532 lines
62 KiB
XML
532 lines
62 KiB
XML
<document id="9BA9C650DAB2C4ED6B914ABD22AE91B8" ID-DOI="10.11646/zootaxa.3635.1.6" ID-GBIF-Dataset="d51bf3ca-437a-4d68-badd-6ced1c730244" ID-ISSN="1175-5326" ID-Zenodo-Dep="223886" ID-ZooBank="44EC57EA-12FB-4382-9C25-21DFCAFE7C90" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460585677443" checkinUser="plazi" docAuthor="Ng, Heok Hee & Kottelat, Maurice" docDate="2013" docId="03CA87EFFFBBFFD602B03A41FB18FADF" docLanguage="en" docName="zt03635p070.pdf" docOrigin="Zootaxa 3635 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rasbora tornieri Ahl 1922" docType="treatment" docVersion="8" masterDocId="FFF3FF97FFBFFFD102273F05FFFDFFA7" masterDocTitle="The identity of the cyprinid fishes Rasbora dusonensis and R. tornieri (Teleostei: Cyprinidae)" updateTime="1698325081030" updateUser="plazi">
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<mods:mods id="E4BC108842F9FF74A4AE584F3F36A50D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="602F3558E615A5E31FAC34D02BD5A993">
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<mods:title id="75937CD29818B901C39C5C04F7FBB5EC">The identity of the cyprinid fishes Rasbora dusonensis and R. tornieri (Teleostei: Cyprinidae)</mods:title>
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<mods:name id="F8A7F85ED53583F9886F2DCEF3ADABA0" type="personal">
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<mods:roleTerm id="ABC7399EF94B2EF8B5D4BA33D765B412">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="31F866EA6E65764FA27272B8F6970A1C">Ng, Heok Hee</mods:namePart>
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</mods:name>
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<mods:name id="4693E5FA6B7CE4030F598E53603C5379" type="personal">
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<mods:roleTerm id="06D2003173977BC30673563D9307D6D1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="08F244653F9288D772E5C4DBB68F9B37">Kottelat, Maurice</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3A496A5261CBC281770194386DF0F3BA">text</mods:typeOfResource>
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<mods:titleInfo id="6FD33E8080523B1562E73EAD895F29CD">
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<mods:title id="E15B80B0DD0993E31327EA1E45E08570">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="57470FEB020CCDC6F5AD06AB111F6A1B">
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<mods:date id="4CB7600D4134C96D7C50EBAD20F43A42">2013</mods:date>
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<mods:detail id="AD38AD1B13112408FD3F6FDE8AB3D5C9" type="volume">
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<mods:number id="5A9A64D6C26CBD140E1FF69347D53054">3635</mods:number>
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</mods:detail>
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<mods:detail id="CF843C747036B21C44B523111DE1A4D6" type="issue">
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<mods:number id="17AC035E4542EA7C09863DDBEB8E608E">1</mods:number>
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</mods:detail>
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<mods:classification id="D7B9CC042A88AF9F7E39243854728605">journal volume</mods:classification>
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<mods:identifier id="883B540F0BC6384C36ADAE5DE3CB76CA" type="DOI">10.11646/zootaxa.3635.1.6</mods:identifier>
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<mods:identifier id="65B70B038C4E3CE4C5DA82E6A8CD4B56" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="19AD37FFD07D4B6F2038ECA565C2E307" type="Zenodo-Dep">223886</mods:identifier>
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<treatment id="03CA87EFFFBBFFD602B03A41FB18FADF" ID-DOI="http://doi.org/10.5281/zenodo.5630988" ID-GBIF-Taxon="119564523" ID-Zenodo-Dep="5630988" LSID="urn:lsid:plazi:treatment:03CA87EFFFBBFFD602B03A41FB18FADF" httpUri="http://treatment.plazi.org/id/03CA87EFFFBBFFD602B03A41FB18FADF" lastPageId="7" pageId="4">
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<subSubSection id="C3796572FFBBFFD502B03A41FBC2FA63" pageId="4" type="nomenclature">
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<paragraph id="8BDC36F9FFBBFFD502B03A41FE21FAFA" blockId="4.[151,476,1347,1406]" box="[151,476,1347,1374]" pageId="4">
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<heading id="D0948195FFBBFFD502B03A41FE21FAFA" bold="true" box="[151,476,1347,1374]" fontSize="11" level="1" pageId="4" reason="1">
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<taxonomicName id="4C634D7AFFBBFFD502B03A41FE21FAFA" ID-CoL="4RLHK" authority="Ahl, 1922" authorityName="Ahl" authorityYear="1922" box="[151,476,1347,1374]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="tornieri">
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<emphasis id="B917EAEBFFBBFFD502B03A41FE21FAFA" bold="true" box="[151,476,1347,1374]" pageId="4">
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<emphasis id="B917EAEBFFBBFFD502B03A41FEA3FAF9" bold="true" box="[151,350,1348,1374]" italics="true" pageId="4">Rasbora tornieri</emphasis>
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Ahl, 1922
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDC36F9FFBBFFD502B03A63FED3FAD9" blockId="4.[151,476,1347,1406]" box="[151,302,1382,1406]" pageId="4">
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(
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<figureCitation id="13582A7CFFBBFFD502B83A63FF17FAD9" box="[159,234,1382,1406]" captionStart="FIGURE 1. a" captionStartId="3.[151,249,1118,1141]" captionTargetBox="[151,1435,337,1096]" captionTargetId="figure@3.[151,1436,337,1097]" captionTargetPageId="3" captionText="FIGURE 1. a. Rasbora dusonensis, ZRC 53470, 70.0 mm SL; b. Rasbora tornieri, ZRC 53469, 73.8 mm SL; both from Sumatra: Sungai Gelam. Figures not to scale." httpUri="https://zenodo.org/record/223887/files/figure.png" pageId="4">Figs. 1</figureCitation>
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b, 2b)
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</paragraph>
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<paragraph id="8BDC36F9FFBBFFD502B03AAAFBC2FA63" blockId="4.[151,1153,1454,1507]" box="[151,1087,1454,1477]" pageId="4">
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<taxonomicName id="4C634D7AFFBBFFD502B03AAAFE2FFA63" authority="Ahl, 1922: 32" authorityName="Ahl" authorityPageNumber="32" authorityYear="1922" box="[151,466,1454,1477]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="tornieri">
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<emphasis id="B917EAEBFFBBFFD502B03AAAFEBCFA62" box="[151,321,1455,1477]" italics="true" pageId="4">Rasbora tornieri</emphasis>
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Ahl, 1922: 32
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</taxonomicName>
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(
