treatments-xml/data/03/CA/87/03CA87D8FFCEFFE3FF0BFB07FC58FD57.xml

<document id="07622C8FCF818BE23B9E93EB5A5BF091" ID-CLB-Dataset="31855" ID-DOI="10.11646/zootaxa.4331.1.1" ID-GBIF-Dataset="14c29731-224f-43a4-ae52-f76823177ce5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1009725" ID-ZooBank="FB50725E-4D03-4507-9CAF-EB939C7F548F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507876567978" checkinUser="plazi" docAuthor="Asami, Takahiro" docDate="2017" docId="03CA87D8FFCEFFE3FF0BFB07FC58FD57" docLanguage="en" docName="zootaxa.4331.1.1.pdf" docOrigin="Zootaxa 4331 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dicharax draco Asami 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FFF3FFA0FFFDFFD6FF9CFFC0FF80FFFF" masterDocTitle="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus" masterLastPageNumber="124" masterPageNumber="1" pageNumber="52" updateTime="1698469440756" updateUser="ExternalLinkService">
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<mods:title id="5CFF5E5029F42FDE43415446E8AD6BA6">Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus</mods:title>
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<mods:date id="AAE6C33EF7F7C5CC6CAF22D00F7B010E">2017</mods:date>
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<treatment id="03CA87D8FFCEFFE3FF0BFB07FC58FD57" ID-DOI="http://doi.org/10.5281/zenodo.6004434" ID-GBIF-Taxon="135435132" ID-Zenodo-Dep="6004434" LSID="urn:lsid:plazi:treatment:03CA87D8FFCEFFE3FF0BFB07FC58FD57" httpUri="http://treatment.plazi.org/id/03CA87D8FFCEFFE3FF0BFB07FC58FD57" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
<subSubSection id="C3796545FFCEFFE5FF0BFB07FE9FFAFC" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8BDC36CEFFCEFFE5FF0BFB07FD58FB1D" blockId="51.[151,728,1223,1283]" box="[151,728,1223,1250]" pageId="51" pageNumber="52">
<heading id="D09481A2FFCEFFE5FF0BFB07FD58FB1D" bold="true" box="[151,728,1223,1250]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B917EADCFFCEFFE5FF0BFB07FD58FB1D" bold="true" box="[151,728,1223,1250]" pageId="51" pageNumber="52">
<taxonomicName id="4C634D4DFFCEFFE5FF0BFB07FD0EFB1D" ID-CoL="35L5Q" authority="Pall-Gergely &amp; Hunyadi" authorityName="Asami" authorityYear="2017" box="[151,654,1223,1250]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFCEFFE5FF0BFB07FED4FB1E" bold="true" box="[151,340,1223,1249]" italics="true" pageId="51" pageNumber="52">Dicharax draco</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFCEFFE5FD09FB08FD58FB1D" box="[661,728,1224,1250]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDC36CEFFCEFFE5FF0BFB2BFE9FFAFC" blockId="51.[151,728,1223,1283]" box="[151,287,1259,1283]" pageId="51" pageNumber="52">
<figureCitation id="13582A4BFFCEFFE5FF0BFB2BFE9BFAFC" box="[151,283,1259,1283]" captionStart="FIGURE 34" captionStartId="52.[151,250,1822,1844]" captionTargetBox="[151,1436,193,1801]" captionTargetId="figure@52.[151,1436,193,1801]" captionTargetPageId="52" captionText="FIGURE 34. Shells of single site endemic Dicharax species. A. Dicharax draco Páll-Gergely &amp; Hunyadi n. sp. (holotype, HNHM 99705). B. Dicharax imitator Páll-Gergely &amp; Hunyadi n. sp. (holotype, HNHM 99706). C. Dicharax microcostatus Páll-Gergely n. sp. (holotype, HNHM 99708). D. Dicharax micropolitus Páll-Gergely &amp; Hunyadi, n. sp. (holotype, HNHM 99709). Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009793/files/figure.png" pageId="51" pageNumber="52">FIGUrE 34A</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFCEFFE5FF0BFAEFFC20FA6F" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BDC36CEFFCEFFE5FF0BFAEFFC20FA6F" blockId="51.[151,1437,1327,2037]" pageId="51" pageNumber="52">
