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<document id="EE4B682E8E1C64C29F13CBFB05161995" ID-DOI="10.11646/zootaxa.3640.1.4" ID-GBIF-Dataset="359d8bed-26b9-408f-b6d2-44ca74cf47a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="283650" ID-ZooBank="3AE00A3A-694B-4343-BF7C-A31182C82CC7" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460592708495" checkinUser="plazi" docAuthor="Bell, Adam J. &amp; Cryan, Jason R." docDate="2013" docId="03CA87E78D38FF9DFF5FF8E7FA8FFC43" docLanguage="en" docName="zt03640p069.pdf" docOrigin="Zootaxa 3640 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kyphomachaerota maaia Bell &amp; Cryan, 2013, sp. n." docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="FFF3FF9F8D32FF96FFC8FF8EFF88FFE7" masterDocTitle="Two new genera of tube-making spittlebugs (Hemiptera: Machaerotidae: Enderleiniini)" masterLastPageNumber="69" masterPageNumber="57" pageNumber="67" updateTime="1698326262506" updateUser="plazi">
<mods:mods id="717742D258BFF3966ED66D8DA36BA103" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3200D2FC220D57D78918396953BB4E45">Two new genera of tube-making spittlebugs (Hemiptera: Machaerotidae: Enderleiniini)</mods:title>
</mods:titleInfo>
<mods:name id="029D3268CBC75A7989FED5370EB7B542" type="personal">
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<mods:namePart id="0B5BE653A8E302417B73DC38DD717C72">Bell, Adam J.</mods:namePart>
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<mods:role id="1EFFAED2C04AF8F3C46CFB94C4658E53">
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<mods:namePart id="50731178C45E84CAF74A175EA9F27508">Cryan, Jason R.</mods:namePart>
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<mods:title id="CF06901170A7908E1BD4D15C408F851C">Zootaxa</mods:title>
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<mods:part id="D89E85366387C8FC250D9FCEDD7F390E">
<mods:date id="E5C1C885423097B454B9165E9FAF8546">2013</mods:date>
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<mods:number id="08ED6BA960DE8BE48DFA23749FCED575">3640</mods:number>
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<mods:identifier id="090ACE7542BEB89D1E0C25588458CDBF" type="DOI">10.11646/zootaxa.3640.1.4</mods:identifier>
<mods:identifier id="9AE5A5B411BE417C4AC1D6183C1BEDB0" type="GBIF-Dataset">359d8bed-26b9-408f-b6d2-44ca74cf47a0</mods:identifier>
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<treatment id="03CA87E78D38FF9DFF5FF8E7FA8FFC43" ID-DOI="http://doi.org/10.5281/zenodo.6151773" ID-GBIF-Taxon="127663835" ID-Zenodo-Dep="6151773" LSID="urn:lsid:plazi:treatment:03CA87E78D38FF9DFF5FF8E7FA8FFC43" httpUri="http://treatment.plazi.org/id/03CA87E78D38FF9DFF5FF8E7FA8FFC43" lastPageId="11" lastPageNumber="68" pageId="10" pageNumber="67">
<subSubSection id="C379657A8D38FF9CFF5FF8E7FEAFF843" pageId="10" pageNumber="67" type="nomenclature">
<paragraph id="8BDC36F18D38FF9CFF5FF8E7FD98F864" blockId="10.[151,528,1897,1956]" box="[151,528,1897,1923]" pageId="10" pageNumber="67">
<heading id="D094819D8D38FF9CFF5FF8E7FD98F864" bold="true" box="[151,528,1897,1923]" fontSize="11" level="1" pageId="10" pageNumber="67" reason="1">
<emphasis id="B917EAE38D38FF9CFF5FF8E7FD98F864" bold="true" box="[151,528,1897,1923]" pageId="10" pageNumber="67">
<taxonomicName id="4C634D728D38FF9CFF5FF8E7FE4EF864" ID-CoL="6NQCL" box="[151,454,1897,1923]" class="Insecta" family="Machaerotidae" genus="Kyphomachaerota" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="67" phylum="Arthropoda" rank="species" species="maaia" status="sp. nov.">
