617 lines
66 KiB
XML
617 lines
66 KiB
XML
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<mods:title id="7211C20F8B8B27B26CB02F42CCC8C300">Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)</mods:title>
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<mods:namePart id="1058D51B665B8586F15A7F4AC93D77E9">Medrano, Miguel</mods:namePart>
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<mods:namePart id="3A4A81AC07E43EC9BB512F0111D561FD">Kury, Adriano Brilhante</mods:namePart>
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<mods:namePart id="3B1C359AB442118ACD9B7FDCA19CABE9">Mendes, Amanda Cruz</mods:namePart>
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<mods:number id="652F6E05298250D722A80E88123C5199">2021-08-30</mods:number>
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<taxonomicName id="4C634D19FF98FFF7FCC5FADEFB13240F" ID-CoL="7YHWG" authority="Simon, 1879: 193" authorityName="Simon" authorityPageNumber="193" authorityYear="1879" box="[827,1204,1384,1406]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="45" pageNumber="630" phylum="Arthropoda" rank="species" species="insolens">
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<emphasis id="B917EA88FF98FFF7FCC5FADEFC4F240C" box="[827,1000,1384,1406]" italics="true" pageId="45" pageNumber="630">Protus insolens</emphasis>
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<bibRefCitation id="EFF24B6BFF98FFF7FC11FADFFB13240F" author="Simon E" box="[1007,1204,1384,1406]" pageId="45" pageNumber="630" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879: 193</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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:
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. [Amazonas]:
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,
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[–3.40357°, –64.76333°],
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(
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, examined)
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.
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: Same data as holotype 1 ind. (
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, not examined)
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.
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.
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:
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,
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[–4.87528°, –65.17417°],
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(
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)
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; [–4.86056°, –65.33444°],
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<collectorName id="2696534CFF98FFF7FC2FF921FB8A27DD" box="[977,1069,1686,1708]" pageId="45" pageNumber="630">SC Dias</collectorName>
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leg.,
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,
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(
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)
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</materialsCitation>
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; [–-4.85306°, –65.13472°],
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<collectingDate id="EF99E9B2FF98FFF7FB9FF903FB7C27BB" box="[1121,1243,1716,1738]" pageId="45" pageNumber="630" value="2006-09-12">12.ix.2006</collectingDate>
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,
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(
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<specimenCode id="DBC59EE1FF98FFF7FAEBF902FA3127BB" box="[1301,1430,1716,1738]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="45" pageNumber="630">MPEG 856</specimenCode>
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)
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</materialsCitation>
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; [–4.86833°, –65.26444°],
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<date id="FFDD105AFF98FFF7FBA7F964FB7D2798" box="[1113,1242,1747,1769]" pageId="45" pageNumber="630" value="2006-07-26">
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<collectingDate id="EF99E9B2FF98FFF7FBA7F964FB7D2798" box="[1113,1242,1747,1769]" pageId="45" pageNumber="630" value="2006-07-26">26.vii.2006</collectingDate>
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</date>
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,
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<specimenCount id="9D65FD13FF98FFF7FB1BF964FAAF2799" box="[1253,1288,1747,1768]" pageId="45" pageNumber="630" type="female">1 ♀</specimenCount>
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(
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<specimenCode id="DBC59EE1FF98FFF7FAEBF964FA312798" box="[1301,1430,1747,1769]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="45" pageNumber="630">MPEG 857</specimenCode>
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)
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</materialsCitation>
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; [–4.86861°, –65.25639°],
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<materialsCitation id="3B0B3CC7FF98FFF7FB93F945FB902657" ID-GBIF-Occurrence="3879212434" collectingDate="2003-07-19" collectionCode="MPEG" collectorName="SO Dias" country="Brazil" pageId="45" pageNumber="630" specimenCode="MPEG 859" specimenCount="1">
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<collectorName id="2696534CFF98FFF7FB93F945FB762679" box="[1133,1233,1778,1800]" pageId="45" pageNumber="630">SO Dias</collectorName>
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leg.,
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<date id="FFDD105AFF98FFF7FAEBF945FA3A2679" box="[1301,1437,1778,1800]" pageId="45" pageNumber="630" value="2003-07-19">
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<collectingDate id="EF99E9B2FF98FFF7FAEBF945FA3A2679" box="[1301,1437,1778,1800]" pageId="45" pageNumber="630" value="2003-07-19">19.vii.2003</collectingDate>
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</date>
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,
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<specimenCount id="9D65FD13FF98FFF7FCC5F8A6FC362657" box="[827,913,1808,1830]" pageId="45" pageNumber="630" type="generic">1 imm.</specimenCount>