<materialsCitation id="3B0B3C93FFCEFFE5FF0BFAEFFC1CFA6F" ID-GBIF-Occurrence="1671715194" collectingDate="2006-01-09" collectionCode="COLL, OKA" collectorName="JUTTING" country="China" elevation="1611" latitude="23.75903" location="Ex" longLatPrecision="1" longitude="104.99259" pageId="51" pageNumber="52" specimenCode="HNHM 99705" specimenCount="5" stateProvince="YUNNaN" typeStatus="holotype">
<emphasis id="B917EADCFFCEFFE5FF0BFAEFFEC7FAB7" bold="true" box="[151,327,1327,1352]" pageId="51" pageNumber="52">
<typeStatus id="54D8886CFFCEFFE5FF0BFAEFFF52FAB7" box="[151,210,1327,1352]" pageId="51" pageNumber="52">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D8886CFFCEFFE5FECCFAF0FE38FAB7" box="[336,440,1328,1352]" pageId="51" pageNumber="52" type="holotype">HOLOTypE</typeStatus>
<specimenCode id="DBC59EB5FFCEFFE5FE5EFAF0FDEAFAB7" box="[450,618,1327,1352]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="51" pageNumber="52">HNHM 99705</specimenCode>
, 
<location id="8EBC6015FFCEFFE5FDEBFAF0FD11FAB8" LSID="urn:lsid:plazi:treatment:03CA87D8FFCEFFE3FF0BFB07FC58FD57:8EBC6015FFCEFFE5FDEBFAF0FD11FAB8" box="[631,657,1328,1351]" country="China" latitude="23.75903" longLatPrecision="1" longitude="104.99259" name="Ex" pageId="51" pageNumber="52" stateProvince="YUNNaN">Ex</location>
<collectionCode id="ED72AE0BFFCEFFE5FD07FAF0FD4AFAB7" box="[667,714,1328,1352]" pageId="51" pageNumber="52">COLL</collectionCode>
. K. OHara, OK/3 
<typeStatus id="54D8886CFFCEFFE5FC0BFAF0FB86FAB7" box="[919,1030,1328,1352]" pageId="51" pageNumber="52" type="paratype">paraTypEs</typeStatus>
, 
<collectionCode id="ED72AE0BFFCEFFE5FB8FFAF0FBD4FAB7" box="[1043,1108,1328,1352]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15165" name="Oksky State Biosphere Reserve" pageId="51" pageNumber="52">OKA</collectionCode>
/1 
<typeStatus id="54D8886CFFCEFFE5FBF2FAF0FB52FAB7" box="[1134,1234,1328,1352]" pageId="51" pageNumber="52" type="paratype">paraTypE</typeStatus>
, PGB/1 
<typeStatus id="54D8886CFFCEFFE5FAA9FAF0FA19FAB7" box="[1333,1433,1328,1352]" pageId="51" pageNumber="52" type="paratype">paraTypE</typeStatus>
, 
<emphasis id="B917EADCFFCEFFE5FF0BFA94FE9BFA92" bold="true" box="[151,283,1364,1389]" pageId="51" pageNumber="52">20060109A</emphasis>
<collectingCountry id="F374765EFFCEFFE5FEB8FA94FEEAFA93" box="[292,362,1364,1388]" name="China" pageId="51" pageNumber="52">CHINa</collectingCountry>
, 
<collectingRegion id="49A7F82CFFCEFFE5FEEBFA95FE50FA93" box="[375,464,1365,1388]" country="China" name="Yunnan" pageId="51" pageNumber="52">YUNNaN</collectingRegion>
, WENsHaNzHUaNG ZUmIaOzU ZIzHIzHO, GUaNGNaN XIaN, LIjI, 
<elevation id="004ED1FDFFCEFFE5FB1AFA94FB5EFA93" box="[1158,1246,1364,1389]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="51" pageNumber="52" unit="m" value="1611.0">