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<typeStatus id="54D8885BFFBBFFD503F83AAAFDF1FA63" box="[479,524,1455,1476]" pageId="4">type</typeStatus>
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locality: Central Sumatra); Kottelat
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<emphasis id="B917EAEBFFBBFFD501583AB5FC4EFA62" box="[895,947,1455,1477]" italics="true" pageId="4">et al.</emphasis>
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, 1993: pl. 20.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796572FFBBFFD502B03AC8FB7CFA44" box="[151,1153,1485,1507]" pageId="4" type="description">
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<paragraph id="8BDC36F9FFBBFFD502B03AC8FB7CFA44" blockId="4.[151,1153,1454,1507]" box="[151,1153,1485,1507]" pageId="4">
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<taxonomicName id="4C634D7AFFBBFFD502B03AC8FE9FFA44" box="[151,354,1485,1507]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="dusonensis">
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<emphasis id="B917EAEBFFBBFFD502B03AC8FE9FFA44" box="[151,354,1485,1507]" italics="true" pageId="4">Rasbora dusonensis</emphasis>
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</taxonomicName>
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(non Bleeker)—Brittan, 1954: 122, Fig. 26; Rainboth, 1996: 77, Pl. V Fig. 34.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796572FFBBFFD502B0390AFBB3F883" pageId="4" type="materials_examined">
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<paragraph id="8BDC36F9FFBBFFD502B0390AFBB3F883" blockId="4.[151,1436,1551,2008]" pageId="4">
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<emphasis id="B917EAEBFFBBFFD502B0390AFE77F98F" bold="true" box="[151,394,1551,1576]" pageId="4">Material examined.</emphasis>
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ZMB 20542 (
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<typeStatus id="54D8885BFFBBFFD50062390AFD4FF980" box="[581,690,1551,1575]" pageId="4" type="lectotype">lectotype</typeStatus>
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),
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<quantity id="4C9B9B1CFFBBFFD500E9390AFCC1F980" box="[718,828,1551,1575]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" pageId="4" unit="mm" value="54.5">54.5 mm</quantity>
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SL; Sumatra: Central Sumatra [Siak/Rokan river drainage].—UMMZ 232649 (4),
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<quantity id="4C9B9B1CFFBBFFD5002B3931FD52F9EB" box="[524,687,1588,1612]" metricMagnitude="-2" metricUnit="m" metricValue="5.484999999999999" metricValueMax="5.84" metricValueMin="5.13" pageId="4" unit="mm" value="54.849999999999994" valueMax="58.4" valueMin="51.3">51.3–58.4 mm</quantity>
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SL;
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<collectingCountry id="F3747669FFBBFFD500C13931FCA3F9EB" box="[742,862,1588,1612]" name="Cambodia" pageId="4">Cambodia</collectingCountry>
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: Kandal, Prek Lo Prampi just upstream from Phum Khleang, at fishing lots 6 and 7,
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<geoCoordinate id="EE57503EFFBBFFD500003952FD7FF9C8" box="[551,642,1623,1647]" direction="north" orientation="latitude" pageId="4" precision="925" value="11.166667">11°10'N</geoCoordinate>
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<geoCoordinate id="EE57503EFFBBFFD500A93952FD1FF9C8" box="[654,738,1623,1647]" direction="east" orientation="longitude" pageId="4" precision="925" value="105.15">105°9'E</geoCoordinate>
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.—CMK 20593 (28),
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<quantity id="4C9B9B1CFFBBFFD501CD3952FB6FF9C8" box="[1002,1170,1623,1647]" metricMagnitude="-2" metricUnit="m" metricValue="5.0649999999999995" metricValueMax="7.409999999999999" metricValueMin="2.7199999999999998" pageId="4" unit="mm" value="50.65" valueMax="74.1" valueMin="27.2">27.2–74.1 mm</quantity>
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SL; ZRC 53469 (26),
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<quantity id="4C9B9B1CFFBBFFD502B03979FEB9F933" box="[151,324,1660,1684]" metricMagnitude="-2" metricUnit="m" metricValue="5.695" metricValueMax="7.38" metricValueMin="4.01" pageId="4" unit="mm" value="56.95" valueMax="73.8" valueMin="40.1">40.1–73.8 mm</quantity>
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SL; Sumatra: Sumatera Selatan, Sungai Gelam, ca. 1 hour downriver from Kenten Laut (Palembang),
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<geoCoordinate id="EE57503EFFBBFFD50314399AFE47F910" box="[307,442,1695,1719]" direction="south" orientation="latitude" pageId="4" precision="1" value="-2.8373888">2°50'14.6"S</geoCoordinate>
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<geoCoordinate id="EE57503EFFBBFFD503E7399AFDA2F910" box="[448,607,1695,1719]" direction="east" orientation="longitude" pageId="4" precision="1" value="104.796974">104°47'49.1"E</geoCoordinate>
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.—CMK 7323 (15),
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<quantity id="4C9B9B1CFFBBFFD50160399AFC14F91F" box="[839,1001,1695,1720]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" metricValueMax="5.9799999999999995" metricValueMin="2.62" pageId="4" unit="mm" value="43.0" valueMax="59.8" valueMin="26.2">26.2–59.8 mm</quantity>
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SL; Sumatra: Riau: Sungei Siak Kecil, small blackwater canal entering mainstream,
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<geoCoordinate id="EE57503EFFBBFFD500B739C1FD22F97B" box="[656,735,1732,1756]" direction="north" orientation="latitude" pageId="4" precision="925" value="1.15">1°09'N</geoCoordinate>
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<geoCoordinate id="EE57503EFFBBFFD500C239C1FCB7F97B" box="[741,842,1732,1756]" direction="east" orientation="longitude" pageId="4" precision="925" value="102.03333">102°02'E</geoCoordinate>
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.—ZRC 42726 (5),
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<quantity id="4C9B9B1CFFBBFFD5060039C1FB37F97B" box="[1063,1226,1732,1756]" metricMagnitude="-2" metricUnit="m" metricValue="6.654999999999999" metricValueMax="8.92" metricValueMin="4.39" pageId="4" unit="mm" value="66.55" valueMax="89.2" valueMin="43.9">43.9–89.2 mm</quantity>
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SL;
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<collectingCountry id="F3747669FFBBFFD506D839C1FAADF97B" box="[1279,1360,1732,1756]" name="Brunei" pageId="4">Brunei</collectingCountry>
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: Belait District, base camp at Sungei Ingei,
