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<document id="E8F24B340E9EC0A057E58DDB379E61D0" ID-DOI="10.11646/zootaxa.5433.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954420" ID-ZooBank="7ADF6A54-E31C-453A-AA25-44DEDBCF2752" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1712752719575" checkinUser="plazi" docAuthor="Souza, Leonardo Santos De, Araújo, Tarciso Almeida De &amp; Pimenta, Alexandre Dias" docDate="2024" docId="03CA878ABA47DE14FF4FFBEA7144F934" docLanguage="en" docName="zootaxa.5433.4.1.pdf" docOrigin="Zootaxa 5433 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graphis rhyssa Souza &amp; Araújo &amp; Pimenta 2024, comb. nov." docType="treatment" docVersion="3" lastPageNumber="492" masterDocId="FFF3FFF2BA6EDE3DFFD8FFD77042FF9F" masterDocTitle="Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena" masterLastPageNumber="499" masterPageNumber="451" pageNumber="492" updateTime="1712859506699" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="A2D2F4D599491BF5A7C64B2E01AC4FDB">Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena</mods:title>
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<mods:namePart id="4F0011814969874EB75B8B48426E5655">Souza, Leonardo Santos De</mods:namePart>
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<heading id="D09481F0BA47DE14FF4FFBEA7237FBC7" bold="true" box="[151,629,1085,1112]" fontSize="11" level="1" pageId="41" pageNumber="492" reason="1">
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<taxonomicName id="4C634D1FBA47DE14FF4FFBEA71A5FBC8" authority="(Dall, 1927)" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Dall" baseAuthorityYear="1927" box="[151,487,1085,1112]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="rhyssa" status="comb. nov.">
<emphasis id="B917EA8EBA47DE14FF4FFBEA710CFBC7" bold="true" box="[151,334,1085,1112]" italics="true" pageId="41" pageNumber="492">Graphis rhyssa</emphasis>
(
<bibRefCitation id="EFF24B6DBA47DE14FE86FBEA719CFBC8" author="Dall, W. H." box="[350,478,1085,1111]" pageId="41" pageNumber="492" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall, 1927</bibRefCitation>
)
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<taxonomicNameLabel id="A22457F5BA47DE14FE36FBE97237FBC7" box="[494,629,1086,1112]" pageId="41" pageNumber="492" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C3796517BA47DE14FF4FFBB6715DFBE3" box="[151,287,1121,1148]" pageId="41" pageNumber="492" type="description">
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<figureCitation id="13582A19BA47DE14FF4FFBB6715DFBE3" box="[151,287,1121,1148]" captionStart="FIGURE 24" captionStartId="41.[151,250,871,895]" captionTargetBox="[151,1435,181,847]" captionTargetId="figure-17@41.[151,1435,181,847]" captionTargetPageId="41" captionText="FIGURE 24. Species incorrectly attributed to Aclis. AC. Graphis rhyssa (Dall, 1927). Holotype, USNM 108389; DE. “Aclis gittenbergeri” (de Jong &amp; Coomans, 1988). Holotype, ZMA.MOLL.138263. A, DE. Shell in frontal view. B. Shell in dorsal view. C. Protoconch in apical view. Scale bars: AB, DE. 1 mm; C. 200 μm. Credits: AC. USNM; DE. Naturalis Biodiversity Center." figureDoi="http://doi.org/10.5281/zenodo.10954479" httpUri="https://zenodo.org/record/10954479/files/figure.png" pageId="41" pageNumber="492">Figs 24AC</figureCitation>
</paragraph>
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<paragraph id="8BDC369CBA47DE14FF4FFB7171F3FB21" blockId="41.[151,704,1190,1275]" box="[151,433,1190,1214]" pageId="41" pageNumber="492">
<treatmentCitationGroup id="AB7311B2BA47DE14FF4FFB7171F3FB21" box="[151,433,1190,1214]" pageId="41" pageNumber="492">
<treatmentCitation id="0AC2108DBA47DE14FF4FFB7171EFFB21" author="Dall, W. H." box="[151,429,1190,1214]" page="71" pageId="41" pageNumber="492" year="1927">
<taxonomicName id="4C634D1FBA47DE14FF4FFB7171EFFB21" ID-CoL="9D69" authority="Dall, 1927: 71" authorityName="Dall" authorityPageNumber="71" authorityYear="1927" box="[151,429,1190,1214]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="rhyssa">
<emphasis id="B917EA8EBA47DE14FF4FFB717150FB21" box="[151,274,1190,1214]" italics="true" pageId="41" pageNumber="492">Aclis rhyssa</emphasis>
<bibRefCitation id="EFF24B6DBA47DE14FEC0FB7171EFFB21" author="Dall, W. H." box="[280,429,1190,1214]" pageId="41" pageNumber="492" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall, 1927: 71</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BDC369CBA47DE14FF4FFB127282FB42" blockId="41.[151,704,1190,1275]" box="[151,704,1221,1245]" pageId="41" pageNumber="492">
<treatmentCitationGroup id="AB7311B2BA47DE14FF4FFB127282FB42" box="[151,704,1221,1245]" pageId="41" pageNumber="492">
<taxonomicName id="4C634D1FBA47DE14FF4FFB127159FB42" ID-CoL="64JXT" authorityName="Bartsch" authorityYear="1911" box="[151,283,1221,1245]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="cubana">
<emphasis id="B917EA8EBA47DE14FF4FFB127159FB42" box="[151,283,1221,1245]" italics="true" pageId="41" pageNumber="492">Aclis cubana</emphasis>
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auct. non
<bibRefCitation id="EFF24B6DBA47DE14FE5CFB127251FB42" author="Bartsch, P." box="[388,531,1221,1245]" pageId="41" pageNumber="492" pagination="435 - 438" refId="ref23751" refString="Bartsch, P. (1911) New mollusks of the genus Aclis from the North Atlantic. Proceedings of the United States National Museum, 40 (1829), 435 - 438. https: // doi. org / 10.5479 / si. 00963801.40 - 1829.435" type="journal article" year="1911">Bartsch, 1911</bibRefCitation>
:
<treatmentCitation id="0AC2108DBA47DE14FDC5FB1272FEFB42" author="Dall, W. H." box="[541,700,1221,1245]" page="71" pageId="41" pageNumber="492" year="1927">
<bibRefCitation id="EFF24B6DBA47DE14FDC5FB1272FEFB42" author="Dall, W. H." box="[541,700,1221,1245]" pageId="41" pageNumber="492" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall (1927: 71)</bibRefCitation>
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.