<quantity id="4C9B9B2BFFCEFFE5FB1AFA94FB5EFA93" box="[1158,1246,1364,1389]" metricMagnitude="3" metricUnit="m" metricValue="1.611" pageId="51" pageNumber="52" unit="m" value="1611.0">1611 m</quantity>
</elevation>
, 
<geoCoordinate id="EE575009FFCEFFE5FB70FA94FA18FA92" box="[1260,1432,1364,1389]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="1" value="23.75903">23°45.54175'N</geoCoordinate>
, 
<geoCoordinate id="EE575009FFCEFFE5FF0BFAB7FECEFA6F" box="[151,334,1399,1424]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="1" value="104.99259">104°59.55567'E</geoCoordinate>
, LEG. K. OHara, K. OkUbO &amp; 
<collectorName id="26965318FFCEFFE5FD0CFAB8FD3DFA70" box="[656,701,1400,1423]" name="JUTTING" pageId="51" pageNumber="52">J.U.</collectorName>
OTaNI, 0 
<collectingDate id="EF99E9E6FFCEFFE5FC83FAB8FC1CFA6F" box="[799,924,1399,1424]" pageId="51" pageNumber="52" value="2006-01-09">
<date id="FFDD100EFFCEFFE5FC83FAB8FC1CFA6F" box="[799,924,1399,1424]" pageId="51" pageNumber="52" value="2006-01-09">9 JaN 2006</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFCEFFE5FF5BFA5CFEC4FA27" pageId="51" pageNumber="52" type="etymology">
<paragraph id="8BDC36CEFFCEFFE5FF5BFA5CFEC4FA27" blockId="51.[151,1437,1327,2037]" pageId="51" pageNumber="52">
<emphasis id="B917EADCFFCEFFE5FF5BFA5CFECEFA4A" bold="true" box="[199,334,1436,1461]" pageId="51" pageNumber="52">Etymology.</emphasis>
THE apErTUrE Of THIs NEw spECIEs rEsEmbLEs a bITING mOUTH Of a draGON (
<taxonomicName id="4C634D4DFFCEFFE5FB1EFA5CFB43FA4A" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[1154,1219,1436,1461]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="111" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">draCO</taxonomicName>
IN LaTIN). DraCO Is UsEd as a NOUN.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFCEFFE5FF5BFA24FC97F9DF" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BDC36CEFFCEFFE5FF5BFA24FC97F9DF" blockId="51.[151,1437,1327,2037]" pageId="51" pageNumber="52">
<emphasis id="B917EADCFFCEFFE5FF5BFA24FEC1FA02" bold="true" box="[199,321,1508,1533]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
A smaLL spECIEs wITH sOmEwHaT ELEvaTEd spIrE, sHOrT R2 aNd R3, TypICaL 
<taxonomicName id="4C634D4DFFCEFFE5FBE9FA24FB5FFA03" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[1141,1247,1508,1532]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="51" pageNumber="52" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFCEFFE5FBE9FA24FB5FFA03" box="[1141,1247,1508,1532]" italics="true" pageId="51" pageNumber="52">Dicharax</emphasis>
</taxonomicName>
- 
<typeStatus id="54D8886CFFCEFFE5FB79FA25FA99FA02" box="[1253,1305,1509,1533]" pageId="51" pageNumber="52">TypE</typeStatus>
rIbs ON R2, sHarp swELLING ON R3, aNd sTrONGLy CrENULaTEd pErIsTOmE.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFCEFFE3FF5BF9ECFF65FF08" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="description">
<paragraph id="8BDC36CEFFCEFFE5FF5BF9ECFA1BF82F" blockId="51.[151,1437,1327,2037]" pageId="51" pageNumber="52">
<emphasis id="B917EADCFFCEFFE5FF5BF9ECFED9F9BA" bold="true" box="[199,345,1580,1605]" pageId="51" pageNumber="52">Description.</emphasis>
SHELL (