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<geoCoordinate id="EE57503EFFBBFFD5006639E2FD43F958" box="[577,702,1767,1791]" direction="north" orientation="latitude" pageId="4" precision="1" value="4.1618333">4°9'42.6"N</geoCoordinate>
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<geoCoordinate id="EE57503EFFBBFFD500EE39E2FC95F958" box="[713,872,1767,1791]" direction="east" orientation="longitude" pageId="4" precision="1" value="114.71652">114°42'59.5"E</geoCoordinate>
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.—ZRC 49994 (25), 11.0–
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<quantity id="4C9B9B1CFFBBFFD506B939E2FAF8F8A7" box="[1182,1285,1767,1792]" metricMagnitude="-2" metricUnit="m" metricValue="8.03" pageId="4" unit="mm" value="80.3">80.3 mm</quantity>
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SL; Borneo: Kalimantan Barat, Kabupaten Sambas, Sungai Sambas,
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<geoCoordinate id="EE57503EFFBBFFD5012D3809FC68F883" box="[778,917,1804,1828]" direction="north" orientation="latitude" pageId="4" precision="1" value="1.3628334">1°21'46.2"N</geoCoordinate>
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<geoCoordinate id="EE57503EFFBBFFD501BB3809FBB7F883" box="[924,1098,1804,1828]" direction="east" orientation="longitude" pageId="4" precision="1" value="109.30801">109° 18'28.8"E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796572FFBBFFD402E0382AFC28FF50" lastPageId="5" pageId="4" type="diagnosis">
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<paragraph id="8BDC36F9FFBBFFD402E0382AFC28FF50" blockId="4.[151,1436,1551,2008]" lastBlockId="5.[151,1437,151,1796]" lastPageId="5" pageId="4">
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<emphasis id="B917EAEBFFBBFFD502E0382AFEBCF8EF" bold="true" box="[199,321,1839,1864]" pageId="4">Diagnosis.</emphasis>
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<taxonomicName id="4C634D7AFFBBFFD5036B3835FDF1F8EF" box="[332,524,1840,1864]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="tornieri">
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<emphasis id="B917EAEBFFBBFFD5036B3835FDF1F8EF" box="[332,524,1840,1864]" italics="true" pageId="4">Rasbora tornieri</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C634D7AFFBBFFD5006D3834FD15F8EF" box="[586,744,1840,1864]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="4" phylum="Chordata" rank="species" species="dusonensis">
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<emphasis id="B917EAEBFFBBFFD5006D3834FD15F8EF" box="[586,744,1840,1864]" italics="true" pageId="4">R. dusonensis</emphasis>
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</taxonomicName>
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are distinguished from congeners in having a broad, darkbrown sharply-defined midlateral stripe on body extending from opercle to caudal-fin base and separated from dark brown supralateral stripe and dorsum by highly contrasting light longitudinal area. The following characters (none unique) also help in identifying them: 12-14 predorsal scales; dorsohypural distance equal to or slightly less than distance between dorsal-fin origin and posterior orbital margin; 14 circumpeduncular scales; brightly coloured caudal fin in life.
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<taxonomicName id="4C634D7AFFBAFFD403783F9DFDE6FF17" box="[351,539,152,176]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="tornieri">
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<emphasis id="B917EAEBFFBAFFD403783F9DFDE6FF17" box="[351,539,152,176]" italics="true" pageId="5">Rasbora tornieri</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C634D7AFFBAFFD400973F9CFCB6FF17" box="[688,843,152,176]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="dusonensis">
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<emphasis id="B917EAEBFFBAFFD400973F9CFCB6FF17" box="[688,843,152,176]" italics="true" pageId="5">R. dusonensis</emphasis>
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</taxonomicName>
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in having a carmine red caudal fin without or with a very narrow black posterior margin (vs. bright yellow with a broad black posterior margin), and a deeper caudal peduncle (7.9–8.8 times in SL vs. 8.5–9.6; 11.4–12.7% SL vs. 10.4–11.7).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796572FFBAFFD402E03E06FD3BF9F7" pageId="5" type="description">
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<paragraph id="8BDC36F9FFBAFFD402E03E06FB2FFE53" blockId="5.[151,1437,151,1796]" pageId="5">
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<emphasis id="B917EAEBFFBAFFD402E03E06FEA4FEBB" bold="true" box="[199,345,259,284]" pageId="5">Description.</emphasis>
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Morphometric data are given in
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<tableCitation id="C6E10342FFBAFFD400CC3E01FCB9FEBB" box="[747,836,260,284]" captionStart="TABLE 1" captionStartId="2.[151,235,152,173]" captionTargetBox="[159,1311,232,1246]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data for Rasbora dusonensis (n = 20; based on ZRC 30452 – 30453, ZRC 40028 and ZRC 53470) and R. tornieri (n = 20; based on UMMZ 232649, ZRC 42726, ZRC 49994 and ZRC 53469)." httpUri="http://table.plazi.org/id/DF1C6671FFBDFFD302B03F9DFBEDFF6B" pageId="5" tableUuid="DF1C6671FFBDFFD302B03F9DFBEDFF6B">Table 1</tableCitation>
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, meristic data in
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<tableCitation id="C6E10342FFBAFFD4063B3E01FB85FEBB" box="[1052,1144,260,284]" captionStart="TABLE 2" captionStartId="4.[151,235,152,173]" captionTargetBox="[159,1420,262,925]" captionTargetPageId="4" captionText="TABLE 2. Meristic data for Rasbora dusonensis (n = 20; based on ZRC 30452 – 30453, ZRC 40028 and ZRC 53470) and R. tornieri (n = 20; based on UMMZ 232649, ZRC 42726, ZRC 49994 and ZRC 53469). Numbers in brackets indicate frequency of character state in material examined." httpUri="http://table.plazi.org/id/DF1C6671FFBBFFD502B03F9DFDDAFF4C" pageId="5" tableUuid="DF1C6671FFBBFFD502B03F9DFDDAFF4C">Table 2</tableCitation>
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. Body slender, elongate, laterally compressed, its greatest depth between pelvic-fin insertion and dorsal-fin origin. Dorsal profile of head sloping evenly from margin of upper lip to supraoccipital. Snout acutely rounded. Dorsal profile of body slightly convex from supraoccipital to dorsal-fin origin, thereafter slightly concave to caudal-fin base. Ventral profile gently convex from margin of lower lip to pelvic-fin origin, with slight concavity at isthmus; almost horizontal between pelvic- and anal-fin origins and gently concave at caudal peduncle. Mouth oblique, superior. Symphyseal knob on dentary strongly developed, fitting into corresponding indentation between premaxillae.
