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<mods:title id="9A380607479CB67AB00A0C7EBC1FB2C3">Bees of the Socotra Archipelago (Hymenoptera: Anthophila), their biogeography and association with parasites</mods:title>
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<mods:affiliation id="8E0C5FE7FF0B562E55A7E9443307C50F">) &amp;) &amp;) Department of Zoology, Charles University in Prague, Viničná 7, CZ- 12844 Praha 2, Czech Republic; e-mails: jakub. straka @ aculeataresearch. com; janbat @ centrum. cz</mods:affiliation>
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<mods:namePart id="CEE5A7C8A5F5B287B4CE0FB853A07534">Batelka, Jan</mods:namePart>
<mods:affiliation id="4613A55D37A2C6DC0EFA6AC008175BC1">) &amp;) &amp;) Department of Zoology, Charles University in Prague, Viničná 7, CZ- 12844 Praha 2, Czech Republic; e-mails: jakub. straka @ aculeataresearch. com; janbat @ centrum. cz</mods:affiliation>
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<mods:namePart id="994DAB5BB1FEC13A75167B808CF34E24">Pauly, Alain</mods:namePart>
<mods:affiliation id="4FB2A2E813FD20EC0D1DC7BA4B204904">) &amp;) Royal Belgian Institute of Natural Sciences, Entomology, Rue Vautier 29, B- 1000 Brussels, Belgium; e-mail: alain. pauly @ brutele. be</mods:affiliation>
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<taxonomicName id="4C77E3FDFF89FF97FE9145346FF0FCC1" ID-CoL="66B68" authority="(de Villers, 1789)" baseAuthorityName="de Villers" baseAuthorityYear="1789" box="[345,932,865,888]" class="Insecta" family="Apidae" genus="Amegilla" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="species" species="quadrifasciata" subGenus="Amegilla">
<emphasis id="B903446CFF89FF97FE9145346FF0FCC1" bold="true" box="[345,932,865,888]" pageId="2" pageNumber="185">
<emphasis id="B903446CFF89FF97FE9145346DE8FCC1" bold="true" box="[345,444,865,888]" italics="true" pageId="2" pageNumber="185">Amegilla</emphasis>
(
<emphasis id="B903446CFF89FF97FE0445346E7BFCC1" bold="true" box="[460,559,865,888]" italics="true" pageId="2" pageNumber="185">Amegilla</emphasis>
)
<emphasis id="B903446CFF89FF97FDF745346E8AFCC1" bold="true" box="[575,734,865,888]" italics="true" pageId="2" pageNumber="185">quadrifasciata</emphasis>
(de Villers, 1789)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF89FF97FDFB45D56E9FFC2A" box="[563,715,896,915]" pageId="2" pageNumber="185" type="description">
<paragraph id="8BC8987EFF89FF97FDFB45D56E9FFC2A" blockId="2.[151,1127,896,1044]" box="[563,715,896,915]" pageId="2" pageNumber="185">
(
<figureCitation id="134C84FBFF89FF97FDF145D56E90FC2A" box="[569,708,896,915]" captionStart="Figs 111. 13" captionStartId="3.[113,150,1511,1530]" captionTargetBox="[113,1082,214,1475]" captionTargetId="figure-11@3.[113,1088,213,1475]" captionTargetPageId="3" captionText="Figs 111. 13 Amegilla pyramidalis (Kirby, 1900). 1 female, dorsal view; 2 male, lateral view; 3 female, lateral view. 47 Amegilla quadrifasciata (de Villers, 1789). 4 female, dorsal view; 5 male, dorsal view; 6 female, lateral view; 7 male, lateral view. 89 Amegilla pyramidalis. 8 female, frons; 9 male, frons. 1011 Amegilla quadrifasciata. 10 female, frons; 11 male, frons." figureDoi="http://doi.org/10.5281/zenodo.5324220" httpUri="https://zenodo.org/record/5324220/files/figure.png" pageId="2" pageNumber="185">Figs 47, 10, 11</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BC8987EFF89FF97FF5F45F76FA1FC0C" blockId="2.[151,1127,896,1044]" box="[151,1013,930,949]" pageId="2" pageNumber="185">
<treatmentCitationGroup id="AB67BF50FF89FF97FF5F45F76FA1FC0C" box="[151,1013,930,949]" pageId="2" pageNumber="185">
<taxonomicName id="4C77E3FDFF89FF97FF5F45F76E4DFC0C" ID-CoL="4KP6D" authority="(de Villers, 1789)" baseAuthorityName="de Villers" baseAuthorityYear="1789" box="[151,537,930,949]" class="Insecta" family="Apidae" genus="Podalirius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="species" species="quadrifasciatus">
<emphasis id="B903446CFF89FF97FF5F45F76D28FC0C" box="[151,380,930,949]" italics="true" pageId="2" pageNumber="185">Podalirius quadrifasciatus</emphasis>
(de Villers, 1789)
</taxonomicName>
:
<treatmentCitation id="0AD6BE6FFF89FF97FDE945F76E95FC0C" author="KIRBY W. F." box="[545,705,930,949]" page="252" pageId="2" pageNumber="185" year="1903">
<bibRefCitation id="EFE6E58FFF89FF97FDE945F76EC2FC0C" author="KIRBY W. F." box="[545,662,930,949]" pageId="2" pageNumber="185" pagination="1898 - 9" refId="ref15844" refString="KIRBY W. F. 1903: Insecta: Hymenoptera. Pp. 235 - 258. In: FORBES H. O. (ed.): The natural history of Sokotra and Abd-el-Kuri: Being the report upon the results of the conjoint expedition to these islands in 1898 - 9. Special Bulletin of the Liverpool Museums, xlvii + 598 pp." type="book chapter" year="1903">KIRBY (1903)</bibRefCitation>
: 252
</treatmentCitation>
;
<treatmentCitation id="0AD6BE6FFF89FF97FD0145F76F1AFC0C" author="KOHL F. F." box="[713,846,930,949]" page="4" pageId="2" pageNumber="185" year="1906">
<bibRefCitation id="EFE6E58FFF89FF97FD0145F76F6CFC0C" author="KOHL F. F." box="[713,824,930,949]" pageId="2" pageNumber="185" refId="ref15911" refString="KOHL F. F. 1906: Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Kaiserlichen Akademie der Wissenschaften, Wien, 133 pp + XI pls. [Preprint of KOHL (1907)]." type="book" year="1906">KOHL (1906)</bibRefCitation>
: 4
</treatmentCitation>
;
<treatmentCitation id="0AD6BE6FFF89FF97FC9F45F76FA5FC0C" author="KOHL F. F." box="[855,1009,930,949]" page="172" pageId="2" pageNumber="185" year="1907">
<bibRefCitation id="EFE6E58FFF89FF97FC9F45F76F93FC0C" author="KOHL F. F." box="[855,967,930,949]" pageId="2" pageNumber="185" pagination="169 - 301" refId="ref15961" refString="KOHL F. F. 1907: Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 71: 169 - 301 + pls I - XI." type="journal article" year="1907">KOHL (1907)</bibRefCitation>
: 172
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C36DCBF5FF89FF97FF5F45996E03FBAD" pageId="2" pageNumber="185" type="materials_examined">
<paragraph id="8BC8987EFF89FF97FF5F45996E03FBAD" blockId="2.[151,1127,896,1044]" pageId="2" pageNumber="185">
<emphasis id="B903446CFF89FF97FF5F45996D1EFC66" bold="true" box="[151,330,972,991]" pageId="2" pageNumber="185">Material examined.</emphasis>
<materialsCitation id="3B1F9223FF89FF97FE9845996F55FC66" box="[336,769,965,991]" collectingDate="1899-01" collectorName="O. Simony" determinerName="F. Kohl" location="Sokotra" pageId="2" pageNumber="185" specimenCount="1" specimenCount-female="1">
<location id="8EA8CEA5FF89FF97FE9845996DC1FC66" LSID="urn:lsid:plazi:treatment:03DE2968FF89FF97FE9145346EF7FB5D:8EA8CEA5FF89FF97FE9845996DC1FC66" box="[336,405,972,991]" name="Sokotra" pageId="2" pageNumber="185">Sokotra</location>
,
<specimenCount id="9D7153F7FF89FF97FE5645996DEBFC67" box="[414,447,965,991]" pageId="2" pageNumber="185" type="female">1 ♀</specimenCount>
,
<date id="FFC9BEBEFF89FF97FE0345996E57FC66" box="[459,515,972,991]" pageId="2" pageNumber="185" value="1899-01">
<collectingDate id="EF8D4756FF89FF97FE0345996E57FC66" box="[459,515,972,991]" pageId="2" pageNumber="185" value="1899-01">i.1899</collectingDate>
</date>
,
<collectorName id="2682FDA8FF89FF97FDC445996E3DFC66" box="[524,617,972,991]" pageId="2" pageNumber="185">O. Simony</collectorName>
lgt.,
<determinerName id="7E1474BAFF89FF97FD5045996E8CFC66" box="[664,728,972,991]" pageId="2" pageNumber="185">F. Kohl</determinerName>
det.
</materialsCitation>
; J. Samha,
<specimenCount id="9D7153F7FF89FF97FCAC45996FC0FC67" box="[868,916,965,991]" pageId="2" pageNumber="185" type="female">3 ♀♀</specimenCount>
,
<date id="FFC9BEBEFF89FF97FC6845996F8CFC66" box="[928,984,972,991]" pageId="2" pageNumber="185" value="1899-01">i.1899</date>
, O. Simony lgt., H. Friese et F. Kohl det.;
<materialsCitation id="3B1F9223FF89FF97FEBA45B36860FC40" box="[370,1076,992,1018]" collectingDate="1899-01" collectionCode="NHMW" collectorName="O. Simony" determinerName="F. Kohl" location="Sokotra" municipality="Ras Shoab" pageId="2" pageNumber="185" specimenCount="1" specimenCount-female="1">
<location id="8EA8CEA5FF89FF97FEBA45B36DE3FC40" LSID="urn:lsid:plazi:treatment:03DE2968FF89FF97FE9145346EF7FB5D:8EA8CEA5FF89FF97FEBA45B36DE3FC40" box="[370,439,998,1017]" municipality="Ras Shoab" name="Sokotra" pageId="2" pageNumber="185">Sokotra</location>
,
<collectingMunicipality id="6BAC0204FF89FF97FE7745B36E48FC40" box="[447,540,998,1017]" pageId="2" pageNumber="185">Ras Shoab</collectingMunicipality>
, 1 J
<specimenCount id="9D7153F7FF89FF97FD8345B36E3FFC40" box="[587,619,992,1017]" pageId="2" pageNumber="185" type="female">1 ♀</specimenCount>
,
<date id="FFC9BEBEFF89FF97FDBE45B36EFAFC40" box="[630,686,998,1017]" pageId="2" pageNumber="185" value="1899-01">
<collectingDate id="EF8D4756FF89FF97FDBE45B36EFAFC40" box="[630,686,998,1017]" pageId="2" pageNumber="185" value="1899-01">i.1899</collectingDate>
</date>
,
<collectorName id="2682FDA8FF89FF97FD7F45B36F40FC40" box="[695,788,998,1017]" pageId="2" pageNumber="185">O. Simony</collectorName>
lgt.,
<determinerName id="7E1474BAFF89FF97FC8945B36FD5FC40" box="[833,897,998,1017]" pageId="2" pageNumber="185">F. Kohl</determinerName>
det. (all in
<collectionCode id="ED6600BBFF89FF97FC2C45B3687AFC40" box="[996,1070,998,1017]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="185" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F9223FF89FF97FBF545B36E07FBAD" collectingDate="2000-02-19" collectionCode="NMPC" collectorName="Wranik" location="wadi Deneghan" pageId="2" pageNumber="185" specimenCount="1" specimenCount-female="1">
<location id="8EA8CEA5FF89FF97FBF545B36CA4FBAD" LSID="urn:lsid:plazi:treatment:03DE2968FF89FF97FE9145346EF7FB5D:8EA8CEA5FF89FF97FBF545B36CA4FBAD" name="wadi Deneghan" pageId="2" pageNumber="185">wadi Deneghan</location>
,
<specimenCount id="9D7153F7FF89FF97FF3042546D4CFBAD" box="[248,280,1019,1044]" pageId="2" pageNumber="185" type="female">1 ♀</specimenCount>
,
<date id="FFC9BEBEFF89FF97FEEB42546D2FFBAD" box="[291,379,1025,1044]" pageId="2" pageNumber="185" value="2000-02-19">
<collectingDate id="EF8D4756FF89FF97FEEB42546D2FFBAD" box="[291,379,1025,1044]" pageId="2" pageNumber="185" value="2000-02-19">19.ii.2000</collectingDate>
</date>
, W.
<collectorName id="2682FDA8FF89FF97FE6842546D8BFBAD" box="[416,479,1025,1044]" pageId="2" pageNumber="185">Wranik</collectorName>
lgt. (
<collectionCode id="ED6600BBFF89FF97FDC442546E19FBAD" box="[524,589,1025,1044]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="185">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF89FF97FF5F42786ED9FBDD" pageId="2" pageNumber="185" type="discussion">
<paragraph id="8BC8987EFF89FF97FF5F42786ED9FBDD" blockId="2.[151,1127,1069,1252]" pageId="2" pageNumber="185">
<emphasis id="B903446CFF89FF97FF5F42786D52FBFD" bold="true" box="[151,262,1069,1092]" pageId="2" pageNumber="185">Remarks.</emphasis>
<bibRefCitation id="EFE6E58FFF89FF97FEC242786D8BFBFD" author="WRANIK W." box="[266,479,1069,1092]" pageId="2" pageNumber="185" refId="ref17108" refString="WRANIK W. 2003: Fauna of the Socotra Archipelago. Field guide. Universitat Rostock, Rostock, 540 pp." type="book" year="2003">WRANIK (2003: 454)</bibRefCitation>
presents a photograph of
<taxonomicName id="4C77E3FDFF89FF97FD2242786FF5FBFD" baseAuthorityName="de Villers" baseAuthorityYear="1789" box="[746,929,1069,1092]" class="Insecta" family="Apidae" genus="Amegilla" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="species" species="quadrifasciata">
<emphasis id="B903446CFF89FF97FD2242786FF5FBFD" box="[746,929,1069,1092]" italics="true" pageId="2" pageNumber="185">A. quadrifasciata</emphasis>
</taxonomicName>
from Hadiboh and incorrectly interprets this species as
<taxonomicName id="4C77E3FDFF89FF97FDD242186ED9FBDD" box="[538,653,1101,1124]" class="Insecta" family="Apidae" genus="Eucera" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="185" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B903446CFF89FF97FDD242186E33FBDD" box="[538,615,1101,1124]" italics="true" pageId="2" pageNumber="185">Eucera</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36DCBF5FF89FF97FF5F42386EF7FB5D" pageId="2" pageNumber="185" type="distribution">
<paragraph id="8BC8987EFF89FF97FF5F42386EF7FB5D" blockId="2.[151,1127,1069,1252]" pageId="2" pageNumber="185">
<emphasis id="B903446CFF89FF97FF5F42386D7EFB3D" bold="true" box="[151,298,1133,1156]" pageId="2" pageNumber="185">Distribution.</emphasis>
A species widely distributed in the entire Palaearctic Region, but absent in the Arabian Peninsula (
<bibRefCitation id="EFE6E58FFF89FF97FE4642D86EFDFB1D" author="ASCHER J. S. &amp; PICKERING J." box="[398,681,1165,1188]" pageId="2" pageNumber="185" refId="ref14368" refString="ASCHER J. S. &amp; PICKERING J. 2017: Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: http: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species (April 26, 2017)." type="url" year="2017">ASCHER &amp; PICKERING 2017</bibRefCitation>
). In
<collectingCountry id="F360D8EEFF89FF97FD1E42D86F73FB1D" box="[726,807,1165,1188]" name="Yemen" pageId="2" pageNumber="185">Socotra</collectingCountry>
reported from Moukaradia (= Gebel Raggit), Homhil, Dinehan Valley, Adho Dimellus (
<bibRefCitation id="EFE6E58FFF89FF97FD3542F86F2AFB7D" author="KIRBY W. F." box="[765,894,1197,1220]" pageId="2" pageNumber="185" pagination="1898 - 9" refId="ref15844" refString="KIRBY W. F. 1903: Insecta: Hymenoptera. Pp. 235 - 258. In: FORBES H. O. (ed.): The natural history of Sokotra and Abd-el-Kuri: Being the report upon the results of the conjoint expedition to these islands in 1898 - 9. Special Bulletin of the Liverpool Museums, xlvii + 598 pp." type="book chapter" year="1903">KIRBY 1903</bibRefCitation>
); Ras Shoab, Haghier Mts., and from Samha island (
<bibRefCitation id="EFE6E58FFF89FF97FE1242986E07FB5D" author="KOHL F. F." box="[474,595,1229,1252]" pageId="2" pageNumber="185" refId="ref15911" refString="KOHL F. F. 1906: Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Kaiserlichen Akademie der Wissenschaften, Wien, 133 pp + XI pls. [Preprint of KOHL (1907)]." type="book" year="1906">KOHL 1906</bibRefCitation>
,
<bibRefCitation id="EFE6E58FFF89FF97FD9642986EC3FB5D" author="KOHL F. F." box="[606,663,1229,1252]" pageId="2" pageNumber="185" pagination="169 - 301" refId="ref15961" refString="KOHL F. F. 1907: Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissenschaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 71: 169 - 301 + pls I - XI." type="journal article" year="1907">1907</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>