<figureCitation id="13582A4BFFCEFFE5FE37F9ECFDB3F9BB" box="[427,563,1580,1604]" captionStart="FIGURE 34" captionStartId="52.[151,250,1822,1844]" captionTargetBox="[151,1436,193,1801]" captionTargetId="figure@52.[151,1436,193,1801]" captionTargetPageId="52" captionText="FIGURE 34. Shells of single site endemic Dicharax species. A. Dicharax draco Páll-Gergely &amp; Hunyadi n. sp. (holotype, HNHM 99705). B. Dicharax imitator Páll-Gergely &amp; Hunyadi n. sp. (holotype, HNHM 99706). C. Dicharax microcostatus Páll-Gergely n. sp. (holotype, HNHM 99708). D. Dicharax micropolitus Páll-Gergely &amp; Hunyadi, n. sp. (holotype, HNHM 99709). Scale represents 5 mm. All photos: B. Páll-Gergely." httpUri="https://zenodo.org/record/1009793/files/figure.png" pageId="51" pageNumber="52">FIGUrE 34A</figureCitation>
) LIGHT GrEy; sHELL OUTLINE rOUNdEd frOm dOrsaL vIEw, spIrE ELEvaTEd, EdGE Of bOdy wHOrL rOUNdEd; aLL ExamINEd spECImENs wErE sOmEwHaT wEaTHErEd, THErEfOrE THEIr OrIGINaL COLOUr COULd NOT bE ObsErvEd; prOTOCONCH CONsIsTs Of 1.75–2 wHOrLs (N = 2), LOw, raTHEr GLOssy; R1 CONsIsTs Of apprOxImaTELy 1.75 wHOrLs (N = 2), wITH LOw, rEGULar rIbs, vIsIbLE ON EdGE Of wHOrLs as wELL; rIbbING wEakEr aT bEGINNING Of R1 (HErE aLmOsT smOOTH), bUT sTrONGEr aT ENd Of R1; bOUNdary bETwEEN R1 aNd R2 CONspICUOUs bECaUsE Of dENsE R2 rIbs; R2 wITH rIbs CUrvEd TOwards apErTUrE; rIbs NEarLy rEaCH EaCH OTHEr, fOrmING aN aLmOsT smOOTH sUrfaCE; CrOss-sECTIONaL vIEw Of R2 COULd NOT bE ExamINEd, bUT prObabLy sImILar TO THOsE Of OTHEr spECIEs HavING CUrvEd rIbs; R2 aNd R3 vEry sHOrT, apprOxImaTELy 90 dEGrEEs COmbINEd; R2 aNd R3 Of abOUT THE samE LENGTH; sCULpTUrE Of R3 sImILar TO THaT Of R1, THErEfOrE bOUNdary bETwEEN R2 aNd R3 CONspICUOUs, INdICaTEd by a dEEp CONsTrICTION; mIddLE Of R3 wITH a prOmINENT, sHarp swELLING; apErTUrE ExTErNaLLy sLIGHTLy, INTErNaLLy vEry sTrONGLy CrENULaTEd by HavING fOUr TrIaNGULar swELLINGs ExTENdING bEyONd THE apErTUrE; THEsE fOUr swELLINGs dETErmINE fIvE LObEs; OUTEr LIp Of THE pErIsTOmE THIN, sHarp, ExpaNdEd aNd sLIGHTLy rEfLECTEd, EspECIaLLy IN dIrECTION Of UmbILICUs; UmbILICUs vEry NarrOw.
</paragraph>
<paragraph id="8BDC36CEFFCEFFE5FF5BF81CFC65F80B" blockId="51.[151,1437,1327,2037]" box="[199,997,2012,2037]" pageId="51" pageNumber="52">
<emphasis id="B917EADCFFCEFFE5FF5BF81CFE63F80A" bold="true" box="[199,483,2012,2037]" pageId="51" pageNumber="52">Measurements (in mm).</emphasis>
D = 3.1–3.3, H = 2.5–2.7 (N = 2, 
<typeStatus id="54D8886CFFCEFFE5FCC1F81DFC0DF80A" box="[861,909,2013,2037]" pageId="51" pageNumber="52">TypE</typeStatus>
sErIEs).
</paragraph>
<caption id="DF1C6646FFC9FFE2FF0BF8DEFD78F86F" httpUri="https://zenodo.org/record/1009793/files/figure.png" pageId="52" pageNumber="53" targetBox="[151,1436,193,1801]" targetPageId="52">
<paragraph id="8BDC36CEFFC9FFE2FF0BF8DEFD78F86F" blockId="52.[151,1436,1822,1936]" pageId="52" pageNumber="53">
<emphasis id="B917EADCFFC9FFE2FF0BF8DEFEA2F8CB" bold="true" box="[151,290,1822,1844]" pageId="52" pageNumber="53">FIGURE 34.</emphasis>
Shells of single site endemic 
<taxonomicName id="4C634D4DFFC9FFE2FDC1F8DEFD3DF8CB" authorityName="Kobelt &amp; Mollendorff" authorityYear="1900" box="[605,701,1822,1844]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="B917EADCFFC9FFE2FDC1F8DEFD3DF8CB" box="[605,701,1822,1844]" italics="true" pageId="52" pageNumber="53">Dicharax</emphasis>
</taxonomicName>
species. 