<emphasis id="B917EAE38D38FF9CFF5FF8E7FE4EF864" bold="true" box="[151,454,1897,1923]" italics="true" pageId="10" pageNumber="67">Kyphomachaerota maaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457988D38FF9CFE05F8E7FD98F864" box="[461,528,1897,1923]" pageId="10" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36F18D38FF9CFF5FF802FEAFF843" blockId="10.[151,528,1897,1956]" box="[151,295,1932,1956]" pageId="10" pageNumber="67">
(
<figureCitation id="13582A748D38FF9CFF57F802FE97F843" box="[159,287,1932,1956]" captionStart-0="FIGURES 29 32" captionStart-1="FIGURES 33 34" captionStart-2="FIGURES 35 41" captionStartId-0="8.[151,264,1957,1980]" captionStartId-1="9.[151,264,1838,1861]" captionStartId-2="10.[151,264,1738,1761]" captionTargetBox-0="[183,1417,287,1929]" captionTargetBox-1="[151,1435,185,1837]" captionTargetBox-2="[151,1435,193,1711]" captionTargetId-0="figure@8.[183,1420,287,1936]" captionTargetId-1="figure@9.[151,1436,185,1837]" captionTargetId-2="figure@10.[151,1436,193,1717]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 29 32. General body features of Kyphomachaerota maaia sp. n. (holotype). 29. Head and pronotum, dorsal view; 30. Head, front view; 31. Lateral habitus; 32. Dorsal habitus." captionText-1="FIGURES 33 34. Wing features of Kyphomachaerota maaia sp. n. (holotype). 33. Forewing (tegmen); 34. Hindwing." captionText-2="FIGURES 35 41. Genitalic features of Kyphomachaerota maaia sp. n. (holotype). 35. Pygofer, lateral view; 36. Pygofer, posterior view; 37. Aedeagus, lateral view; 38. Aedeagus, dorsal view; 39. Parameres, dorsal view; 40. Paramere, external view; 41. Paramere, internal view." httpUri-0="https://zenodo.org/record/283654/files/figure.png" httpUri-1="https://zenodo.org/record/283655/files/figure.png" httpUri-2="https://zenodo.org/record/283656/files/figure.png" pageId="10" pageNumber="67">Figs 2941</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D38FF9CFF5FF85BFAACF80B" box="[151,1316,2004,2029]" pageId="10" pageNumber="67" type="materials_examined">
<paragraph id="8BDC36F18D38FF9CFF5FF85BFAACF80B" blockId="10.[151,1316,2004,2029]" box="[151,1316,2004,2029]" pageId="10" pageNumber="67">
<emphasis id="B917EAE38D38FF9CFF5FF85BFEC8F80A" bold="true" box="[151,320,2004,2029]" pageId="10" pageNumber="67">
<typeStatus id="54D888538D38FF9CFF5FF85BFF58F80A" box="[151,208,2005,2029]" pageId="10" pageNumber="67">Type</typeStatus>
Locality.
</emphasis>
Gunung Mulu World Heritage Area (
<geoCoordinate id="EE5750368D38FF9CFD2DF85AFC0AF80B" box="[741,898,2004,2028]" direction="north" orientation="latitude" pageId="10" pageNumber="67" precision="1" value="4.042444">N04°02'32.8&quot;</geoCoordinate>
,
<geoCoordinate id="EE5750368D38FF9CFC45F85AFBBBF80B" box="[909,1075,2004,2028]" direction="east" orientation="longitude" pageId="10" pageNumber="67" precision="1" value="114.81378">E114°48'49.6&quot;</geoCoordinate>
),
<collectingCountry id="F37476618D38FF9CFB8FF85AFB22F80B" box="[1095,1194,2004,2028]" name="Malaysia" pageId="10" pageNumber="67">Sarawak</collectingCountry>
,
<collectingCountry id="F37476618D38FF9CFB7EF85AFAA8F80B" box="[1206,1312,2004,2028]" name="Malaysia" pageId="10" pageNumber="67">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFF19FCE8FF10" pageId="11" pageNumber="68" type="diagnosis">
<paragraph id="8BDC36F18D39FF9DFF0FFF19FCE8FF10" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFF19FEC9FF57" bold="true" box="[199,321,151,176]" pageId="11" pageNumber="68">Diagnosis.</emphasis>
Head pale yellow-green, basal edge of postclypeus, anteclypeus, lorum and basal half maxillary plate dark brown; prothorax yellow green, posterolateral angle dark brown; scutellum dark brown; pro- and mesothoracic legs dark brown, metathoracic legs lighter brown.