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</paragraph>
|
||
<paragraph id="8BDC36F9FFBAFFD402E03EFAFE9EFCB3" blockId="5.[151,1437,151,1796]" pageId="5">Scales cycloid. Lateral line complete (all scales perforated; 27–30+2) its anterior portion curving ventrally. Dorsal fin pointed at tip, with slightly convex posterior margin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray. Dorsohypural distance equal to or slightly less than distance between dorsal-fin origin and posterior orbital margin. Pectoral-fin subtriangular, with subrectangular axillary lobe dorsal to base of first unbranched ray. Tip of adpressed pectoral fin just reaching vertical through pelvic-fin insertion. Pelvic-fin profile triangular, with lanceolate axillary scale located dorsal to pelvic-fin base. Tip of adpressed pelvic fin reaching to anus. Anal fin with falcate posterior margin. Caudal fin deeply forked, with acutely pointed asymmetrical lobes, lower lobe longer.</paragraph>
|
||
<paragraph id="8BDC36F9FFBAFFD402E03C1AFB3BFA67" blockId="5.[151,1437,151,1796]" pageId="5">
|
||
<emphasis id="B917EAEBFFBAFFD402E03C1AFEB2FC9F" bold="true" box="[199,335,799,824]" pageId="5">Coloration.</emphasis>
|
||
In 70% ethanol: Dorsal and dorsolateral surfaces of head and body brown; ventrolateral and ventral surfaces unpigmented. Opercle pale brown dorsally, with distinct brown midopercular stripe. Middorsal stripe one-fourth scale wide, extending from nape to dorsal part of caudal peduncle. Pigments on posterior edge of scales distinct and present on up to three longitudinal scale rows on dorsolateral portion of body. Subopercle and pectoral axillary lobe with concentration of melanophores imparting brown appearance. Patch of pigments on scale pocket restricted to pore-bearing scales of laterosensory canal system between pectoral- and anal-fin origins, and in a few specimens on row immediately above it. Pigmentation most intense along anterior portion of lateral line, decreasing gradually in intensity posteriorly. In some specimens, additional pigments on scales of lateral line row forming a faint stripe. Brown midlateral stripe prominent, slightly more intense posteriorly, of approximately oneand-a-half scales width throughout its length, with very discrete and darker upper and lower edges. Stripe extending from behind pectoral girdle to tip of hypural plate. Dorsolateral stripe conspicuous and dark brown or blackish anteriorly, becoming less distinct and paler posteriorly; lower edge of stripe and of dorsal background pigmentation very discrete. Light area between midlateral and supralateral stripes almost totally devoid of pigments, of approximately half-scale width. Posterior extremity of axial streak overlapping midlateral stripe; extending forward along its upper edge until under dorsal-fin base and then for 3–4 scales in light area. Supraanal pigmentation originating slightly anterior to anal-fin origin, terminating at base of posteriormost anal-fin ray. Subpeduncular pigmentation present. Pectoral, dorsal, pelvic and anal fins hyaline, with scattered melanophores on rays. Caudal fin hyaline, with a diffuse brownish patch of pigments on proximal part of 4-5 outermost principal rays of each lobe; a very narrow black margin along posterior edge present in some individuals.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFBAFFD402E03AC9FD3BF9F7" blockId="5.[151,1437,151,1796]" pageId="5">
|
||
Coloration in life similar, with dark gray to black replacing brown coloration on dorsal and dorsolateral surfaces of head and body, reflective yellowish or greenish coloration on light longitudinal area; diffuse pigmentation along fin rays of pectoral, dorsal, pelvic and anal fins more prominent. Caudal fin carmine red, with very thin gray or black posterior margin (
|
||
<figureCitation id="13582A7CFFBAFFD40041393DFD54F9F7" box="[614,681,1592,1616]" captionStart="FIGURE 2" captionStartId="3.[151,250,1953,1976]" captionTargetBox="[172,1415,1240,1913]" captionTargetId="figure@3.[151,1436,1220,1931]" captionTargetPageId="3" captionText="FIGURE 2. Live coloration of: a. Rasbora dusonensis, ZRC 53470, 84.4 mm SL; b. Rasbora tornieri, ZRC ZRC 53469, 73.8 mm SL; both from Sumatra: Sungai Gelam. Figures not to scale." httpUri="https://zenodo.org/record/223888/files/figure.png" pageId="5">Fig. 2</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796572FFBAFFD402E0395EFAC1F947" pageId="5" type="distribution">
|
||
<paragraph id="8BDC36F9FFBAFFD402E0395EFAC1F947" blockId="5.[151,1437,151,1796]" pageId="5">
|
||
<emphasis id="B917EAEBFFBAFFD402E0395EFE9DF9D3" bold="true" box="[199,352,1627,1652]" pageId="5">Distribution.</emphasis>
|
||
<taxonomicName id="4C634D7AFFBAFFD4034B3959FDD1F9D3" box="[364,556,1628,1652]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFBAFFD4034B3959FDD1F9D3" box="[364,556,1628,1652]" italics="true" pageId="5">Rasbora tornieri</emphasis>
|
||
</taxonomicName>
|
||
is known from the Mekong River drainage in
|
||
<collectingCountry id="F3747669FFBAFFD406783959FB28F9D3" box="[1119,1237,1628,1652]" name="Cambodia" pageId="5">Cambodia</collectingCountry>
|
||
, as well as river drainages in the Malay Peninsula (from the Perak River drainage, according to Brittan, 1954), Sumatra (from the Siak River drainage southwards to the Musi River drainage) and Borneo (recorded from the Belait River drainage southwards and westwards to the Sambas River drainage; it possibly occurs in other drainages eastwards).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796572FFBAFFD602E039E9FB18FADF" lastPageId="7" pageId="5" type="discussion">
|
||
<paragraph id="8BDC36F9FFBAFFD402E039E9FBD9F8A3" blockId="5.[151,1437,151,1796]" box="[199,1060,1772,1796]" pageId="5">
|
||
<emphasis id="B917EAEBFFBAFFD402E039E9FEC6F8A3" bold="true" box="[199,315,1772,1796]" pageId="5">Remarks.</emphasis>
|
||
The
|
||
<taxonomicName id="4C634D7AFFBAFFD4035339E8FE15F8A3" box="[372,488,1773,1796]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFBAFFD4035339E8FE15F8A3" box="[372,488,1773,1796]" italics="true" pageId="5">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
figured by Rainboth (1996: pl. 6) is
|
||
<taxonomicName id="4C634D7AFFBAFFD401A339E8FBE0F8A3" box="[900,1053,1773,1796]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="aurotaenia">
|
||
<emphasis id="B917EAEBFFBAFFD401A339E8FBE0F8A3" box="[900,1053,1773,1796]" italics="true" pageId="5">R. aurotaenia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFBAFFD402B03852FEE8F8D7" blockId="5.[151,277,1879,1904]" box="[151,277,1879,1904]" pageId="5">
|
||
<emphasis id="B917EAEBFFBAFFD402B03852FEE8F8D7" bold="true" box="[151,277,1879,1904]" pageId="5">Discussion</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFBAFFD702B038A5FA86FE98" blockId="5.[151,1436,1952,2013]" lastBlockId="6.[151,1436,151,1076]" lastPageId="6" pageId="5">
|
||
Bleeker (1850: 14) described
|
||
<taxonomicName id="4C634D7AFFBAFFD403CE38A4FD78F81F" box="[489,645,1952,1976]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="5" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFBAFFD403CE38A4FD78F81F" box="[489,645,1952,1976]" italics="true" pageId="5">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
from Banjarmasin on the basis of a single specimen
|
||
<quantity id="4C9B9B1CFFBAFFD406C338A5FABCF81F" box="[1252,1345,1952,1976]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="5" unit="mm" value="115.0">115 mm</quantity>