<emphasis id="B917EADCFFC9FFE2FC82F8DEFCB6F8CB" bold="true" box="[798,822,1822,1844]" pageId="52" pageNumber="53">A.</emphasis>
<taxonomicName id="4C634D4DFFC9FFE2FCA3F8DEFB68F8CB" authority="Pall-Gergely &amp; Hunyadi" authorityName="Asami" authorityYear="2017" box="[831,1256,1822,1844]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC9FFE2FCA3F8DEFC62F8CB" box="[831,994,1822,1844]" italics="true" pageId="52" pageNumber="53">Dicharax draco</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC9FFE2FB6DF8DEFAACF8CB" box="[1265,1324,1822,1844]" pageId="52" pageNumber="53" rank="species">
<emphasis id="B917EADCFFC9FFE2FB6DF8DEFAACF8CB" bold="true" box="[1265,1324,1822,1844]" pageId="52" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
(holotype, HNHM 99705). 
<emphasis id="B917EADCFFC9FFE2FEDFF8FDFEDAF8AC" bold="true" box="[323,346,1853,1875]" pageId="52" pageNumber="53">B.</emphasis>
<taxonomicName id="4C634D4DFFC9FFE2FEFEF8FDFCA0F8AC" authority="Pall-Gergely &amp; Hunyadi" authorityName="Asami" authorityYear="2017" box="[354,800,1853,1875]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="imitator" status="sp. nov.">
<emphasis id="B917EADCFFC9FFE2FEFEF8FDFD9CF8AC" box="[354,540,1853,1875]" italics="true" pageId="52" pageNumber="53">Dicharax imitator</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC9FFE2FCB4F8FDFCE3F8AC" box="[808,867,1853,1875]" pageId="52" pageNumber="53" rank="species">
<emphasis id="B917EADCFFC9FFE2FCB4F8FDFCE3F8AC" bold="true" box="[808,867,1853,1875]" pageId="52" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
(holotype, HNHM 99706). 
<emphasis id="B917EADCFFC9FFE2FB1AF8FDFB1FF8AC" bold="true" box="[1158,1183,1853,1875]" pageId="52" pageNumber="53">C.</emphasis>
<taxonomicName id="4C634D4DFFC9FFE2FB3BF8FDFE98F88E" authority="Pall-Gergely" authorityName="Asami" authorityYear="2017" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="microcostatus" status="sp. nov.">
<emphasis id="B917EADCFFC9FFE2FB3BF8FDFA1CF8AC" box="[1191,1436,1853,1875]" italics="true" pageId="52" pageNumber="53">Dicharax microcostatus</emphasis>
Páll-Gergely
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC9FFE2FEBCF89BFEDBF88E" box="[288,347,1883,1905]" pageId="52" pageNumber="53" rank="species">
<emphasis id="B917EADCFFC9FFE2FEBCF89BFEDBF88E" bold="true" box="[288,347,1883,1905]" pageId="52" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
(holotype, HNHM 99708). 
<emphasis id="B917EADCFFC9FFE2FDE3F89BFD19F88E" bold="true" box="[639,665,1883,1905]" pageId="52" pageNumber="53">D.</emphasis>
<taxonomicName id="4C634D4DFFC9FFE2FD3CF89BFB0FF88E" authority="Pall-Gergely &amp; Hunyadi" authorityName="Asami" authorityYear="2017" box="[672,1167,1883,1905]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="micropolitus" status="sp. nov.">
<emphasis id="B917EADCFFC9FFE2FD3CF89BFC08F88E" box="[672,904,1883,1905]" italics="true" pageId="52" pageNumber="53">Dicharax micropolitus</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
, 
<taxonomicNameLabel id="A22457A7FFC9FFE2FB07F89BFB57F88E" box="[1179,1239,1883,1905]" pageId="52" pageNumber="53" rank="species">
<emphasis id="B917EADCFFC9FFE2FB07F89BFB57F88E" bold="true" box="[1179,1239,1883,1905]" pageId="52" pageNumber="53">n. sp.</emphasis>
</taxonomicNameLabel>
(holotype, HNHM 99709). Scale represents 5 mm. All photos: B. Páll-Gergely.