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFE8DFB3AFD63" pageId="11" pageNumber="68" type="description">
<paragraph id="8BDC36F18D39FF9DFF0FFE8DFC5FFDDB" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFE8DFED1FEFB" bold="true" box="[199,345,259,284]" pageId="11" pageNumber="68">Description.</emphasis>
<emphasis id="B917EAE38D39FF9DFE97FE8AFE14FEFB" box="[351,412,260,284]" italics="true" pageId="11" pageNumber="68">Head</emphasis>
(
<figureCitation id="13582A748D39FF9DFE62FE8AFDA7FEFB" box="[426,559,260,284]" captionStart="FIGURES 29 32" captionStartId="8.[151,264,1957,1980]" captionTargetBox="[183,1417,287,1929]" captionTargetId="figure@8.[183,1420,287,1936]" captionTargetPageId="8" captionText="FIGURES 29 32. General body features of Kyphomachaerota maaia sp. n. (holotype). 29. Head and pronotum, dorsal view; 30. Head, front view; 31. Lateral habitus; 32. Dorsal habitus." httpUri="https://zenodo.org/record/283654/files/figure.png" pageId="11" pageNumber="68">Figs 2932</figureCitation>
): bright yellow-green; eyes yellow green; face yellow; basal ¼ of postclypeus, anteclypeus, lorum, and basal ½ of maxillary plate dark brown; vertex smooth, lacking median carina.
<emphasis id="B917EAE38D39FF9DFAC3FEA6FAD4FEA7" box="[1291,1372,296,320]" italics="true" pageId="11" pageNumber="68">Thorax</emphasis>
(
<figureCitation id="13582A748D39FF9DFAA2FEA9FF6AFE83" captionStart-0="FIGURES 29 32" captionStart-1="FIGURES 33 34" captionStartId-0="8.[151,264,1957,1980]" captionStartId-1="9.[151,264,1838,1861]" captionTargetBox-0="[183,1417,287,1929]" captionTargetBox-1="[151,1435,185,1837]" captionTargetId-0="figure@8.[183,1420,287,1936]" captionTargetId-1="figure@9.[151,1436,185,1837]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 29 32. General body features of Kyphomachaerota maaia sp. n. (holotype). 29. Head and pronotum, dorsal view; 30. Head, front view; 31. Lateral habitus; 32. Dorsal habitus." captionText-1="FIGURES 33 34. Wing features of Kyphomachaerota maaia sp. n. (holotype). 33. Forewing (tegmen); 34. Hindwing." httpUri-0="https://zenodo.org/record/283654/files/figure.png" httpUri-1="https://zenodo.org/record/283655/files/figure.png" pageId="11" pageNumber="68">Figs 2934</figureCitation>
): pronotum bright yellow-green; dark brown coloration wraps around body to frontal edge of scutellum encompassing posterolateral angle of pronotum; legs light to dark brown; tegmen apically hyaline, becoming increasingly opaque along basal half; veins strongly pigmented, flanked by dark punctures; anal veins fused medially.