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(
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<specimenCode id="DBC59EE1FF98FFF7FC5CF8A6FB892657" box="[930,1070,1808,1830]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="45" pageNumber="630">MPEG 859</specimenCode>
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)
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</materialsCitation>
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; [–4.80639°, –65.03472°],
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<materialsCitation id="3B0B3CC7FF98FFF7FA7EF8A6FAA92634" ID-GBIF-Occurrence="3879212358" collectingDate="2006-07-09" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" pageId="45" pageNumber="630" specimenCode="MPEG 860" specimenCount="1" specimenCount-male="1">
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<collectorName id="2696534CFF98FFF7FA7EF8A6FCD62634" pageId="45" pageNumber="630">SC Dias</collectorName>
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leg.,
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<date id="FFDD105AFF98FFF7FC4AF898FB9D2634" box="[948,1082,1839,1861]" pageId="45" pageNumber="630" value="2006-07-09">
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<collectingDate id="EF99E9B2FF98FFF7FC4AF898FB9D2634" box="[948,1082,1839,1861]" pageId="45" pageNumber="630" value="2006-07-09">09.vii.2006</collectingDate>
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</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF98FFF7FBB6F898FBC82635" box="[1096,1135,1839,1860]" pageId="45" pageNumber="630" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF98FFF7FB81F898FAA02634" box="[1151,1287,1839,1861]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="45" pageNumber="630">MPEG 860</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.84222°, –65.06889°],
|
||
<materialsCitation id="3B0B3CC7FF98FFF4FC2CF8F9FF6327AD" ID-GBIF-Occurrence="3879212406" collectingDate="2006-09-05" collectionCode="MPEG" collectorName="DF Candiani" country="Brazil" lastPageId="46" lastPageNumber="631" pageId="45" pageNumber="630" specimenCode="MPEG 861" specimenCount="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF98FFF7FC2CF8F9FBCE2615" box="[978,1129,1870,1892]" pageId="45" pageNumber="630">DF Candiani</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF98FFF7FB59F8F9FAB42615" box="[1191,1299,1870,1892]" pageId="45" pageNumber="630" value="2006-09-05">
|
||
<collectingDate id="EF99E9B2FF98FFF7FB59F8F9FAB42615" box="[1191,1299,1870,1892]" pageId="45" pageNumber="630" value="2006-09-05">5.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF98FFF7FAE1F8F9FAE32612" box="[1311,1348,1870,1891]" pageId="45" pageNumber="630" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF98FFF4FAAAF8F9FF1A27AD" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" lastPageId="46" lastPageNumber="631" name="Museu Paraense Emilio Goeldi" pageId="45" pageNumber="630">MPEG 861</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.89167°, –65.33528°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FE0DF970FE6A278A" ID-GBIF-Occurrence="3879212392" collectingDate="2006-11-01" collectionCode="MPEG" collectorName="NF Lo Man Hung" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 862" specimenCount="2" specimenCount-male="2">
|
||
<collectorName id="2696534CFF9BFFF4FE0DF970FD6527AD" box="[499,706,1735,1756]" pageId="46" pageNumber="631">NF Lo Man Hung</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FF6FF952FF59278A" box="[145,254,1765,1787]" pageId="46" pageNumber="631" value="2006-11-01">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FF6FF952FF59278A" box="[145,254,1765,1787]" pageId="46" pageNumber="631" value="2006-11-01">1.xi.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FEF5F952FE96278B" box="[267,305,1765,1786]" pageId="46" pageNumber="631" type="male">2 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FEBFF952FE61278A" box="[321,454,1765,1787]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 862</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.85722°, –65.10167°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FF6FF8B3FE6A266B" ID-GBIF-Occurrence="3879212309" box="[145,461,1796,1818]" collectingDate="2006-09-09" collectionCode="MPEG" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 863" specimenCount="1" specimenCount-male="1">
|
||
<date id="FFDD105AFF9BFFF4FF6FF8B3FF59266B" box="[145,254,1796,1818]" pageId="46" pageNumber="631" value="2006-09-09">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FF6FF8B3FF59266B" box="[145,254,1796,1818]" pageId="46" pageNumber="631" value="2006-09-09">9.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FEF5F8B3FE962669" box="[267,305,1796,1817]" pageId="46" pageNumber="631" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FEBFF8B3FE61266B" box="[321,454,1796,1818]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 863</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.89167°, –65.33528°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FF6FF895FE6A2649" ID-GBIF-Occurrence="3879212384" box="[145,461,1826,1848]" collectingDate="2006-11-01" collectionCode="MPEG" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 864" specimenCount="1" specimenCount-male="1">
|
||
<date id="FFDD105AFF9BFFF4FF6FF895FF592649" box="[145,254,1826,1848]" pageId="46" pageNumber="631" value="2006-11-01">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FF6FF895FF592649" box="[145,254,1826,1848]" pageId="46" pageNumber="631" value="2006-11-01">1.xi.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FEF5F895FE962646" box="[267,305,1826,1847]" pageId="46" pageNumber="631" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FEBFF895FE612649" box="[321,454,1826,1848]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 864</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.83361°, –65.06472°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FF6FF8F6FDCC2626" ID-GBIF-Occurrence="3879212457" box="[145,619,1857,1879]" collectingDate="2006-09-03" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 865" specimenCount="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF9BFFF4FF6FF8F6FF492626" box="[145,238,1857,1879]" pageId="46" pageNumber="631">SC Dias</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FED4F8F6FE032626" box="[298,420,1857,1879]" pageId="46" pageNumber="631" value="2006-09-03">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FED4F8F6FE032626" box="[298,420,1857,1879]" pageId="46" pageNumber="631" value="2006-09-03">03.