</paragraph>
</caption>
<paragraph id="8BDC36CEFFC8FFE3FF5BFF57FF65FF08" blockId="53.[151,1437,151,680]" pageId="53" pageNumber="54">
<emphasis id="B917EADCFFC8FFE3FF5BFF57FED6FF4F" bold="true" box="[199,342,151,176]" pageId="53" pageNumber="54">Operculum.</emphasis>
THE OpErCULUm Is prEsENT IN ONE wEaTHErEd spECImEN (COLL. OK). THE OUTEr sIdE Is CONCavE, NEarLy smOOTH, bUT ITs mULTIspIraL sTrUCTUrE Is jUsT vIsIbLE; THE INNEr sUrfaCE Is wITH a sIGNIfICaNTLy ELEvaTEd, pOINTEd NIppLE.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFC8FFE3FF5BFEC4FB18FEE3" box="[199,1176,260,285]" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BDC36CEFFC8FFE3FF5BFEC4FB18FEE3" blockId="53.[151,1437,151,680]" box="[199,1176,260,285]" pageId="53" pageNumber="54">
<emphasis id="B917EADCFFC8FFE3FF5BFEC4FE5DFEE2" bold="true" box="[199,477,260,285]" pageId="53" pageNumber="54">Variation in specimens.</emphasis>
NO TraITs fOUNd TO bE varIabLE IN 
<collectionCode id="ED72AE0BFFC8FFE3FCCDFEC5FCF3FEE3" box="[849,883,261,284]" pageId="53" pageNumber="54">THE</collectionCode>
sINGLE kNOwN pOpULaTION.
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFC8FFE3FF5BFEE7FA09FE0A" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="8BDC36CEFFC8FFE3FF5BFEE7FC1CFE77" blockId="53.[151,1437,151,680]" pageId="53" pageNumber="54">
<emphasis id="B917EADCFFC8FFE3FF5BFEE7FE4FFEBF" bold="true" box="[199,463,295,320]" pageId="53" pageNumber="54">Differential diagnosis.</emphasis>
<taxonomicName id="4C634D4DFFC8FFE3FE44FEE8FD0CFEBF" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[472,652,296,320]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FE44FEE8FD0CFEBF" box="[472,652,296,320]" italics="true" pageId="53" pageNumber="54">Dicharax draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FD08FEE7FD55FEBF" box="[660,725,295,320]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FD08FEE7FD55FEBF" bold="true" box="[660,725,295,320]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
CaN bE dIsTINGUIsHEd frOm aLL CONGENErs INHabITING 
<collectingCountry id="F374765EFFC8FFE3FAB4FEE8FAEBFEBF" box="[1320,1387,296,320]" name="China" pageId="53" pageNumber="54">CHINa</collectingCountry>
aNd 
<collectingCountry id="F374765EFFC8FFE3FF0BFE8DFF76FE9B" box="[151,246,333,356]" name="Vietnam" pageId="53" pageNumber="54">VIETNam</collectingCountry>
ON THE basIs Of THE vEry sTrONGLy CrENULaTEd pErIsTOmE, THE sHOrT R3 HavING a prOmINENT swELLING IN ITs mIddLE (CLOsE TO THE apErTUrE). SEE aLsO UNdEr 
<taxonomicName id="4C634D4DFFC8FFE3FD05FEB1FCD7FE78" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[665,855,368,392]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="microcostatus" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FD05FEB1FCD7FE78" box="[665,855,368,392]" italics="true" pageId="53" pageNumber="54">D. microcostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FCC2FEAFFC1CFE77" box="[862,924,367,392]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FCC2FEAFFC1CFE77" bold="true" box="[862,924,367,392]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDC36CEFFC8FFE3FF5BFE55FA09FE0A" blockId="53.[151,1437,151,680]" pageId="53" pageNumber="54">
THE INdIaN 
<taxonomicName id="4C634D4DFFC8FFE3FED1FE54FDD5FE53" box="[333,597,404,428]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="plectocheilus">
<emphasis id="B917EADCFFC8FFE3FED1FE54FDD5FE53" box="[333,597,404,428]" italics="true" pageId="53" pageNumber="54">Dicharax plectocheilus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF24B3FFFC8FFE3FDFBFE54FC85FE52" author="BENsON" box="[615,773,404,429]" pageId="53" pageNumber="54" refString="Benson, W. H. (1859) A Sectional Distribution of the genus Alycaeus, Gray, with Characters of six new species and of other Cyclostomidae collected at Darjiling by W. T. Blanford, Esq., Geol. Survey. The Annals and Magazine of Natural History, 3 (3), 176 - 184." type="journal article" year="1859">BENsON, 1859</bibRefCitation>
) Is sImILar TO 
<taxonomicName id="4C634D4DFFC8FFE3FC36FE55FB91FE53" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[938,1041,404,428]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FC36FE55FB91FE53" box="[938,1041,404,428]" italics="true" pageId="53" pageNumber="54">D. draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FB86FE54FBDBFE52" box="[1050,1115,404,429]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FB86FE54FBDBFE52" bold="true" box="[1050,1115,404,429]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
IN sHapE, bUT ITs R2 Has NO dIsTINCT rIbs, ONLy wITH NarrOw LIGHT aNd THICk dark TraNsvErsE sTrIpEs aLTErNaTE, fOrmING a NEarLy smOOTH arEa. MOrEOvEr, THE pErIsTOmE Is CrENULaTEd IN 
<taxonomicName id="4C634D4DFFC8FFE3FDFFFE1CFCE7FE0B" box="[611,871,476,500]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="plectocheilus">
<emphasis id="B917EADCFFC8FFE3FDFFFE1CFCE7FE0B" box="[611,871,476,500]" italics="true" pageId="53" pageNumber="54">Dicharax plectocheilus</emphasis>
</taxonomicName>
, bUT sTraIGHT frOm LaTEraL vIEw IN 
<taxonomicName id="4C634D4DFFC8FFE3FB43FE1DFAC3FE0B" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[1247,1347,476,500]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FB43FE1DFAC3FE0B" box="[1247,1347,476,500]" italics="true" pageId="53" pageNumber="54">D. draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FAD6FE1CFA09FE0A" box="[1354,1417,476,501]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FAD6FE1CFA09FE0A" bold="true" box="[1354,1417,476,501]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFC8FFE3FF5BFE3FFBFCFDE7" box="[199,1148,511,536]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8BDC36CEFFC8FFE3FF5BFE3FFBFCFDE7" blockId="53.[151,1437,151,680]" box="[199,1148,511,536]" pageId="53" pageNumber="54">
<emphasis id="B917EADCFFC8FFE3FF5BFE3FFEE0FDE7" bold="true" box="[199,352,511,536]" pageId="53" pageNumber="54">Distribution.</emphasis>
<taxonomicName id="4C634D4DFFC8FFE3FEFBFDC0FD9AFDE7" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[359,538,512,536]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FEFBFDC0FD9AFDE7" box="[359,538,512,536]" italics="true" pageId="53" pageNumber="54">Dicharax draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FDBCFE3FFDDFFDE7" box="[544,607,511,536]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FDBCFE3FFDDFFDE7" bold="true" box="[544,607,511,536]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
Is kNOwN frOm THE 
<typeStatus id="54D8886CFFC8FFE3FCA6FDC0FCEAFDE7" box="[826,874,512,536]" pageId="53" pageNumber="54">TypE</typeStatus>
LOCaLITy ONLy (
<figureCitation id="13582A4BFFC8FFE3FB97FDC0FBEEFDE7" box="[1035,1134,512,536]" captionStart="FIGURE 7" captionStartId="15.[151,250,1122,1144]" captionTargetBox="[151,1436,193,1101]" captionTargetId="figure@15.[151,1436,193,1101]" captionTargetPageId="15" captionText="FIGURE 7. Distribution of narrow range or single-site endemic Dicharax species. Filled circle. Dicharax tangmaiensis (Chen &amp; Zhang, 2001). Filled triangle, top up. Dicharax (?) alticola Páll-Gergely &amp; Hunyadi n. sp. Filled diamond. Dicharax (?) immaculatus Páll-Gergely n. sp. Empty diamond. Dicharax microcostatus Páll-Gergely n. sp., Dicharax micropolitus Páll- Gergely &amp; Hunyadi, n. sp. and Dicharax bison Páll-Gergely &amp; Hunyadi n. sp. Filled triangle, top down. Dicharax robustus Páll-Gergely &amp; Hunyadi, n. sp. Empty triangle, top down. Dicharax draco Páll-Gergely &amp; Hunyadi n. sp. Empty square. Dicharax imitator Páll-Gergely &amp; Hunyadi n. sp. Empty triangle, top up. Dicharax (?) moellendorffi (Kobelt &amp; Möllendorff, 1886). Filled square. Dicharax (?) ellipticus n. sp. Cross. Dicharax (?) abdoui Páll-Gergely n. sp. Empty circle shows the position of Tchen K’eou (Chengkou), which is the type locality of Alycaeus pentagonus Heude, 1886 (synonym of Alycaeus anthostoma Möllendorff, 1885), Dicharax diminutus (Heude, 1885), Dicharax (?) fargesianus (Heude, 1885), Dicharax planorbulus (Heude, 1885), Metalycaeus muciferus (Heude, 1885), Alycaeus dolomiticus Heude, 1890 (synonym of Alycaeus rathouisianus Heude, 1882), Dicharax (?) dolichodeiros (Heude, 1890), Metalycaeus (?) elevatus (Heude, 1886), Metalycaeus (?) expansus (Heude, 1890)." httpUri="https://zenodo.org/record/1009739/files/figure.png" pageId="53" pageNumber="54">FIGUrE 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3796545FFC8FFE3FF5BFDE4FC58FD57" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BDC36CEFFC8FFE3FF5BFDE4FC58FD57" blockId="53.[151,1437,151,680]" pageId="53" pageNumber="54">
<emphasis id="B917EADCFFC8FFE3FF5BFDE4FEBBFDC2" bold="true" box="[199,315,548,573]" pageId="53" pageNumber="54">Remarks.</emphasis>
WE HavE CONsIdErEd INTErprETING 
<taxonomicName id="4C634D4DFFC8FFE3FD2CFDE4FCE2FDC3" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" box="[688,866,548,572]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FD2CFDE4FCE2FDC3" box="[688,866,548,572]" italics="true" pageId="53" pageNumber="54">Dicharax draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FCF4FDE4FC26FDC2" box="[872,934,548,573]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FCF4FDE4FC26FDC2" bold="true" box="[872,934,548,573]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
as aN UNUsUaL LOCaL fOrm Of THE vEry varIabLE 
<taxonomicName id="4C634D4DFFC8FFE3FF0BFD89FEADFD9F" box="[151,301,584,608]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="fimbriatus">
<emphasis id="B917EADCFFC8FFE3FF0BFD89FEADFD9F" box="[151,301,584,608]" italics="true" pageId="53" pageNumber="54">D. fimbriatus</emphasis>
</taxonomicName>
. HOwEvEr, THE GENEraL sHELL sHapE, THE LENGTH Of THE sHELL rEGIONs aNd THE pOsITION Of THE swELLING Of R3 faLL OUTsIdE Of THE mOrpHOLOGICaL raNGE Of 
<taxonomicName id="4C634D4DFFC8FFE3FD2AFDADFCCFFD7B" box="[694,847,620,644]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="fimbriatus">
<emphasis id="B917EADCFFC8FFE3FD2AFDADFCCFFD7B" box="[694,847,620,644]" italics="true" pageId="53" pageNumber="54">D. fimbriatus</emphasis>
</taxonomicName>
. MOrEOvEr, THE sINGLE kNOwN LOCaLITy Of 
<taxonomicName id="4C634D4DFFC8FFE3FAAEFDACFF59FD57" authority="Asami, 2017" authorityName="Asami" authorityYear="2017" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="draco" status="sp. nov.">
<emphasis id="B917EADCFFC8FFE3FAAEFDACFF59FD57" italics="true" pageId="53" pageNumber="54">Dicharax draco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457A7FFC8FFE3FF43FD4FFE9EFD57" box="[223,286,655,680]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B917EADCFFC8FFE3FF43FD4FFE9EFD57" bold="true" box="[223,286,655,680]" pageId="53" pageNumber="54">n. sp.</emphasis>
</taxonomicNameLabel>
Is sUrrOUNdEd by &quot;NOrmaL LOOkING&quot; 
<taxonomicName id="4C634D4DFFC8FFE3FD2CFD51FCC6FD57" box="[688,838,656,680]" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="fimbriatus">
<emphasis id="B917EADCFFC8FFE3FD2CFD51FCC6FD57" box="[688,838,656,680]" italics="true" pageId="53" pageNumber="54">D. fimbriatus</emphasis>
</taxonomicName>
pOpULaTIONs.
</paragraph>
</subSubSection>
</treatment>
</document>