<emphasis id="B917EAE38D39FF9DFEC3FE36FEFDFE37" box="[267,373,440,464]" italics="true" pageId="11" pageNumber="68">Genitalia</emphasis>
:
<emphasis id="B917EAE38D39FF9DFE42FE36FE4CFE37" box="[394,452,440,464]" italics="true" pageId="11" pageNumber="68">Male</emphasis>
: pygofer (
<figureCitation id="13582A748D39FF9DFD8DFE39FD43FE28" box="[581,715,439,463]" captionStart="FIGURES 35 41" captionStartId="10.[151,264,1738,1761]" captionTargetBox="[151,1435,193,1711]" captionTargetId="figure@10.[151,1436,193,1717]" captionTargetPageId="10" captionText="FIGURES 35 41. Genitalic features of Kyphomachaerota maaia sp. n. (holotype). 35. Pygofer, lateral view; 36. Pygofer, posterior view; 37. Aedeagus, lateral view; 38. Aedeagus, dorsal view; 39. Parameres, dorsal view; 40. Paramere, external view; 41. Paramere, internal view." httpUri="https://zenodo.org/record/283656/files/figure.png" pageId="11" pageNumber="68">Figs 3536</figureCitation>
) lightly sclerotized, consisting of upper and lower sclerites, scabrous area projecting slightly from apical sides; subgenital plates (
<figureCitation id="13582A748D39FF9DFC6BFE52FBABFE13" box="[931,1059,476,500]" captionStart="FIGURES 35 41" captionStartId="10.[151,264,1738,1761]" captionTargetBox="[151,1435,193,1711]" captionTargetId="figure@10.[151,1436,193,1717]" captionTargetPageId="10" captionText="FIGURES 35 41. Genitalic features of Kyphomachaerota maaia sp. n. (holotype). 35. Pygofer, lateral view; 36. Pygofer, posterior view; 37. Aedeagus, lateral view; 38. Aedeagus, dorsal view; 39. Parameres, dorsal view; 40. Paramere, external view; 41. Paramere, internal view." httpUri="https://zenodo.org/record/283656/files/figure.png" pageId="11" pageNumber="68">Figs 3536</figureCitation>
) rounded at apex; aedeagus (
<figureCitation id="13582A748D39FF9DFAA2FE52FF56FDF0" captionStart="FIGURES 35 41" captionStartId="10.[151,264,1738,1761]" captionTargetBox="[151,1435,193,1711]" captionTargetId="figure@10.[151,1436,193,1717]" captionTargetPageId="10" captionText="FIGURES 35 41. Genitalic features of Kyphomachaerota maaia sp. n. (holotype). 35. Pygofer, lateral view; 36. Pygofer, posterior view; 37. Aedeagus, lateral view; 38. Aedeagus, dorsal view; 39. Parameres, dorsal view; 40. Paramere, external view; 41. Paramere, internal view." httpUri="https://zenodo.org/record/283656/files/figure.png" pageId="11" pageNumber="68">Figs 3738</figureCitation>
) cylindrical, heavily scaled dorsally and ventrally; parameres (
<figureCitation id="13582A748D39FF9DFC74FE71FBC8FDF0" box="[956,1088,511,535]" captionStart="FIGURES 35 41" captionStartId="10.[151,264,1738,1761]" captionTargetBox="[151,1435,193,1711]" captionTargetId="figure@10.[151,1436,193,1717]" captionTargetPageId="10" captionText="FIGURES 35 41. Genitalic features of Kyphomachaerota maaia sp. n. (holotype). 35. Pygofer, lateral view; 36. Pygofer, posterior view; 37. Aedeagus, lateral view; 38. Aedeagus, dorsal view; 39. Parameres, dorsal view; 40. Paramere, external view; 41. Paramere, internal view." httpUri="https://zenodo.org/record/283656/files/figure.png" pageId="11" pageNumber="68">Figs 3941</figureCitation>
) with short styli and rounded medially with an apical lobe with setae on dorsal edge.
<emphasis id="B917EAE38D39FF9DFCCCFDAAFCD0FDDB" box="[772,856,548,572]" italics="true" pageId="11" pageNumber="68">Female</emphasis>
: unknown.
</paragraph>
<paragraph id="8BDC36F18D39FF9DFF0FFDC6FB3AFD63" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFDC6FE45FD87" bold="true" box="[199,461,583,608]" pageId="11" pageNumber="68">Measurements (mm).</emphasis>
Body length: male 5.0; pronotal length: male 1.51; forewing length: male 4.5; maximum width of head, including eyes: male 1.54; width between humeral angles: male 2.2.
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFD01FB84FD4F" box="[199,1036,655,680]" pageId="11" pageNumber="68" type="distribution">
<paragraph id="8BDC36F18D39FF9DFF0FFD01FB84FD4F" blockId="11.[151,1436,151,932]" box="[199,1036,655,680]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFD01FEE8FD4F" bold="true" box="[199,352,655,680]" pageId="11" pageNumber="68">Distribution.</emphasis>
<collectingCountry id="F37476618D39FF9DFEAFFD01FE75FD40" box="[359,509,655,679]" name="Malaysia" pageId="11" pageNumber="68">MALAYSIA</collectingCountry>
:
<collectingCountry id="F37476618D39FF9DFDCFFD01FDE2FD40" box="[519,618,655,679]" name="Malaysia" pageId="11" pageNumber="68">Sarawak</collectingCountry>
, Gunung Mulu World Heritage Area.
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFD3AFC2FFD08" pageId="11" pageNumber="68" type="discussion">
<paragraph id="8BDC36F18D39FF9DFF0FFD3AFC2FFD08" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFD3AFEB3FD2B" bold="true" box="[199,315,692,716]" pageId="11" pageNumber="68">Remarks.</emphasis>
While
<taxonomicName id="4C634D728D39FF9DFE5BFD3BFE75FD2B" box="[403,509,693,716]" class="Insecta" family="Machaerotidae" genus="Kyphomachaerota" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="maaia">
<emphasis id="B917EAE38D39FF9DFE5BFD3BFE75FD2B" box="[403,509,693,716]" italics="true" pageId="11" pageNumber="68">K. maaia</emphasis>
</taxonomicName>
may superficially resemble species of
<taxonomicName id="4C634D728D39FF9DFC0CFD3AFBB9FD2B" box="[964,1073,692,716]" class="Insecta" family="Machaerotidae" genus="Taihorina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAE38D39FF9DFC0CFD3AFBB9FD2B" box="[964,1073,692,716]" italics="true" pageId="11" pageNumber="68">Taihorina</emphasis>
</taxonomicName>
, it differs in the male genitalia (lacking the apical process) and the forewing venation (being visible).
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFD72FECDFCD0" pageId="11" pageNumber="68" type="etymology">
<paragraph id="8BDC36F18D39FF9DFF0FFD72FECDFCD0" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFD72FEC6FCF3" bold="true" box="[199,334,764,788]" pageId="11" pageNumber="68">Etymology.</emphasis>
This species is named in honor of Tsing-Chao Maa, for his important work on the systematics of
<taxonomicName id="4C634D728D39FF9DFF5FFC91FEC9FCD0" box="[151,321,799,823]" class="Insecta" family="Machaerotidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="family">Machaerotidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C379657A8D39FF9DFF0FFCCDFA8FFC43" pageId="11" pageNumber="68" type="materials_examined">
<paragraph id="8BDC36F18D39FF9DFF0FFCCDFA8FFC43" blockId="11.[151,1436,151,932]" pageId="11" pageNumber="68">
<emphasis id="B917EAE38D39FF9DFF0FFCCDFE3FFCBB" bold="true" box="[199,439,835,860]" pageId="11" pageNumber="68">Material Examined.</emphasis>
<typeStatus id="54D888538D39FF9DFE76FCCAFDAEFCBB" box="[446,550,836,860]" pageId="11" pageNumber="68" type="holotype">Holotype</typeStatus>
3 (dissected, DNA extracted, deposited at USNM), “
<collectingCountry id="F37476618D39FF9DFB4AFCCAFA92FCBB" box="[1154,1306,836,860]" name="Malaysia" pageId="11" pageNumber="68">MALAYSIA</collectingCountry>
:
<collectingCountry id="F37476618D39FF9DFAECFCCAFA00FCBB" box="[1316,1416,836,860]" name="Malaysia" pageId="11" pageNumber="68">Sarawak</collectingCountry>
; / Gunung Mulu National Park; / Park HQ /
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<geoCoordinate id="EE5750368D39FF9DFCDDFCE6FC37FC98" box="[789,959,871,895]" direction="east" orientation="longitude" pageId="11" pageNumber="68" precision="1" value="114.81378">E 114°48'49.6&quot;</geoCoordinate>
/
<date id="FFDD10318D39FF9DFC1DFCE9FB44FC98" box="[981,1228,871,895]" pageId="11" pageNumber="68" value="2009-06-30" valueMax="2009-07-28" valueMin="2009-06-30">30 Jun 28 July 2009</date>
/ A.J. Bell &amp; J.M. Urban”, “CER379 / JCRC
<date id="FFDD10318D39FF9DFE0CFC02FDE8FC43" box="[452,608,908,932]" pageId="11" pageNumber="68" value="1965-09-01" valueMax="1965-09-15" valueMin="1965-09-01">09011565</date>
”; “
<typeStatus id="54D888538D39FF9DFD4BFC02FC91FC43" box="[643,793,908,932]" pageId="11" pageNumber="68" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C634D728D39FF9DFCE7FC02FBCEFC43" box="[815,1094,908,932]" class="Insecta" family="Machaerotidae" genus="Kyphomachaerota" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="maaia">Kyphomachaerota maaia</taxonomicName>
/ Bell &amp; Cryan”.
</paragraph>
</subSubSection>
</treatment>
</document>