|
||
TL. He described it as having, among others, a green caudal fin with a blackish posterior margin. This is clearly illustrated in the Atlas (Bleeker,
|
||
<date id="FFDD1039FFB9FFD703A83F92FE09FF08" box="[399,500,151,175]" pageId="6" value="1863" valueMax="1864">1863–64</date>
|
||
: pl. 120), but this figure does not show the
|
||
<typeStatus id="54D8885BFFB9FFD701F83F92FBB8FF08" box="[991,1093,151,175]" pageId="6" type="holotype">holotype</typeStatus>
|
||
. Figures in Bleeker's Atlas are natural size and this specimen is
|
||
<quantity id="4C9B9B1CFFB9FFD700353FB9FD8CFF73" box="[530,625,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="6" unit="mm" value="130.0">130 mm</quantity>
|
||
SL, which is too large to be the
|
||
<typeStatus id="54D8885BFFB9FFD701CA3FB9FBAEFF73" box="[1005,1107,188,212]" pageId="6" type="holotype">holotype</typeStatus>
|
||
. We have examined material from Banjarmasin (ZRC 40028) and the Barito River drainage (ZRC 38866, ZRC 53468) in agreement with Bleeker's figure and description in both coloration and morphometry (and in disagreement with Brittan's (1954: 122) description of
|
||
<taxonomicName id="4C634D7AFFB9FFD703543E2CFDF0FEE7" box="[371,525,296,320]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD703543E2CFDF0FEE7" box="[371,525,296,320]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
). The
|
||
<typeStatus id="54D8885BFFB9FFD700703E22FD44FE98" box="[599,697,295,319]" pageId="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C634D7AFFB9FFD700F93E2CFC85FEE7" box="[734,888,296,320]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD700F93E2CFC85FEE7" box="[734,888,296,320]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
could not be borrowed for direct comparison.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB9FFD702E03E49FEF7FE53" blockId="6.[151,1436,151,1076]" pageId="6">
|
||
<taxonomicName id="4C634D7AFFB9FFD702E03E49FE74FEC3" box="[199,393,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD702E03E49FE74FEC3" box="[199,393,332,356]" italics="true" pageId="6">Rasbora tornieri</emphasis>
|
||
</taxonomicName>
|
||
was described from an unspecified location in “Central Sumatra” by Ahl (1922). The collector, Max Moszkowski, travelled in the upper and middle parts of the Rokan Kanan, Rokan Kiri and Siak drainages ("Sultanate of Siak" [=the modern day regency (kabupaten) of Siak in Riau Province, Sumatra] and the "Rokan states"; Moszkowski, 1909a–b: maps). The
|
||
<typeStatus id="54D8885BFFB9FFD700D23EBDFCD8FE77" box="[757,805,440,464]" pageId="6">type</typeStatus>
|
||
locality can thus be restricted to the Rokan and Siak drainages.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB9FFD702E03EFAFC96FB93" blockId="6.[151,1436,151,1076]" pageId="6">
|
||
Brittan (1954) misidentified material referable to
|
||
<taxonomicName id="4C634D7AFFB9FFD700D33D04FC94FDBF" box="[756,873,513,536]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD700D33D04FC94FDBF" box="[756,873,513,536]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="4C634D7AFFB9FFD701A83D04FBD4FDBF" box="[911,1065,512,536]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD701A83D04FBD4FDBF" box="[911,1065,512,536]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
(see Kottelat, 1991), at the same time considering
|
||
<taxonomicName id="4C634D7AFFB9FFD7037D3D20FE32FD9B" box="[346,463,549,572]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD7037D3D20FE32FD9B" box="[346,463,549,572]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
a junior synonym of
|
||
<taxonomicName id="4C634D7AFFB9FFD700E63D20FC83FD9B" box="[705,894,548,572]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="cephalotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD700E63D20FC83FD9B" box="[705,894,548,572]" italics="true" pageId="6">R. cephalotaenia</emphasis>
|
||
</taxonomicName>
|
||
. After examining the
|
||
<typeStatus id="54D8885BFFB9FFD706513D20FB25FD9A" box="[1142,1240,549,573]" pageId="6" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C634D7AFFB9FFD706D93D20FA8EFD9B" box="[1278,1395,549,572]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD706D93D20FA8EFD9B" box="[1278,1395,549,572]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
, he (1972: unnumbered page between pp. 192–193, in the addendum of the reprint of the original 1954 edition) concluded that they are “examples of
|
||
<taxonomicName id="4C634D7AFFB9FFD7001B3D68FD2BFD23" box="[572,726,620,644]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD7001B3D68FD2BFD23" box="[572,726,620,644]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
, almost perfectly resembling fig. 26, p. 123 [of Brittan, 1954]”. The confusion was exacerbated by Alfred (1963: 130), who examined the
|
||
<typeStatus id="54D8885BFFB9FFD701F43D8AFBC8FD00" box="[979,1077,655,679]" pageId="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C634D7AFFB9FFD7067E3D94FB0EFD0F" box="[1113,1267,656,680]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD7067E3D94FB0EFD0F" box="[1113,1267,656,680]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and concluded that it was conspecific with
|
||
<taxonomicName id="4C634D7AFFB9FFD703FA3DB0FDB4FD6B" box="[477,585,693,716]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD703FA3DB0FDB4FD6B" box="[477,585,693,716]" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
. The fact that the
|
||
<typeStatus id="54D8885BFFB9FFD701063DB1FC7EFD6B" box="[801,899,692,716]" pageId="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C634D7AFFB9FFD701893DB0FBB6FD6B" box="[942,1099,692,716]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD701893DB0FBB6FD6B" box="[942,1099,692,716]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
is in poor condition and that “[o]nly a weak reticulate colouration is visible” may have prompted Alfred to synonymize
|
||
<taxonomicName id="4C634D7AFFB9FFD706EA3DDCFAC6FD57" box="[1229,1339,729,752]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD706EA3DDCFAC6FD57" box="[1229,1339,729,752]" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C634D7AFFB9FFD707A33DDCFEEFFCB3" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD707A33DDCFEEFFCB3" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
. This led to Kottelat's (1991) usage of
|
||
<taxonomicName id="4C634D7AFFB9FFD700E83DF8FC97FCB3" box="[719,874,764,788]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD700E83DF8FC97FCB3" box="[719,874,764,788]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
for the large plain gray species then known as
|
||
<taxonomicName id="4C634D7AFFB9FFD707A33DF8FF1FFC9F" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD707A33DF8FF1FFC9F" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
. However, we note that
|
||
<taxonomicName id="4C634D7AFFB9FFD703D33C24FDA2FC9F" box="[500,607,801,824]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD703D33C24FDA2FC9F" box="[500,607,801,824]" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
is a distinctly different species with the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to the distance between the dorsal-fin origin to the middle of the orbit), fewer scale rows between the lateral line and the midventral row in front of the pelvic fins (2 vs. 3) and with a much less distinct and contrasting dark lateral stripe (compare
|
||
<figureCitation id="13582A7CFFB9FFD7032C3CAAFEA7FC60" box="[267,346,943,967]" captionStart="FIGURE 1. a" captionStartId="3.[151,249,1118,1141]" captionTargetBox="[151,1435,337,1096]" captionTargetId="figure@3.[151,1436,337,1097]" captionTargetPageId="3" captionText="FIGURE 1. a. Rasbora dusonensis, ZRC 53470, 70.0 mm SL; b. Rasbora tornieri, ZRC 53469, 73.8 mm SL; both from Sumatra: Sungai Gelam. Figures not to scale." httpUri="https://zenodo.org/record/223887/files/figure.png" pageId="6">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="13582A7CFFB9FFD703B13CAAFE5BFC60" box="[406,422,943,967]" captionStart="FIGURE 3" captionStartId="6.[151,250,1569,1592]" captionTargetBox="[151,1436,1129,1547]" captionTargetId="figure@6.[151,1436,1129,1548]" captionTargetPageId="6" captionText="FIGURE 3. Rasbora myersi, CMK 6960, 58.6 mm SL: Borneo: Danau Temuan, a shallow blackwater lake near Nanga Embaluh." httpUri="https://zenodo.org/record/223889/files/figure.png" pageId="6">3</figureCitation>
|
||
). Although we were unable to examine the
|
||
<typeStatus id="54D8885BFFB9FFD701813CAAFBF5FC60" box="[934,1032,943,967]" pageId="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C634D7AFFB9FFD706143CB4FB2DFC6F" box="[1075,1232,944,968]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD706143CB4FB2DFC6F" box="[1075,1232,944,968]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
, we note that the diagnostic characters of
|
||
<taxonomicName id="4C634D7AFFB9FFD7038B3CD0FDE5FC4B" box="[428,536,981,1004]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD7038B3CD0FDE5FC4B" box="[428,536,981,1004]" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
by (e.g.) Brittan (1954) (absence of a distinct dark lateral stripe and caudal fin without black margin, or with only a blackish edge) are explicitly contradicted by the original description of
|
||
<taxonomicName id="4C634D7AFFB9FFD707A33CFCFEEFFB93" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD707A33CFCFEEFFB93" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
, thus ruling out the possibility of their conspecificity.
|
||
</paragraph>
|
||
<caption id="DF1C6671FFB9FFD702B03924FF07F9F1" httpUri="https://zenodo.org/record/223889/files/figure.png" pageId="6" targetBox="[151,1436,1129,1547]" targetPageId="6">
|
||
<paragraph id="8BDC36F9FFB9FFD702B03924FF07F9F1" blockId="6.[151,1436,1569,1622]" pageId="6">
|
||
<emphasis id="B917EAEBFFB9FFD702B03924FEEAF991" bold="true" box="[151,279,1569,1592]" pageId="6">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C634D7AFFB9FFD703053924FE3BF990" box="[290,454,1569,1591]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD703053924FE3BF990" box="[290,454,1569,1591]" italics="true" pageId="6">Rasbora myersi</emphasis>
|
||
</taxonomicName>
|
||
, CMK 6960, 58.6 mm SL: Borneo: Danau Temuan, a shallow blackwater lake near Nanga Embaluh.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDC36F9FFB9FFD702E03987FBC8F919" blockId="6.[151,1436,1666,2015]" pageId="6">
|
||
Kottelat (1991) examined the
|
||
<typeStatus id="54D8885BFFB9FFD700023986FDA8F93C" box="[549,597,1667,1691]" pageId="6">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C634D7AFFB9FFD700EB3986FCBEF93D" box="[716,835,1667,1690]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD700EB3986FCBEF93D" box="[716,835,1667,1690]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
, and designated and illustrated the
|
||
<typeStatus id="54D8885BFFB9FFD706C23987FAB1F93D" box="[1253,1356,1666,1690]" pageId="6" type="lectotype">lectotype</typeStatus>
|
||
(ZMB 20542). It has all the characters that we consider here as diagnostic for the species.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB9FFD602E039CFFE6CFF73" blockId="6.[151,1436,1666,2015]" lastBlockId="7.[151,1436,151,1400]" lastPageId="7" pageId="6">
|
||
Both
|
||
<taxonomicName id="4C634D7AFFB9FFD7032339CEFE63F945" box="[260,414,1738,1762]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD7032339CEFE63F945" box="[260,414,1738,1762]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB9FFD703F439CEFDB5F945" box="[467,584,1739,1762]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD703F439CEFDB5F945" box="[467,584,1739,1762]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
belong to the
|
||
<taxonomicName id="4C634D7AFFB9FFD700C139CEFC68F945" box="[742,917,1739,1762]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD700C139CEFC68F945" box="[742,917,1739,1762]" italics="true" pageId="6">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
group (sensu Brittan, 1954), which is unlikely to be monophyletic (Liao
|
||
<emphasis id="B917EAEBFFB9FFD7039939F5FE0AF8A1" box="[446,503,1774,1799]" italics="true" pageId="6">et al.</emphasis>
|
||
, 2009), but is used here for easier comparison among congeners. Members of this group are characterized by 14 circumpeduncular scale rows, the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin, 1 or 1½ scale rows between the lateral line and the origin of the pelvic fin, and a continuous dark midlateral stripe from the opercle to the base of the caudal fin (Kottelat, 2012). Species in the
|
||
<taxonomicName id="4C634D7AFFB9FFD700B23885FCB7F831" box="[661,842,1919,1943]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD700B23885FCB7F831" box="[661,842,1919,1943]" italics="true" pageId="6">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
group include:
|
||
<taxonomicName id="4C634D7AFFB9FFD7062D3885FB42F831" box="[1034,1215,1919,1943]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD7062D3885FB42F831" box="[1034,1215,1919,1943]" italics="true" pageId="6">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD706F53885FA8CF831" box="[1234,1393,1919,1943]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="aurotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD706F53885FA8CF831" box="[1234,1393,1919,1943]" italics="true" pageId="6">R. aurotaenia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD707A33885FEEDF81D" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis id="B917EAEBFFB9FFD707A33885FEEDF81D" italics="true" pageId="6">R. borneensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD7033938A6FE72F81D" box="[286,399,1955,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="B917EAEBFFB9FFD7033938A6FE72F81D" box="[286,399,1955,1978]" italics="true" pageId="6">R. everetti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD703BA38A6FDDBF81D" box="[413,550,1954,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="laticlavia">
|
||
<emphasis id="B917EAEBFFB9FFD703BA38A6FDDBF81D" box="[413,550,1954,1978]" italics="true" pageId="6">R. laticlavia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD7001438A6FD38F81D" box="[563,709,1954,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="leptosoma">
|
||
<emphasis id="B917EAEBFFB9FFD7001438A6FD38F81D" box="[563,709,1954,1978]" italics="true" pageId="6">R. leptosoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD700F438A6FCC1F81D" box="[723,828,1955,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB9FFD700F438A6FCC1F81D" box="[723,828,1955,1978]" italics="true" pageId="6">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD7016D38A6FC26F81D" box="[842,987,1954,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B917EAEBFFB9FFD7016D38A6FC26F81D" box="[842,987,1954,1978]" italics="true" pageId="6">R. philippina</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD701CE38A6FB89F81D" box="[1001,1140,1954,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="rheophila">
|
||
<emphasis id="B917EAEBFFB9FFD701CE38A6FB89F81D" box="[1001,1140,1954,1978]" italics="true" pageId="6">R. rheophila</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C634D7AFFB9FFD706A538A6FABAF81D" box="[1154,1351,1954,1978]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="B917EAEBFFB9FFD706A538A6FABAF81D" box="[1154,1351,1954,1978]" italics="true" pageId="6">R. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C634D7AFFB9FFD707A338A6FF01F879" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="vaillantii">
|
||
<emphasis id="B917EAEBFFB9FFD707A338A6FF01F879" italics="true" pageId="6">R. vaillantii</emphasis>
|
||
</taxonomicName>
|
||
. Both
|
||
<taxonomicName id="4C634D7AFFB9FFD7036F38CDFE1EF879" box="[328,483,1990,2015]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB9FFD7036F38CDFE1EF879" box="[328,483,1990,2015]" italics="true" pageId="6">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB9FFD7003D38CDFD72F879" box="[538,655,1991,2015]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB9FFD7003D38CDFD72F879" box="[538,655,1991,2015]" italics="true" pageId="6">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from congeners in the
|
||
<taxonomicName id="4C634D7AFFB9FFD706A738CDFAD2F879" box="[1152,1327,1991,2015]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="6" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB9FFD706A738CDFAD2F879" box="[1152,1327,1991,2015]" italics="true" pageId="6">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
group by the color pattern on the caudal fin in life (see diagnoses), except
|
||
<taxonomicName id="4C634D7AFFB8FFD601403F9CFBFDFF17" box="[871,1024,153,176]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="aurotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD601403F9CFBFDFF17" box="[871,1024,153,176]" italics="true" pageId="7">R. aurotaenia</emphasis>
|
||
</taxonomicName>
|
||
; additional distinguishing characters are highlighted below.
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB8FFD602E03FE5FE56FE68" blockId="7.[151,1436,151,1400]" pageId="7">
|
||
Among the species of the
|
||
<taxonomicName id="4C634D7AFFB8FFD600243FE4FD4AFF5F" box="[515,695,225,248]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD600243FE4FD4AFF5F" box="[515,695,225,248]" italics="true" pageId="7">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<taxonomicName id="4C634D7AFFB8FFD601303FE4FC48FF5F" box="[791,949,225,248]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="aurotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD601303FE4FC48FF5F" box="[791,949,225,248]" italics="true" pageId="7">R. aurotaenia</emphasis>
|
||
</taxonomicName>
|
||
is the only other species with a brightly coloured caudal fin. Like
|
||
<taxonomicName id="4C634D7AFFB8FFD603913E00FDADFEBB" box="[438,592,260,284]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD603913E00FDADFEBB" box="[438,592,260,284]" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
, the caudal fin is bright yellow with a conspicuous black poterior margin in life.
|
||
<taxonomicName id="4C634D7AFFB8FFD602E93E2DFE4FFEE7" box="[206,434,296,320]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD602E93E2DFE4FFEE7" box="[206,434,296,320]" italics="true" pageId="7">Rasbora dusonensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished fom
|
||
<taxonomicName id="4C634D7AFFB8FFD600963E2CFCB3FEE7" box="[689,846,297,320]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="aurotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD600963E2CFCB3FEE7" box="[689,846,297,320]" italics="true" pageId="7">R. aurotaenia</emphasis>
|
||
</taxonomicName>
|
||
by having a subpeduncular stripe (vs. absent), the midlateral and supralateral stripes with discrete edges and sharply contrasted, and the light area between them almost totally devoid of pigments (vs. stripes with indistinct edges and light area partly covered by pigments), a shorter pelvic axillary scale (about one-fifth of length of fin vs. one-third), and a more slender body (depth 4.0–4.8 times in SL vs. 3.7–4.0).
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB8FFD602E03ED9FC90FAAB" blockId="7.[151,1436,151,1400]" pageId="7">
|
||
The identity of
|
||
<taxonomicName id="4C634D7AFFB8FFD6035D3ED8FDD7FE53" box="[378,554,477,500]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD6035D3ED8FDD7FE53" box="[378,554,477,500]" italics="true" pageId="7">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
awaits further clarification (Kottelat, 2012); it was originally described from Java. Besides the striking colour differences mentioned above, both
|
||
<taxonomicName id="4C634D7AFFB8FFD6019D3D04FBA4FDBF" box="[954,1113,512,536]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD6019D3D04FBA4FDBF" box="[954,1113,512,536]" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB8FFD606BF3D04FAEFFDBF" box="[1176,1298,513,536]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD606BF3D04FAEFFDBF" box="[1176,1298,513,536]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
differ from Javanese populations identified as
|
||
<taxonomicName id="4C634D7AFFB8FFD6003B3D20FD36FD9B" box="[540,715,549,572]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="argyrotaenia">
|
||
<emphasis id="B917EAEBFFB8FFD6003B3D20FD36FD9B" box="[540,715,549,572]" italics="true" pageId="7">R. argyrotaenia</emphasis>
|
||
</taxonomicName>
|
||
in having the pectoral fin reaching (vs. not reaching) the pelvicfin origin. Both
|
||
<taxonomicName id="4C634D7AFFB8FFD6036B3D4CFE1AFDC7" box="[332,487,584,608]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD6036B3D4CFE1AFDC7" box="[332,487,584,608]" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB8FFD600393D4CFD69FDC7" box="[542,660,585,608]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD600393D4CFD69FDC7" box="[542,660,585,608]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
are further distinguished from
|
||
<taxonomicName id="4C634D7AFFB8FFD601D43D4CFB71FDC7" box="[1011,1164,584,608]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis id="B917EAEBFFB8FFD601D43D4CFB71FDC7" box="[1011,1164,584,608]" italics="true" pageId="7">R. borneensis</emphasis>
|
||
</taxonomicName>
|
||
in having a longer head (3.4–4.0 times in SL vs. 4.2–4.5) and fewer predorsal scales (13–14 vs. 15–16), from
|
||
<taxonomicName id="4C634D7AFFB8FFD606A33D68FB04FD23" box="[1156,1273,621,644]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="everetti">
|
||
<emphasis id="B917EAEBFFB8FFD606A33D68FB04FD23" box="[1156,1273,621,644]" italics="true" pageId="7">R. everetti</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to distance between the dorsal-fin origin and a point between the nostril and the anterior orbital margin) and fewer vertebrae (32–34 vs. 36–38), and from
|
||
<taxonomicName id="4C634D7AFFB8FFD601B23DDCFBDCFD57" box="[917,1057,728,752]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="laticlavia">
|
||
<emphasis id="B917EAEBFFB8FFD601B23DDCFBDCFD57" box="[917,1057,728,752]" italics="true" pageId="7">R. laticlavia</emphasis>
|
||
</taxonomicName>
|
||
in having more predorsal scales (13–14 vs. 11–12).