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FE51F8F6FE732627" box="[431,468,1857,1878]" pageId="46" pageNumber="631" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FE1CF8F6FDC42626" box="[482,611,1857,1879]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 865</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.88528°, –65.26472°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FED8F8D7FCFA27AD" ID-GBIF-Occurrence="3879212326" box="[294,861,1734,1910]" collectingDate="2003-07-21" collectionCode="MPEG" collectorName="AB Bonaldo" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 866" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF9BFFF4FED8F8D7FE172607" box="[294,432,1888,1910]" pageId="46" pageNumber="631">AB Bonaldo</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FE12F8D7FDC92607" box="[492,622,1888,1910]" pageId="46" pageNumber="631" value="2003-07-21">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FE12F8D7FDC92607" box="[492,622,1888,1910]" pageId="46" pageNumber="631" value="2003-07-21">21.vii.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FD87F8D7FD3B2605" box="[633,668,1888,1909]" pageId="46" pageNumber="631" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FD54F8D7FCF227AD" box="[682,853,1734,1909]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 866</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.84222°, –65.06889°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FB7EF971FC45278A" ID-GBIF-Occurrence="3879212401" collectingDate="2006-09-05" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 867" specimenCount="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF9BFFF4FB7EF971FB7C27AD" box="[1152,1243,1734,1756]" pageId="46" pageNumber="631">SC Dias</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FAEDF971FA2B27AD" box="[1299,1420,1734,1756]" pageId="46" pageNumber="631" value="2006-09-05">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FAEDF971FA2B27AD" box="[1299,1420,1734,1756]" pageId="46" pageNumber="631" value="2006-09-05">05.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FCD7F952FCEB278B" box="[809,844,1765,1786]" pageId="46" pageNumber="631" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FCA7F952FC7D278A" box="[857,986,1765,1787]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 867</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.86861°, –65.26472°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FAF6F952FB4E266B" ID-GBIF-Occurrence="3879212453" collectingDate="2003-07-15" collectionCode="MPEG" collectorName="AB Bonaldo" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 870" specimenCount="2" specimenCount-male="2">
|
||
<collectorName id="2696534CFF9BFFF4FAF6F952FA36278A" box="[1288,1425,1765,1787]" pageId="46" pageNumber="631">AB Bonaldo</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FC94F8B4FC5B2668" box="[874,1020,1795,1817]" pageId="46" pageNumber="631" value="2003-07-15">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FC94F8B4FC5B2668" box="[874,1020,1795,1817]" pageId="46" pageNumber="631" value="2003-07-15">15.vii.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FBF0F8B3FB9E2669" box="[1038,1081,1796,1817]" pageId="46" pageNumber="631" type="male">2 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FBB3F8B3FB47266B" box="[1101,1248,1796,1818]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 870</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FB04F8B3FB9B2649" ID-GBIF-Occurrence="3879212458" collectingDate="2003-07-19" collectionCode="MPEG" collectorName="AB Bonaldo" country="Brazil" isTailEnumeration="true" pageId="46" pageNumber="631" specimenCode="MPEG 871" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<date id="FFDD105AFF9BFFF4FB04F8B3FA2D266B" box="[1274,1418,1796,1818]" pageId="46" pageNumber="631" value="2003-07-19">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FB04F8B3FA2D266B" box="[1274,1418,1796,1818]" pageId="46" pageNumber="631" value="2003-07-19">19.vii.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FCD7F895FCF32646" box="[809,852,1826,1847]" pageId="46" pageNumber="631" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="9D65FD13FF9BFFF4FC9FF895FC2B2646" box="[865,908,1826,1847]" pageId="46" pageNumber="631" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FC5EF895FB932649" box="[928,1076,1826,1848]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 871</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.84222°, –65.06889°], CAC
|
||
<materialsCitation id="3B0B3CC7FF9BFFF4FC95F8F6FA2C2626" ID-GBIF-Occurrence="3879212304" box="[875,1419,1857,1879]" collectingDate="2006-09-05" collectionCode="MPEG" collectorName="Santos Jr" country="Brazil" pageId="46" pageNumber="631" specimenCode="MPEG 872" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF9BFFF4FC95F8F6FC432627" box="[875,996,1857,1878]" pageId="46" pageNumber="631">Santos Jr</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FBD2F8F6FB092626" box="[1068,1198,1857,1879]" pageId="46" pageNumber="631" value="2006-09-05">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FBD2F8F6FB092626" box="[1068,1198,1857,1879]" pageId="46" pageNumber="631" value="2006-09-05">05.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FB40F8F6FB422627" box="[1214,1253,1857,1878]" pageId="46" pageNumber="631" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF9BFFF4FB09F8F6FA242626" box="[1271,1411,1857,1879]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="46" pageNumber="631">MPEG 872</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.90444°, –65.32694°],
|
||
<materialsCitation id="3B0B3CC7FF9BFFEAFBB4F8D7FE8E21AA" ID-GBIF-Occurrence="3879212362" collectingDate="2006-09-21" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" lastPageId="48" lastPageNumber="633" pageId="46" pageNumber="631" specimenCode="MPEG 873" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF9BFFF4FBB4F8D7FB0E2607" box="[1098,1193,1888,1910]" pageId="46" pageNumber="631">SC Dias</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF9BFFF4FB19F8D7FAC62607" box="[1255,1377,1888,1910]" pageId="46" pageNumber="631" value="2006-09-21">
|
||
<collectingDate id="EF99E9B2FF9BFFF4FB19F8D7FAC62607" box="[1255,1377,1888,1910]" pageId="46" pageNumber="631" value="2006-09-21">21.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF9BFFF4FA93F8D7FA362605" box="[1389,1425,1888,1909]" pageId="46" pageNumber="631" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFF66FF72FE8621AA" box="[152,289,197,219]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 873</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.86861°, –65.26472°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFD96FF72FDC6218B" ID-GBIF-Occurrence="3879212424" collectingDate="2003-07-18" collectionCode="MPEG" collectorName="AB Bonaldo" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 874" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF85FFEAFD96FF72FD5F21AA" box="[616,760,197,219]" pageId="48" pageNumber="633">AB Bonaldo</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFF32FF53FEF2218B" box="[204,341,228,250]" pageId="48" pageNumber="633" value="2003-07-18">
|
||
<collectingDate id="EF99E9B2FF85FFEAFF32FF53FEF2218B" box="[204,341,228,250]" pageId="48" pageNumber="633" value="2003-07-18">18.vii.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFE9AFF53FE2B2188" box="[356,396,228,249]" pageId="48" pageNumber="633" type="male">1 ♂</specimenCount>
|
||
<specimenCount id="9D65FD13FF85FFEAFE68FF53FE1A2188" box="[406,445,228,249]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFE31FF53FDFF218B" box="[463,600,228,250]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 874</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.87361°, –65.15139°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFED9FEB4FD532069" ID-GBIF-Occurrence="3879212337" box="[295,756,258,280]" collectingDate="2006-10-14" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 875" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF85FFEAFED9FEB4FE232069" box="[295,388,258,280]" pageId="48" pageNumber="633">SC Dias</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFE41FEB5FD972069" box="[447,560,258,280]" pageId="48" pageNumber="633" value="2006-10-14">
|
||
<collectingDate id="EF99E9B2FF85FFEAFE41FEB5FD972069" box="[447,560,258,280]" pageId="48" pageNumber="633" value="2006-10-14">14.x.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFDC5FEB4FDF92066" box="[571,606,259,280]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFD95FEB4FD4B2069" box="[619,748,258,280]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 875</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.88722°, –65.22694°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFE4FFE96FD532046" ID-GBIF-Occurrence="3879212416" box="[433,756,289,311]" collectingDate="2006-09-22" collectionCode="MPEG" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 876" specimenCount="2" specimenCount-male="2">
|
||
<date id="FFDD105AFF85FFEAFE4FFE96FD8C2046" box="[433,555,289,311]" pageId="48" pageNumber="633" value="2006-09-22">
|
||
<collectingDate id="EF99E9B2FF85FFEAFE4FFE96FD8C2046" box="[433,555,289,311]" pageId="48" pageNumber="633" value="2006-09-22">22.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFDC9FE96FDFC2047" box="[567,603,289,310]" pageId="48" pageNumber="633" type="male">2 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFD94FE96FD4B2046" box="[618,748,289,311]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 876</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.87944°, –65.16389°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFE4CFEF7FD532027" ID-GBIF-Occurrence="3879212314" box="[434,756,320,342]" collectingDate="2006-09-12" collectionCode="MPEG" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 877" specimenCount="1" specimenCount-female="1">
|
||
<date id="FFDD105AFF85FFEAFE4CFEF7FD8B2027" box="[434,556,320,342]" pageId="48" pageNumber="633" value="2006-09-12">
|
||
<collectingDate id="EF99E9B2FF85FFEAFE4CFEF7FD8B2027" box="[434,556,320,342]" pageId="48" pageNumber="633" value="2006-09-12">12.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFDC9FEF7FDFC2024" box="[567,603,320,341]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFD94FEF7FD4B2027" box="[618,748,320,342]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 877</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFF6FFEE9FDCB2004" ID-GBIF-Occurrence="3879212439" box="[145,620,350,373]" collectingDate="2006-09-12" collectionCode="MPEG" country="Brazil" isTailEnumeration="true" pageId="48" pageNumber="633" specimenCode="MPEG 878" specimenCount="1" specimenCount-male="1">
|
||
[–4.86972°, –65.13472°],
|
||
<specimenCount id="9D65FD13FF85FFEAFE4EFEE8FE732002" box="[432,468,351,372]" pageId="48" pageNumber="633" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFE1CFEE8FDC32004" box="[482,612,351,373]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 878</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.81139°, –65.03361°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFECBFEC9FF6020C3" ID-GBIF-Occurrence="3879212414" collectingDate="2006-07" collectionCode="MPEG" collectorName="LT Miglio" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 879" specimenCount="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF85FFEAFECBFEC9FE0920E2" box="[309,430,381,403]" pageId="48" pageNumber="633">LT Miglio</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFE09FECAFDFB20E2" box="[503,604,381,403]" pageId="48" pageNumber="633" value="2006-07">
|
||
<collectingDate id="EF99E9B2FF85FFEAFE09FECAFDFB20E2" box="[503,604,381,403]" pageId="48" pageNumber="633" value="2006-07">vii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFD95FECAFD3320E3" box="[619,660,381,402]" pageId="48" pageNumber="633" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFD58FECAFF1820C3" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 879</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.90444°, –65.32694°], CAC
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFDB6FE2BFE7420A0" ID-GBIF-Occurrence="3879212311" collectingDate="2006-09-21" collectionCode="MPEG" collectorName="Santos Jr" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 880" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF85FFEAFDB6FE2BFD1920C0" box="[584,702,412,433]" pageId="48" pageNumber="633">Santos Jr</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFF6FFE0DFEAC20A1" box="[145,267,442,464]" pageId="48" pageNumber="633" value="2006-09-21">
|
||
<collectingDate id="EF99E9B2FF85FFEAFF6FFE0DFEAC20A1" box="[145,267,442,464]" pageId="48" pageNumber="633" value="2006-09-21">21.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFEE8FE0CFE9D20BE" box="[278,314,443,464]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFEB7FE0CFE6C20A0" box="[329,459,443,465]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 880</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.86833°, –65.26444°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFF6FFE6EFDC1209E" ID-GBIF-Occurrence="3879212402" box="[145,614,473,495]" collectingDate="2006-07" collectionCode="MPEG" collectorName="SC Dias" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 881" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF85FFEAFF6FFE6EFF55209E" box="[145,242,473,495]" pageId="48" pageNumber="633">SC Dias</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFECDFE6EFE32209E" box="[307,405,473,495]" pageId="48" pageNumber="633" value="2006-07">
|
||
<collectingDate id="EF99E9B2FF85FFEAFECDFE6EFE32209E" box="[307,405,473,495]" pageId="48" pageNumber="633" value="2006-07">vii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFE5CFE6EFE60209F" box="[418,455,473,494]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFE29FE6EFDFA209E" box="[471,605,473,495]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 881</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.88722°, –65.22694°], CAC
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFE84FE4FFE86235C" ID-GBIF-Occurrence="3879212463" collectingDate="2006-09-22" collectionCode="MPEG" collectorName="Santos Jr" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 882" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF85FFEAFE84FE4FFE53237C" box="[378,500,504,525]" pageId="48" pageNumber="633">Santos Jr</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFDC3FE4FFD67237F" box="[573,704,504,526]" pageId="48" pageNumber="633" value="2006-09-22">
|
||
<collectingDate id="EF99E9B2FF85FFEAFDC3FE4FFD67237F" box="[573,704,504,526]" pageId="48" pageNumber="633" value="2006-09-22">22.ix.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFD2FFE4FFD5F237C" box="[721,760,504,525]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFF66FDA0FEBE235C" box="[152,281,535,557]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 882</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.87361°, –65.15139°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFDB4FDA0FE6E233A" ID-GBIF-Occurrence="3879212440" collectingDate="2006-10-01" collectionCode="MPEG" collectorName="NC Bastos" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 883" specimenCount="1" specimenCount-male="1">
|
||
<collectorName id="2696534CFF85FFEAFDB4FDA0FD63235D" box="[586,708,535,556]" pageId="48" pageNumber="633">NC Bastos</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFF6FFD82FF50233A" box="[145,247,565,587]" pageId="48" pageNumber="633" value="2006-10-01">
|
||
<collectingDate id="EF99E9B2FF85FFEAFF6FFD82FF50233A" box="[145,247,565,587]" pageId="48" pageNumber="633" value="2006-10-01">1.x.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFEFAFD82FE8C233B" box="[260,299,565,586]" pageId="48" pageNumber="633" type="male">1 ♂</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFEC5FD82FE66233A" box="[315,449,565,587]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 883</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; [–4.90417°, –65.20583°],
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFF6FFDE3FDCE231B" ID-GBIF-Occurrence="3879212430" box="[145,617,596,618]" collectingDate="2006-07" collectionCode="MPEG" collectorName="LT Miglio" country="Brazil" pageId="48" pageNumber="633" specimenCode="MPEG 884" specimenCount="1" specimenCount-female="1">
|
||
<collectorName id="2696534CFF85FFEAFF6FFDE3FEA5231B" box="[145,258,596,618]" pageId="48" pageNumber="633">LT Miglio</collectorName>
|
||
leg.,
|
||
<date id="FFDD105AFF85FFEAFEC1FDE3FE39231B" box="[319,414,596,618]" pageId="48" pageNumber="633" value="2006-07">
|
||
<collectingDate id="EF99E9B2FF85FFEAFEC1FDE3FE39231B" box="[319,414,596,618]" pageId="48" pageNumber="633" value="2006-07">vii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD13FF85FFEAFE54FDE3FE692318" box="[426,462,596,617]" pageId="48" pageNumber="633" type="female">1 ♀</specimenCount>
|
||
(
|
||
<specimenCode id="DBC59EE1FF85FFEAFE23FDE3FDC6231B" box="[477,609,596,618]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="48" pageNumber="633">MPEG 884</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF1C6612FF9BFFF4FF6FF9F8FEEC27D1" ID-DOI="http://doi.org/10.5281/zenodo.6959602" ID-Zenodo-Dep="6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="46" pageNumber="631" startId="46.[145,223,1615,1637]" targetBox="[145,1425,195,1575]" targetPageId="46">
|
||
<paragraph id="8BDC369AFF9BFFF4FF6FF9F8FEEC27D1" blockId="46.[145,1425,1615,1696]" pageId="46" pageNumber="631">
|
||
<emphasis id="B917EA88FF9BFFF4FF6FF9F8FEA12715" bold="true" box="[145,262,1615,1637]" pageId="46" pageNumber="631">Figure 35.</emphasis>
|
||
<taxonomicName id="4C634D19FF9BFFF4FEEFF9E7FE172714" authorityName="Simon" authorityYear="1879" box="[273,432,1615,1637]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="46" pageNumber="631" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B917EA88FF9BFFF4FEEFF9E7FE172714" box="[273,432,1615,1637]" italics="true" pageId="46" pageNumber="631">Protus insolens</emphasis>
|
||
</taxonomicName>
|
||
, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1C6612FF9AFFF5FF5DF94CFE512618" ID-DOI="http://doi.org/10.5281/zenodo.6959604" ID-Zenodo-Dep="6959604" httpUri="https://zenodo.org/record/6959604/files/figure.png" pageId="47" pageNumber="632" startId="47.[163,241,1787,1809]" targetBox="[243,1362,196,1746]" targetPageId="47">
|
||
<paragraph id="8BDC369AFF9AFFF5FF5DF94CFE512618" blockId="47.[163,1444,1787,1897]" pageId="47" pageNumber="632">
|
||
<emphasis id="B917EA88FF9AFFF5FF5DF94CFEB02661" bold="true" box="[163,279,1787,1809]" pageId="47" pageNumber="632">Figure 36.</emphasis>
|
||
SEM images of some structures of
|
||
<taxonomicName id="4C634D19FF9AFFF5FD77F94BFC802660" authorityName="Simon" authorityYear="1879" box="[649,807,1787,1809]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="47" pageNumber="632" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B917EA88FF9AFFF5FD77F94BFC802660" box="[649,807,1787,1809]" italics="true" pageId="47" pageNumber="632">Protus insolens</emphasis>
|
||
</taxonomicName>
|
||
, male (MPEG 867). A–C, lenis: A, dorsal view; B, lateral view; C, ventral view. D. Left basichelicerite, dorsal view. E, left cheliceral hand, dorsal view. F, left pedipalpal tarsus, ventral view. G, left pedipalpus, mesal view. H, left claw IV, ventral view. I, left claw III, ventro-lateral view. Scale bars: 100 μm (A–C, F, H, I), 200 μm (D, E), 500 μm (G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDC369AFF85FFEAFF6FFD25FEA923B7" blockId="48.[145,761,658,710]" pageId="48" pageNumber="633">
|
||
<emphasis id="B917EA88FF85FFEAFF6FFD25FF5723D6" box="[145,240,658,679]" italics="true" pageId="48" pageNumber="633">Records:</emphasis>
|
||
<collectingRegion id="49A7F878FF85FFEAFEFFFD24FED023D9" box="[257,375,659,680]" country="Venezuela" name="Amazonas" pageId="48" pageNumber="633">Amazonas</collectingRegion>
|
||
(H.
|
||
<bibRefCitation id="EFF24B6BFF85FFEAFE5BFD24FD9323D9" author="Soares HEM" box="[421,564,659,680]" pageId="48" pageNumber="633" pagination="323 - 338" refId="ref55298" refString="Soares HEM. 1970. Novas especies de opilioes da Regiao Amazonica (Opiliones, Cosmetidae, Gonyleptidae, Phalangiidae, Stygnidae). Revista Brasileira de Biologia 30: 323 - 338." type="journal article" year="1970">Soares, 1970</bibRefCitation>
|
||
), probably Upper
|
||
<collectingRegion id="49A7F878FF85FFEAFF6FFD06FEAD23B7" box="[145,266,689,710]" country="Venezuela" name="Amazonas" pageId="48" pageNumber="633">Amazonas</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF6FFD47FD5E22CC" blockId="48.[145,762,752,958]" pageId="48" pageNumber="633">
|
||
<emphasis id="B917EA88FF85FFEAFF6FFD47FEB22274" box="[145,277,752,773]" italics="true" pageId="48" pageNumber="633">Diagnosis:</emphasis>
|
||
It differs from other
|
||
<taxonomicName id="4C634D19FF85FFEAFDB6FD47FD3D2274" authorityName="Simon" authorityYear="1879" box="[584,666,752,773]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="48" pageNumber="633" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FF85FFEAFDB6FD47FD3D2274" box="[584,666,752,773]" italics="true" pageId="48" pageNumber="633">Protus</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following characteristics: (1) coxa
|
||
<collectionCode id="ED72AE5FFF85FFEAFD20FCB8FD5D2255" box="[734,762,783,804]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
uniformly reddish without any yellow spots (
|
||
<figureCitation id="13582A1FFF85FFEAFD70FC9AFD4D2232" box="[654,746,813,835]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35F</figureCitation>
|
||
), (2) DS only with paired dots in areas I and
|
||
<collectionCode id="ED72AE5FFF85FFEAFD9AFCFBFD262210" box="[612,641,844,865]" pageId="48" pageNumber="633">III</collectionCode>
|
||
and minor dots in laterals of areas I–
|
||
<collectionCode id="ED72AE5FFF85FFEAFE52FCDCFE6F22F1" box="[428,456,875,896]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
of mesotergum, borders the same colour of the rest of DS (
|
||
<figureCitation id="13582A1FFF85FFEAFE1AFC3EFDE522EE" box="[484,578,905,927]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35A</figureCitation>
|
||
), (3) stylus with an extended wattle going to the base (
|
||
<figureCitation id="13582A1FFF85FFEAFDCCFC1FFD3322CF" box="[562,660,936,958]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Figs 35G</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF85FFEAFD5EFC1FFD4C22CF" box="[672,747,936,958]" captionStart="Figure 36" captionStartId="47.[163,241,1787,1809]" captionTargetBox="[243,1362,196,1746]" captionTargetId="figure-25@47.[243,1363,195,1747]" captionTargetPageId="47" captionText="Figure 36. SEM images of some structures of Protus insolens, male (MPEG 867). A–C, lenis: A, dorsal view; B, lateral view; C, ventral view. D. Left basichelicerite, dorsal view. E, left cheliceral hand, dorsal view. F, left pedipalpal tarsus, ventral view. G, left pedipalpus, mesal view. H, left claw IV, ventral view. I, left claw III, ventro-lateral view. Scale bars: 100 μm (A–C, F, H, I), 200 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6959604" httpUri="https://zenodo.org/record/6959604/files/figure.png" pageId="48" pageNumber="633">36A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF6FFC46FE082554" blockId="48.[145,760,1009,1062]" pageId="48" pageNumber="633">
|
||
<emphasis id="B917EA88FF85FFEAFF6FFC46FE852577" box="[145,290,1009,1030]" italics="true" pageId="48" pageNumber="633">Distribution:</emphasis>
|
||
Upper
|
||
<collectingRegion id="49A7F878FF85FFEAFE7BFC46FE5C2577" box="[389,507,1009,1030]" country="Venezuela" name="Amazonas" pageId="48" pageNumber="633">Amazonas</collectingRegion>
|
||
in Juruá-Purus moist forest ecoregion (
|
||
<figureCitation id="13582A1FFF85FFEAFEADFBA7FE382554" box="[339,415,1040,1062]" captionStart="Figure 30" captionStartId="41.[163,241,928,950]" captionTargetBox="[165,1446,197,888]" captionTargetId="figure-513@41.[163,1446,195,889]" captionTargetPageId="41" captionText="Figure 30. Map of known distribution of Protus. Numbers correspond to the following WWF ecoregions: 1 = Ucayali moist forest, 2 = Iquitos varzea, 3 = Southwest Amazon moist forest, 4 = Solimões-Japurá moist forest, 5 = Juruá-Purus moist forest, 6 = Madeira-Tapajos moist forest, 7 = Japurá-Solimões-Negro moist forest, 8 = Uatuma-Trombetas moist forest, 9 = Mato Grosso seasonal forest." figureDoi="http://doi.org/10.5281/zenodo.6959592" httpUri="https://zenodo.org/record/6959592/files/figure.png" pageId="48" pageNumber="633">Fig. 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF6FFBEDFD9925DD" blockId="48.[145,761,1113,1840]" pageId="48" pageNumber="633">
|
||
<emphasis id="B917EA88FF85FFEAFF6FFBEDFEB225FC" italics="true" pageId="48" pageNumber="633">
|
||
Description of
|
||
<typeStatus id="54D88838FF85FFEAFEC4FBEEFE3D251F" box="[314,410,1113,1134]" pageId="48" pageNumber="633" type="holotype">holotype</typeStatus>
|
||
(with extra figures from other specimens):
|
||
</emphasis>
|
||
Measurements: CL: 1.62, CW: 2.55, AL: 3.47, AW: 4.44, IOD: 0.66, FeIV: 11.28.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF57FB01FE03248F" blockId="48.[145,761,1113,1840]" pageId="48" pageNumber="633">
|
||
Dorsum (
|
||
<figureCitation id="13582A1FFF85FFEAFEECFB02FE2925BA" box="[274,398,1205,1227]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35A–F</figureCitation>
|
||
). DS beta-type and lenticular in lateral view, without remarkable ornamentation, only low, coloured tubercles in areas I and
|
||
<collectionCode id="ED72AE5FFF85FFEAFDCBFB44FDF22479" box="[565,597,1267,1288]" pageId="48" pageNumber="633">III</collectionCode>
|
||
. Anterior edge of dorsal scutum with protoglyphs slightly concave with lateral rounded borders. Ocularium low and with a few granules near the eyes. Mesotergum convex, higher at level of area
|
||
<collectionCode id="ED72AE5FFF85FFEAFE6AFADAFE0E24F3" box="[404,425,1389,1410]" pageId="48" pageNumber="633">II</collectionCode>
|
||
, scutal areas I and
|
||
<collectionCode id="ED72AE5FFF85FFEAFD74FADAFD0F24F3" box="[650,680,1389,1410]" pageId="48" pageNumber="633">III</collectionCode>
|
||
armed with a pair of rounded tubercles, other areas unarmed. Posterior border and free tergites each containing a transverse row of minute granules, anal operculum with scattered granules.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF57F9B0FEFB2782" blockId="48.[145,761,1113,1840]" pageId="48" pageNumber="633">
|
||
Venter (
|
||
<figureCitation id="13582A1FFF85FFEAFEFAF9B1FEC5276D" box="[260,354,1542,1564]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35C</figureCitation>
|
||
). Coxae I–
|
||
<collectionCode id="ED72AE5FFF85FFEAFE22F9B0FE5E276D" box="[476,505,1543,1564]" pageId="48" pageNumber="633">III</collectionCode>
|
||
triangular, transverse to the main body axis, with a longitudinal row of granules, more remarkable in coxa I. Coxae
|
||
<collectionCode id="ED72AE5FFF85FFEAFD49F9F3FD6D2728" box="[695,714,1604,1625]" pageId="48" pageNumber="633">II</collectionCode>
|
||
–
|
||
<collectionCode id="ED72AE5FFF85FFEAFD27F9F3FD502728" box="[729,759,1604,1625]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
connected by tubercular bridges. Coxa
|
||
<collectionCode id="ED72AE5FFF85FFEAFDB1F9D4FDCC2709" box="[591,619,1635,1656]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
pentagonal, greatly developed, oriented obliquely, but almost parallel to the body axis. Stigmatic area T-shaped with stigmata large, unconcealed. Free sternites each with a row of granules.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFF57F94BFB702005" blockId="48.[145,761,1113,1840]" lastBlockId="48.[809,1425,197,1415]" pageId="48" pageNumber="633">
|
||
Chelicerae (
|
||
<figureCitation id="13582A1FFF85FFEAFEB2F94BFE422660" box="[332,485,1788,1810]" captionStart="Figure 36" captionStartId="47.[163,241,1787,1809]" captionTargetBox="[243,1362,196,1746]" captionTargetId="figure-25@47.[243,1363,195,1747]" captionTargetPageId="47" captionText="Figure 36. SEM images of some structures of Protus insolens, male (MPEG 867). A–C, lenis: A, dorsal view; B, lateral view; C, ventral view. D. Left basichelicerite, dorsal view. E, left cheliceral hand, dorsal view. F, left pedipalpal tarsus, ventral view. G, left pedipalpus, mesal view. H, left claw IV, ventral view. I, left claw III, ventro-lateral view. Scale bars: 100 μm (A–C, F, H, I), 200 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6959604" httpUri="https://zenodo.org/record/6959604/files/figure.png" pageId="48" pageNumber="633">Fig. 36D, E</figureCitation>
|
||
). Hand not swollen. Basichelicerite short, with well-marked bulla and uniformly tuberculated dorsally. Meso-distal corner of bulla with one setiferous tubercle larger than dorsal ones. Posterior and ectal margins of bulla fringed with many tubercles. Movable finger of cheliceral hand with ten trapezoidal tubercles. Fixed finger with six triangular tubercles of different sizes.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFCBFFECAFA872318" blockId="48.[809,1425,197,1415]" pageId="48" pageNumber="633">
|
||
Pedipalps (
|
||
<figureCitation id="13582A1FFF85FFEAFC2BFECAFBC120E3" box="[981,1126,381,403]" captionStart="Figure 36" captionStartId="47.[163,241,1787,1809]" captionTargetBox="[243,1362,196,1746]" captionTargetId="figure-25@47.[243,1363,195,1747]" captionTargetPageId="47" captionText="Figure 36. SEM images of some structures of Protus insolens, male (MPEG 867). A–C, lenis: A, dorsal view; B, lateral view; C, ventral view. D. Left basichelicerite, dorsal view. E, left cheliceral hand, dorsal view. F, left pedipalpal tarsus, ventral view. G, left pedipalpus, mesal view. H, left claw IV, ventral view. I, left claw III, ventro-lateral view. Scale bars: 100 μm (A–C, F, H, I), 200 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6959604" httpUri="https://zenodo.org/record/6959604/files/figure.png" pageId="48" pageNumber="633">Fig. 36F, G</figureCitation>
|
||
). Elongate trochanter, foliaceus femur; this is convex dorsally, with a dorsal row of seven setiferous tubercles and a ventral row of 14 setiferous tubercles. Tibia only slightly convex in ectal border, with a ventro-marginal row of small setae, mesal region three equally spaced large setae. Tarsus conical, with scattered dorsal setae and three well-marked ventral rows of subequal setae.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFCBFFDC4FC192250" blockId="48.[809,1425,197,1415]" pageId="48" pageNumber="633">
|
||
Legs (
|
||
<figureCitation id="13582A1FFF85FFEAFC76FDC5FBAF23F9" box="[904,1032,626,648]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35A–F</figureCitation>
|
||
). Long and unarmed legs, femora straight. Coxa
|
||
<collectionCode id="ED72AE5FFF85FFEAFC1BFD26FBA523D7" box="[997,1026,657,678]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
convex in dorsal view with few apical granules, no clavi inguines (also known as groin warts). Bipectinate claws in legs
|
||
<collectionCode id="ED72AE5FFF85FFEAFAE6FD78FA8A2395" box="[1304,1325,719,740]" pageId="48" pageNumber="633">II</collectionCode>
|
||
and
|
||
<collectionCode id="ED72AE5FFF85FFEAFA8EFD78FA2B2395" box="[1392,1420,719,740]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
, inner row with larger spines. Tarsal counts: 7(3), 16(3), 10, 11.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFCBFFC9DFB8622CA" blockId="48.[809,1425,197,1415]" pageId="48" pageNumber="633">
|
||
Colour (in alcohol,
|
||
<figureCitation id="13582A1FFF85FFEAFBCFFC9DFB1E2231" box="[1073,1209,810,832]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Fig. 35A–F</figureCitation>
|
||
). Dorsal scutum, venter and free tergites vivid orange (48), with paramedian rounded pale yellow (89) spots in areas I,
|
||
<collectionCode id="ED72AE5FFF85FFEAFC34FC30FC4922ED" box="[970,1006,903,924]" pageId="48" pageNumber="633">III</collectionCode>
|
||
and
|
||
<collectionCode id="ED72AE5FFF85FFEAFBC2FC30FBFA22ED" box="[1084,1117,903,924]" pageId="48" pageNumber="633">IV</collectionCode>
|
||
. Appendages brilliant greenish-yellow (98).
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFCBFFC74FB7E24F6" blockId="48.[809,1425,197,1415]" pageId="48" pageNumber="633">
|
||
<materialsCitation id="3B0B3CC7FF85FFEAFCBFFC74FB7224F6" ID-GBIF-Occurrence="3879212419" collectionCode="VP, MS-D, MS-A" pageId="48" pageNumber="633" specimenCode="MS-D1, MS-D2, T1" specimenCount="1">
|
||
Penis (
|
||
<figureCitation id="13582A1FFF85FFEAFC72FC74FBAA22A8" box="[908,1037,963,985]" captionStart="Figure 35" captionStartId="46.[145,223,1615,1637]" captionTargetBox="[145,1425,195,1575]" captionTargetId="figure-117@46.[145,1425,195,1575]" captionTargetPageId="46" captionText="Figure 35. Protus insolens, male holotype (MNHN ES 03021). A, dorsal view. B, panoramic dorsal view. C, ventral view. D, frontal view. E, posterior view. F, lateral view. G, penis, dorsal view. H, same, lateral view. Scale bars: 1 mm. Abbreviation: MS = macrosetae." figureDoi="http://doi.org/10.5281/zenodo.6959602" httpUri="https://zenodo.org/record/6959602/files/figure.png" pageId="48" pageNumber="633">Figs 35G, H</figureCitation>
|
||
,
|
||
<figureCitation id="13582A1FFF85FFEAFBE8FC74FBC422A8" box="[1046,1123,963,985]" captionStart="Figure 36" captionStartId="47.[163,241,1787,1809]" captionTargetBox="[243,1362,196,1746]" captionTargetId="figure-25@47.[243,1363,195,1747]" captionTargetPageId="47" captionText="Figure 36. SEM images of some structures of Protus insolens, male (MPEG 867). A–C, lenis: A, dorsal view; B, lateral view; C, ventral view. D. Left basichelicerite, dorsal view. E, left cheliceral hand, dorsal view. F, left pedipalpal tarsus, ventral view. G, left pedipalpus, mesal view. H, left claw IV, ventral view. I, left claw III, ventro-lateral view. Scale bars: 100 μm (A–C, F, H, I), 200 μm (D, E), 500 μm (G)." figureDoi="http://doi.org/10.5281/zenodo.6959604" httpUri="https://zenodo.org/record/6959604/files/figure.png" pageId="48" pageNumber="633">36A–C</figureCitation>
|
||
). Ventral plate rectangular elongate, with apical corners rounded, distal margin convex. Sub-distal lateral margin of
|
||
<collectionCode id="ED72AE5FFF85FFEAFB3AFBB6FB412567" box="[1220,1254,1025,1046]" pageId="48" pageNumber="633">VP</collectionCode>
|
||
with two pairs of MS-C, large, slender and conical, flattened at the tip. Two pairs of
|
||
<collectionCode id="ED72AE5FFF85FFEAFC42FB89FC592522" box="[956,1022,1086,1107]" pageId="48" pageNumber="633">MS-D</collectionCode>
|
||
located in apical half of
|
||
<collectionCode id="ED72AE5FFF85FFEAFAEBFB88FA912525" box="[1301,1334,1087,1108]" pageId="48" pageNumber="633">VP</collectionCode>
|
||
,
|
||
<specimenCode id="DBC59EE1FF85FFEAFABFFB89FA372522" box="[1345,1424,1086,1107]" collectionCode="MS-D" pageId="48" pageNumber="633">MS-D1</specimenCode>
|
||
twice longer than
|
||
<specimenCode id="DBC59EE1FF85FFEAFBFCFBEAFBF32503" box="[1026,1108,1117,1138]" collectionCode="MS-D" pageId="48" pageNumber="633">MS-D2</specimenCode>
|
||
and closer to MS-C2, both
|
||
<collectionCode id="ED72AE5FFF85FFEAFCD7FBCBFCCB25E0" box="[809,876,1148,1169]" pageId="48" pageNumber="633">MS-D</collectionCode>
|
||
straight and conical. One pair of
|
||
<collectionCode id="ED72AE5FFF85FFEAFB0AFBCBFA9125E0" box="[1268,1334,1148,1169]" pageId="48" pageNumber="633">MS-A</collectionCode>
|
||
basally in
|
||
<collectionCode id="ED72AE5FFF85FFEAFCBBFB2CFCC125C1" box="[837,870,1179,1200]" pageId="48" pageNumber="633">VP</collectionCode>
|
||
, straight, conical and as long as MS-C. Two pairs of MS-E in apical portion of lateroventral surface and one pair of MS-B in the basal portion of ventral surface of
|
||
<collectionCode id="ED72AE5FFF85FFEAFCBCFB40FCC3247D" box="[834,868,1271,1292]" pageId="48" pageNumber="633">VP</collectionCode>
|
||
. Ventral surface glabrous apically, two wide basal regions with microsetae
|
||
<typeStatus id="54D88838FF85FFEAFBB1FAA2FB25245B" box="[1103,1154,1301,1322]" pageId="48" pageNumber="633">type</typeStatus>
|
||
1 (
|
||
<specimenCode id="DBC59EE1FF85FFEAFB55FAA2FB6E245A" box="[1195,1225,1301,1323]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="48" pageNumber="633">T1</specimenCode>
|
||
) near MS-B and invading truncus. Glans with dorsal process thin and acuminate, stylus with a large wattle with an extended serrate part and without stylar barbs
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC369AFF85FFEAFCB8FA03FAD424BA" blockId="48.[838,1395,1460,1514]" box="[838,1395,1460,1483]" pageId="48" pageNumber="633">
|
||
<taxonomicName id="4C634D19FF85FFEAFCB8FA03FB8224BB" authorityName="Medrano, Kury & Mendes" authorityYear="2022" box="[838,1061,1460,1483]" class="Arachnida" family="Cosmetidae" genus="Protus" kingdom="Animalia" order="Opiliones" pageId="48" pageNumber="666" phylum="Arthropoda" rank="species" species="marllusi">
|
||
<emphasis id="B917EA88FF85FFEAFCB8FA03FB8224BB" bold="true" box="[838,1061,1460,1483]" italics="true" pageId="48" pageNumber="633">PROTUS MARLLUSI</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B917EA88FF85FFEAFBD2FA03FAD424BA" box="[1068,1395,1460,1483]" italics="true" pageId="48" pageNumber="633">
|
||
<collectionCode id="ED72AE5FFF85FFEAFBD2FA03FB0724BA" box="[1068,1184,1460,1483]" pageId="48" pageNumber="633">MEDRANO</collectionCode>
|
||
,
|
||
<collectionCode id="ED72AE5FFF85FFEAFB53FA03FB4A24BA" box="[1197,1261,1460,1483]" pageId="48" pageNumber="633">KURY</collectionCode>
|
||
&
|
||
<emphasis id="B917EA88FF85FFEAFAEFFA03FAD424BA" box="[1297,1395,1460,1483]" italics="true" pageId="48" pageNumber="633">
|
||
<collectionCode id="ED72AE5FFF85FFEAFAEFFA03FAD424BA" box="[1297,1395,1460,1483]" pageId="48" pageNumber="633">MENDES</collectionCode>
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |