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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.4049.1.1" ID-GBIF-Dataset="13ce6790-fb60-4602-b68b-6d06c9f011e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="245367" ID-ZooBank="A8BD434E-9A7C-40A5-BDDE-D45356517FF3" checkinTime="1461348552233" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2015" docId="03CA87EF70718956F99FFCE24B156C64" docLanguage="en" docName="zt04049p098.pdf" docOrigin="Zootaxa 4049 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@77e7ad7b" docStyleName="zootaxa.2013.monograph" docTitle="Pagodatrochus variabilis H. Adams 1873" docType="treatment" docVersion="14" lastPageNumber="84" masterDocId="FFF3FF9770228905F908FFB44F5A6936" masterDocTitle="An annotated catalogue and bibliography of the taxonomy, synonymy and distribution of the Recent Vetigastropoda of South Africa (Mollusca)" masterLastPageNumber="98" masterPageNumber="1" pageNumber="84" updateTime="1661740910064" updateUser="ExternalLinkService">
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<mods:title>An annotated catalogue and bibliography of the taxonomy, synonymy and distribution of the Recent Vetigastropoda of South Africa (Mollusca)</mods:title>
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<mods:namePart>Herbert, David G.</mods:namePart>
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<mods:date>2015</mods:date>
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<mods:number>4049</mods:number>
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<heading bold="true" box="[151,655,854,879]" fontSize="10" level="2" pageId="83" pageNumber="84" reason="2">
<taxonomicName authority="H. Adams, 1873" authorityName="H. Adams" authorityYear="1873" box="[151,655,854,879]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis bold="true" box="[151,655,854,879]" pageId="83" pageNumber="84">
<emphasis bold="true" box="[151,439,854,879]" italics="true" pageId="83" pageNumber="84">Pagodatrochus variabilis</emphasis>
(
<bibRefCitation author="Adams" box="[455,646,854,879]" pageId="83" pageNumber="84" refString="Adams, H. (1873) Description of seventeen new species of land and marine shells. Proceedings of the Zoological Society of London, 41 (1873), 205 - 209." type="journal article" year="1873">
H. Adams,
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)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph blockId="83.[151,1436,854,1005]" pageId="83" pageNumber="84">
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<taxonomicName authority="H. Adams, 1873: 207" authorityName="H. Adams" authorityPageNumber="207" authorityYear="1873" box="[151,567,890,912]" class="Gastropoda" family="Solariellidae" genus="Minolia" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis box="[151,336,890,912]" italics="true" pageId="83" pageNumber="84">Minolia variabilis</emphasis>
<treatmentCitation author="Adams" box="[343,567,890,912]" page="207" pageId="83" pageNumber="84" year="1873">
<bibRefCitation author="Adams" box="[343,515,890,912]" pageId="83" pageNumber="84" refString="Adams, H. (1873) Description of seventeen new species of land and marine shells. Proceedings of the Zoological Society of London, 41 (1873), 205 - 209." type="journal article" year="1873">
H. Adams,
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:
<number box="[527,567,890,912]" pageId="83" pageNumber="84" value="207.0">207</number>
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</taxonomicName>
, pl.
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, fig.
<number box="[694,722,890,912]" pageId="83" pageNumber="84" value="10.0">10</number>
.
<typeStatus box="[733,783,891,912]" pageId="83" pageNumber="84">Type</typeStatus>
loc.: Persian Gulf;
<typeStatus box="[986,1075,890,912]" pageId="83" pageNumber="84">holotype</typeStatus>
in
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(
<collectionCode LSID="urn:lsid:biocol.org:col:34665" box="[1183,1280,890,912]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84">NHMUK</collectionCode>
1878.1.28.96), figured by
<bibRefCitation author="Herbert" box="[310,469,921,943]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">
Herbert (
<number box="[401,455,921,943]" pageId="83" pageNumber="84" value="1989.0">1989</number>
b
</bibRefCitation>
: fig.
<number box="[519,529,921,943]" pageId="83" pageNumber="84" value="1.0">1</number>
e).
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</paragraph>
<paragraph blockId="83.[151,1436,854,1005]" pageId="83" pageNumber="84">
<treatmentCitationGroup pageId="83" pageNumber="84">
<taxonomicName box="[151,409,952,974]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="variabilis">
<emphasis box="[151,409,952,974]" italics="true" pageId="83" pageNumber="84">Pagodatrochus variabilis</emphasis>
</taxonomicName>
<treatmentCitation author="Herbert" box="[435,645,952,974]" page="366" pageId="83" pageNumber="84" year="1989">
<bibRefCitation author="Herbert" box="[435,593,952,974]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">
Herbert,
<number box="[526,580,952,974]" pageId="83" pageNumber="84" value="1989.0">1989</number>
b
</bibRefCitation>
:
<number box="[604,645,952,974]" pageId="83" pageNumber="84" value="366.0">366</number>
</treatmentCitation>
, figs
<number box="[701,710,952,974]" pageId="83" pageNumber="84" value="1.0">1</number>
af,
<number box="[761,801,952,974]" pageId="83" pageNumber="84" value="2.3">2, 3</number>
,
<number box="[814,827,952,974]" pageId="83" pageNumber="84" value="4.0">4</number>
ac,
<number box="[876,888,952,974]" pageId="83" pageNumber="84" value="5.0">5</number>
(detailed synonymy).
<treatmentCitation author="Bosch" box="[1120,1350,952,974]" page="37" pageId="83" pageNumber="84" year="1995">
<bibRefCitation author="Bosch" box="[1120,1311,952,974]" pageId="83" pageNumber="84" refString="Bosch, D. T, Dance, S. P., Moolenbeek, R. G. &amp; Oliver, P. G. (1995) Seashells of Eastern Arabia. Motivate Publishing, Abu Dhabi, Dubai and London, 296 pp." type="book" year="1995">
Bosch
<emphasis box="[1191,1237,952,974]" italics="true" pageId="83" pageNumber="84">et al</emphasis>
.,
<number box="[1257,1311,952,974]" pageId="83" pageNumber="84" value="1995.0">1995</number>
</bibRefCitation>
:
<number box="[1323,1350,952,974]" pageId="83" pageNumber="84" value="37.0">37</number>
</treatmentCitation>
, fig.
<number box="[1404,1432,952,974]" pageId="83" pageNumber="84" value="52.0">52</number>
.
<treatmentCitation author="Zuschin" box="[199,441,982,1005]" page="99" pageId="83" pageNumber="84" year="2009">
<bibRefCitation author="Zuschin" box="[199,404,982,1005]" pageId="83" pageNumber="84" refString="Zuschin, M., Janssen, R. &amp; Baal, C. (2009) Gastropods and their habitats from the northern Red Sea (Egypt: Safaga). Part 1: Patellogastropoda, Vetigastropoda and Cycloneritimorpha. Annalen des Naturhistorischen Museums in Wien, 111 (A), 73 - 158." type="journal article" year="2009">
Zuschin
<emphasis box="[287,331,982,1005]" italics="true" pageId="83" pageNumber="84">et al</emphasis>
.,
<number box="[350,404,982,1004]" pageId="83" pageNumber="84" value="2009.0">2009</number>
</bibRefCitation>
:
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</treatmentCitation>
, pl.
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, figs
<number box="[554,594,982,1004]" pageId="83" pageNumber="84" value="2.5" valueMax="4.0" valueMin="1.0">14</number>
.
<treatmentCitation author="Bandel" box="[604,793,982,1004]" page="461" pageId="83" pageNumber="84" year="2010">
<bibRefCitation author="Bandel" box="[604,744,982,1004]" pageId="83" pageNumber="84" refString="Bandel, K. (2010) Relationships of the Triassic Eucycloidea Koken, 1897 (Mollusca, Gastropoda) to modern genera such as Pagodatrochus, Calliotropis and Euchelus, based on morphology of the early shell. Bulletin of Geosciences, 85, 435 - 486. http: // dx. doi. org / 10.3140 / bull. geosci. 1208" type="journal article" year="2010">
Bandel,
<number box="[689,744,982,1004]" pageId="83" pageNumber="84" value="2010.0">2010</number>
</bibRefCitation>
:
<number box="[753,793,982,1004]" pageId="83" pageNumber="84" value="461.0">461</number>
</treatmentCitation>
, fig.
<number box="[844,872,982,1004]" pageId="83" pageNumber="84" value="10.0">10</number>
.
<bibRefCitation author="Jay" box="[882,981,982,1004]" pageId="83" pageNumber="84" refString="Jay, M. (2014) Mollusques de l'ile de la Reunion. Version 24. Available from: http: // vieoceane. free. fr / mollusques / intro _ mollusque. htm (accessed 15 April 2015)" type="journal article" year="2014">
Jay,
<number box="[927,981,982,1004]" pageId="83" pageNumber="84" value="2014.0">2014</number>
</bibRefCitation>
.
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</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="distribution">
<paragraph blockId="83.[151,1436,1049,1362]" pageId="83" pageNumber="84">
<emphasis bold="true" box="[151,304,1049,1074]" pageId="83" pageNumber="84">Distribution.</emphasis>
Western Indian Ocean, from the Persian Gulf and Red Sea through East Africa, the
<collectingCountry box="[1283,1386,1050,1074]" name="Comoros" pageId="83" pageNumber="84">Comoros</collectingCountry>
and Mascarene Islands to the KwaZulu-Natal south coast (off Park Rynie); local material
<quantity box="[1133,1250,1085,1110]" metricMagnitude="2" metricUnit="m" metricValue="1.34" metricValueMax="2.5" metricValueMin="0.18" pageId="83" pageNumber="84" unit="m" value="134.0" valueMax="250.0" valueMin="18.0">18250 m</quantity>
, but mostly less than
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(living
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).
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<paragraph blockId="83.[151,1436,1049,1362]" pageId="83" pageNumber="84">
<emphasis bold="true" box="[199,273,1157,1182]" pageId="83" pageNumber="84">Notes.</emphasis>
The phylogenetic relationships of this genus are unclear.
<bibRefCitation author="Herbert" box="[932,1115,1157,1182]" pageId="83" pageNumber="84" refString="Herbert, D. G. (1989 b) Pagodatrochus, a new genus for Minolia variabilis H. Adams, 1873 (Gastropoda: Trochidae). Journal of Molluscan Studies, 55, 365 - 372. http: // dx. doi. org / 10.1093 / mollus / 55.3.365" type="journal article" year="1989" yearSuffix="b">
Herbert (
<number box="[1034,1092,1157,1182]" pageId="83" pageNumber="84" value="1989.0">1989</number>
b)
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tentatively referred it to the
<taxonomicName box="[151,276,1193,1218]" class="Gastropoda" family="Trochidae" kingdom="Animalia" pageId="83" pageNumber="84" phylum="Mollusca" rank="subFamily" subFamily="Gibbulinae">Gibbulinae</taxonomicName>
[=
<taxonomicName box="[304,470,1193,1218]" class="Gastropoda" family="Trochidae" kingdom="Animalia" pageId="83" pageNumber="84" phylum="Mollusca" rank="subFamily" subFamily="Cantharidinae">Cantharidinae</taxonomicName>
], but more recently
<bibRefCitation author="Bandel" box="[692,853,1193,1218]" pageId="83" pageNumber="84" refString="Bandel, K. (2010) Relationships of the Triassic Eucycloidea Koken, 1897 (Mollusca, Gastropoda) to modern genera such as Pagodatrochus, Calliotropis and Euchelus, based on morphology of the early shell. Bulletin of Geosciences, 85, 435 - 486. http: // dx. doi. org / 10.3140 / bull. geosci. 1208" type="journal article" year="2010">
Bandel (
<number box="[786,845,1194,1218]" pageId="83" pageNumber="84" value="2010.0">2010</number>
)
</bibRefCitation>
suggested that the genus was related to seguenzioid (eucycloid) taxa and created for it a new family, the Pagodatrochidae.
<bibRefCitation author="Herbert" box="[991,1163,1229,1254]" pageId="83" pageNumber="84" refString="Herbert, D. G. (2012) A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean. African Invertebrates, 53, 381 - 502. http: // dx. doi. org / 10.5733 / afin. 053.0209" type="journal article" year="2012">
Herbert (
<number box="[1096,1155,1230,1254]" pageId="83" pageNumber="84" value="2012.0">2012</number>
)
</bibRefCitation>
noted that while some characters of the shell and
<taxonomicName box="[477,547,1265,1290]" class="Gastropoda" family="Calliostomatidae" genus="Falsimargarita" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="species" species="radula">radula</taxonomicName>
of
<taxonomicName box="[596,766,1265,1290]" class="Gastropoda" family="Chilodontidae" genus="Pagodatrochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="83" pageNumber="84" phylum="Mollusca" rank="genus">
<emphasis box="[596,766,1265,1290]" italics="true" pageId="83" pageNumber="84">Pagodatrochus</emphasis>
</taxonomicName>
are indeed similar to those of chilodontid seguenzioids, features of the external anatomy are less consistent with such a relationship. In the absence of molecular data I consider it preferable to treat the genus as a vetigastropod taxon of uncertain affinity.
</paragraph>
</subSubSection>
</treatment>
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