|
||
<taxonomicName id="4C634D7AFFB8FFD603513DF9FDA4FCB3" box="[374,601,764,788]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD603513DF9FDA4FCB3" box="[374,601,764,788]" italics="true" pageId="7">Rasbora dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB8FFD600B33DF8FCF7FCB3" box="[660,778,765,788]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD600B33DF8FCF7FCB3" box="[660,778,765,788]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
further differ from
|
||
<taxonomicName id="4C634D7AFFB8FFD601C83DF8FB79FCB3" box="[1007,1156,764,788]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="leptosoma">
|
||
<emphasis id="B917EAEBFFB8FFD601C83DF8FB79FCB3" box="[1007,1156,764,788]" italics="true" pageId="7">R. leptosoma</emphasis>
|
||
</taxonomicName>
|
||
in having a longer head (3.4–4.0 times in SL vs. 4.5–4.6), from
|
||
<taxonomicName id="4C634D7AFFB8FFD600723C24FD3DFC9F" box="[597,704,801,824]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="myersi">
|
||
<emphasis id="B917EAEBFFB8FFD600723C24FD3DFC9F" box="[597,704,801,824]" italics="true" pageId="7">R. myersi</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to the distance between the dorsal-fin origin to the middle of the orbit), fewer scale rows between the lateral line and the midventral row in front of the pelvic fins (2 vs. 3) and a much less distinct and contrasting dark lateral stripe (compare
|
||
<figureCitation id="13582A7CFFB8FFD606DC3C89FABAFC03" box="[1275,1351,908,932]" captionStart="FIGURE 1. a" captionStartId="3.[151,249,1118,1141]" captionTargetBox="[151,1435,337,1096]" captionTargetId="figure@3.[151,1436,337,1097]" captionTargetPageId="3" captionText="FIGURE 1. a. Rasbora dusonensis, ZRC 53470, 70.0 mm SL; b. Rasbora tornieri, ZRC 53469, 73.8 mm SL; both from Sumatra: Sungai Gelam. Figures not to scale." httpUri="https://zenodo.org/record/223887/files/figure.png" pageId="7">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="13582A7CFFB8FFD6075A3C89FA71FC03" box="[1405,1420,908,932]" captionStart="FIGURE 3" captionStartId="6.[151,250,1569,1592]" captionTargetBox="[151,1436,1129,1547]" captionTargetId="figure@6.[151,1436,1129,1548]" captionTargetPageId="6" captionText="FIGURE 3. Rasbora myersi, CMK 6960, 58.6 mm SL: Borneo: Danau Temuan, a shallow blackwater lake near Nanga Embaluh." httpUri="https://zenodo.org/record/223889/files/figure.png" pageId="7">3</figureCitation>
|
||
), and from
|
||
<taxonomicName id="4C634D7AFFB8FFD603213CB4FE64FC6F" box="[262,409,944,968]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="philippina">
|
||
<emphasis id="B917EAEBFFB8FFD603213CB4FE64FC6F" box="[262,409,944,968]" italics="true" pageId="7">R. philippina</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to distance between the dorsal-fin origin and a point slightly posterior to the nostril) and a more slender body (depth 3.8–4.5 times in SL vs. 3.1–3.8). Both
|
||
<taxonomicName id="4C634D7AFFB8FFD607A33CFCFEEFFB93" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD607A33CFCFEEFFB93" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB8FFD6036F3B18FE40FB93" box="[328,445,1053,1076]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD6036F3B18FE40FB93" box="[328,445,1053,1076]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
are further distinguished from
|
||
<taxonomicName id="4C634D7AFFB8FFD6013D3B18FC58FB93" box="[794,933,1052,1076]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="rheophila">
|
||
<emphasis id="B917EAEBFFB8FFD6013D3B18FC58FB93" box="[794,933,1052,1076]" italics="true" pageId="7">R. rheophila</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to the distance between the dorsal-fin origin and a point anterior to the nostril), fewer lateral-line scales (27–31+2–3 vs. 31–34+2–3) and vertebrae (32–34 vs. 36–38), from
|
||
<taxonomicName id="4C634D7AFFB8FFD600FA3B8CFC5FFB07" box="[733,930,1160,1184]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="B917EAEBFFB8FFD600FA3B8CFC5FFB07" box="[733,930,1160,1184]" italics="true" pageId="7">R. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
in lacking (vs. having) a distinct dark spot at the base of the median caudal-fin rays, and from
|
||
<taxonomicName id="4C634D7AFFB8FFD600F83BA8FC9AFB63" box="[735,871,1196,1220]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="vaillantii">
|
||
<emphasis id="B917EAEBFFB8FFD600F83BA8FC9AFB63" box="[735,871,1196,1220]" italics="true" pageId="7">R. vaillantii</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsohypural distance equal to or slightly less than the distance between the dorsal-fin origin and the posterior orbital margin (vs. equal to the distance between the dorsal-fin origin to the middle of the orbit).
|
||
</paragraph>
|
||
<paragraph id="8BDC36F9FFB8FFD602E03A1DFB18FADF" blockId="7.[151,1436,151,1400]" pageId="7">
|
||
Although both
|
||
<taxonomicName id="4C634D7AFFB8FFD603513A1CFDEEFA97" box="[374,531,1304,1328]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD603513A1CFDEEFA97" box="[374,531,1304,1328]" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D7AFFB8FFD600683A1CFD3AFA97" box="[591,711,1305,1328]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD600683A1CFD3AFA97" box="[591,711,1305,1328]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
were collected syntopically, we have not been able to observe whether they form mixed schools. Even though the two species can be found syntopically, we note that
|
||
<taxonomicName id="4C634D7AFFB8FFD607A33A38FEEFFADF" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="dusonensis">
|
||
<emphasis id="B917EAEBFFB8FFD607A33A38FEEFFADF" italics="true" pageId="7">R. dusonensis</emphasis>
|
||
</taxonomicName>
|
||
is more frequently seen in large rivers with murky water and
|
||
<taxonomicName id="4C634D7AFFB8FFD601E23A64FBC4FADF" box="[965,1081,1377,1400]" class="Actinopterygii" family="Cyprinidae" genus="Rasbora" kingdom="Animalia" order="Cypriniformes" pageId="7" phylum="Chordata" rank="species" species="tornieri">
|
||
<emphasis id="B917EAEBFFB8FFD601E23A64FBC4FADF" box="[965,1081,1377,1400]" italics="true" pageId="7">R. tornieri</emphasis>
|
||
</taxonomicName>
|
||
in black water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |