="4" pageNumber="25">Pre</emphasis>
= epiproct;
<emphasis id="B903EA93FFDB955367CEFB3CFE61C6EC" bold="true" box="[247,268,1237,1258]" pageId="4" pageNumber="25">sc</emphasis>
= efferent duct (sperm canal);
<emphasis id="B903EA93FFDB9553657FFB3DFD37C6ED" bold="true" box="[582,602,1236,1259]" pageId="4" pageNumber="25">St</emphasis>
= sternite;
<emphasis id="B903EA93FFDB955365F7FB3CFD99C6EC" bold="true" box="[718,756,1237,1258]" pageId="4" pageNumber="25">sub</emphasis>
= hypoproct;
<emphasis id="B903EA93FFDB955364BAFB3CFCF9C6EC" bold="true" box="[899,916,1237,1258]" pageId="4" pageNumber="25">T</emphasis>
= telopodite;
<emphasis id="B903EA93FFDB9553631BFB3CFB2CC6EC" bold="true" box="[1058,1089,1237,1258]" pageId="4" pageNumber="25">Tp</emphasis>
= telopodite process. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BC83681FFDB955367FCFADFFBDAC7BC" blockId="4.[151,1437,1334,1935]" pageId="4" pageNumber="25">
<emphasis id="B903EA93FFDB955367FCFADFFEE1C748" box="[197,396,1334,1358]" italics="true" pageId="4" pageNumber="25">Anterior gonopod</emphasis>
sternite neither slender nor massive (
<figureCitation id="134C2A04FFDB95536415FADFFC03C748" box="[812,878,1334,1358]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
C), elongated into a wide, well-rounded lobe (
<figureCitation id="134C2A04FFDB95536257FADFFFC8C774" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
C). Sternite in anterior view well-visible (
<figureCitation id="134C2A04FFDB95536545FAB3FDA9C774" box="[636,708,1370,1394]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs 2</figureCitation>
E, 3D), without discernable apodemes. Coxite without processes (
<figureCitation id="134C2A04FFDB955367A6FA97FF84C790" box="[159,233,1406,1430]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs 2</figureCitation>
E, 3C). Telopodite with slender process arising mesally (
<figureCitation id="134C2A04FFDB9553644CFA97FCD7C790" box="[885,954,1406,1430]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
E), process conspicuously curved laterally (
<figureCitation id="134C2A04FFDB955367A6FA4BFF8FC7BC" box="[159,226,1442,1466]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
C, D), short with a well-rounded tip, not reaching lateral margin of telopodite (
<figureCitation id="134C2A04FFDB9553636AFA4BFBFBC7BC" box="[1107,1174,1442,1466]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Fig. 2</figureCitation>
E).
</paragraph>
<paragraph id="8BC83681FFDB955367FCFA2EFB3AC524" blockId="4.[151,1437,1334,1935]" pageId="4" pageNumber="25">
<emphasis id="B903EA93FFDB955367FCFA2EFECBC7D8" box="[197,422,1478,1502]" italics="true" pageId="4" pageNumber="25">Posterior gonopods</emphasis>
consisting of three parts, each separated by articulations (
<figureCitation id="134C2A04FFDB95536377FA2FFBF9C7D8" box="[1102,1172,1478,1502]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
E): (1) basally a large, bulky coxite, followed by (2) a slender, elongated process of coxite, and (3) apically a telopodite with free sperm channel discharging laterally (
<figureCitation id="134C2A04FFDB955366D3F9E7FD5DC420" box="[490,560,1550,1574]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Figs 3</figureCitation>
F, G). Process of coxite and telopodite are standing in same axis (
<figureCitation id="134C2A04FFDB9553622DF9E7FA36C420" box="[1300,1371,1550,1574]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs 2</figureCitation>
F, G). Pair of posterior gonopods located parallel to each other, connected by a small, sclerotized and visible sternite (
<figureCitation id="134C2A04FFDB95536257F9DBFFC8C468" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
E). Basal part of coxite wide, mesally with small sclerite located on lower level than remaining part (
<figureCitation id="134C2A04FFDB9553620FF9BFFA16C468" box="[1334,1403,1622,1646]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
E). Coxite elongated in long, slender coxite process, coxite and coxite process separated by suture (
<figureCitation id="134C2A04FFDB955363ECF993FA75C494" box="[1237,1304,1658,1682]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
E). Efferent duct running at mesal margin of coxite (
<figureCitation id="134C2A04FFDB9553656EF977FDF0C4B0" box="[599,669,1694,1718]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Figs 3</figureCitation>
E, F). Telopodite with torsion, moving discharge opening of efferent duct from mesally to laterally (
<figureCitation id="134C2A04FFDB955366C9F92BFD5BC4DC" box="[496,566,1730,1754]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs 2</figureCitation>
F, G). Telopodite shorter than coxite process (
<figureCitation id="134C2A04FFDB9553630CF92BFB10C4DC" box="[1077,1149,1730,1754]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs 2</figureCitation>
E, F), latero-apically with a wide, conspicuous membranous fringe (
<figureCitation id="134C2A04FFDB9553655EF90FFDD9C4F8" box="[615,692,1766,1790]" captionStart="FIGURE 2" captionStartId="3.[151,250,1044,1067]" captionTargetBox="[151,1436,301,1023]" captionTargetId="figure@3.[151,1436,301,1024]" captionTargetPageId="3" captionText="FIGURE 2. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), multi-layer photographs. A: head and anterior five rings, lateral view; B: midbody with ozopores, lateral view; C: posterior body end; D: head and anterior five rings, ventral view; E: anterior gonopods, posterior view; F: posterior gonopods, posterior view; G: posterior gonopods, anterior view. Abbreviations: av = paraproct; Co = collum; Cx = coxite; Gn = gnathochilarium; IL = incisura lateralis; Md = mandible; Mes = mesozonite; Met = metazonite; O = ozopore; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Not to scale." httpUri="https://zenodo.org/record/278637/files/figure.png" pageId="4" pageNumber="25">Figs. 2</figureCitation>
F, G, 3EG). Efferent duct running free starting at the apical part of process of coxite, protruding latero-apically above margins of telopodite (
<figureCitation id="134C2A04FFDB955364E9F8E3FB78C524" box="[976,1045,1802,1826]" captionStart="FIGURE 3" captionStartId="4.[151,250,1113,1136]" captionTargetBox="[409,1179,194,1093]" captionTargetId="figure@4.[409,1180,193,1094]" captionTargetPageId="4" captionText="FIGURE 3. Zehntnerobolus hoffmani n. sp., holotype (FMMC 4014), schematic drawings. A: posterior body end; B: left midbody leg, posterior view; C: anterior gonopods, anterior view; D: anterior gonopods, posterior view; E: right posterior gonopod, overview, posterior view; F: right posterior gonopod, apical part, mesal view; G: right posterior gonopods, apical part, anterior view. Abbreviations: av = paraproct; Cx = coxite; Cx-p = coxite process; mem-fr = membraneous fringe; Pre = epiproct; sc = efferent duct (sperm canal); St = sternite; sub = hypoproct; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." httpUri="https://zenodo.org/record/278638/files/figure.png" pageId="4" pageNumber="25">Figs 3</figureCitation>
F, G).
</paragraph>
</subSubSection>
<subSubSection id="C36D650AFFDB955367FEF8C7FC9AC588" pageId="4" pageNumber="25" type="distribution">
<paragraph id="8BC83681FFDB955367FEF8C7FC9AC588" blockId="4.[151,1437,1334,1935]" pageId="4" pageNumber="25">
<emphasis id="B903EA93FFDB955367FEF8C7FE9FC540" bold="true" box="[199,498,1838,1863]" pageId="4" pageNumber="25">Distribution and ecology.</emphasis>
This species is only known from the
<typeStatus id="54CC8823FFDB955364ACF8C6FCA8C541" box="[917,965,1839,1863]" pageId="4" pageNumber="25">type</typeStatus>
locality. Most likely living within the leaf litter (see genus description above), a cryptic lifestyle that explains the scarce records of this species despite largescale arthropod inventory programs conducted on
<collectingCountry id="F3607611FFDB955365F3F89FFC3CC588" box="[714,849,1910,1934]" name="Madagascar" pageId="4" pageNumber="25">Madagascar</collectingCountry>
(
<bibRefCitation id="EFE64B70FFDB95536466F89FFC86C588" author="Fisher" box="[863,1003,1910,1934]" pageId="4" pageNumber="25" refString="Fisher, B. L. (2005) A Model for a Global Inventory of Ants: A Case Study in Madagascar. In: N. G. Jablonski (Ed.), Biodiversity: A Symposium Held on the Occasion of the 150 th Anniversary of the California Academy of Sciences, San Francisco, U. S. A., pp. 78 - 89." type="book chapter" year="2005">Fisher 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>