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<paragraph id="8BDC369CBA47DE14FF4FFB3472C6FB64" blockId="41.[151,704,1190,1275]" box="[151,644,1251,1275]" pageId="41" pageNumber="492">
<treatmentCitationGroup id="AB7311B2BA47DE14FF4FFB3472C6FB64" box="[151,644,1251,1275]" pageId="41" pageNumber="492">
<taxonomicName id="4C634D1FBA47DE14FF4FFB34717CFB64" baseAuthorityName="R.B.Watson" baseAuthorityYear="1881" box="[151,318,1251,1275]" class="Gastropoda" family="Eulimidae" genus="Costaclis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="rhyssa">
<emphasis id="B917EA8EBA47DE14FF4FFB34717CFB64" box="[151,318,1251,1275]" italics="true" pageId="41" pageNumber="492">Costaclis rhyssa</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AC2108DBA47DE14FE93FB3471B7FB64" author="Bartsch, P." box="[331,501,1251,1275]" page="9" pageId="41" pageNumber="492" year="1947">
<bibRefCitation id="EFF24B6DBA47DE14FE93FB3471B7FB64" author="Bartsch, P." box="[331,501,1251,1275]" pageId="41" pageNumber="492" pagination="1 - 29" refId="ref23800" refString="Bartsch, P. (1947) A monograph of the west Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29." type="journal article" year="1947">Bartsch (1947: 9</bibRefCitation>
</treatmentCitation>
, pl. 1,
<figureCitation id="13582A19BA47DE14FDE2FB347235FB64" box="[570,631,1251,1275]" captionStart="FIGURE 5" captionStartId="12.[151,250,1825,1849]" captionTargetBox="[151,1435,597,1801]" captionTargetId="figure-218@12.[151,1435,597,1801]" captionTargetPageId="12" captionText="FIGURE 5. Aclis kanelaAbsalão, 2009. A, DG. MNRJ 28293. BC. MNRJ 26713. AC. Shell in frontal view; rectangle in A indicates detail in E. D. Detail of body whorl in frontal view. E, F. Detail of teleoconch sculpture; rectangle in E indicates detail in F. G. Detail of apical whorls. Scale bars: AC. 1 mm; D. 500 μm; EF. 50 μm; G. 200 μm." figureDoi="http://doi.org/10.5281/zenodo.10954434" httpUri="https://zenodo.org/record/10954434/files/figure.png" pageId="41" pageNumber="492">Fig. 5</figureCitation>
).
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<emphasis id="B917EA8EBA47DE14FF4FFAFD7107FADC" bold="true" box="[151,325,1321,1348]" pageId="41" pageNumber="492">Type material.</emphasis>
<materialsCitation id="3B0B3CC1BA47DE14FE94FAFE7308FADB" box="[332,842,1321,1348]" collectionCode="USNM" pageId="41" pageNumber="492" specimenCode="USNM 108389" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D8883EBA47DE14FE94FAFE71F6FADC" box="[332,436,1321,1347]" pageId="41" pageNumber="492" type="holotype">Holotype</typeStatus>
(herein examined):
<specimenCode id="DBC59EE7BA47DE14FD4EFAFD7304FADB" box="[662,838,1322,1348]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="41" pageNumber="492" type="Museum">USNM 108389</specimenCode>
.
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<paragraph id="8BDC369CBA47DE14FF1FFA99710DFA13" blockId="41.[151,1437,1321,1707]" pageId="41" pageNumber="492">
<materialsCitation id="3B0B3CC1BA47DE14FF1FFA997109FA13" collectingDate="1885-04-01" collectorName="Steamer Albatross" country="United States of America" elevation="805" latitude="30.733334" location="Fernandina" longLatPrecision="1220" longitude="-79.433334" pageId="41" pageNumber="492" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
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<typeStatus id="54D8883EBA47DE14FF1FFA997140FAF7" box="[199,258,1358,1384]" pageId="41" pageNumber="492">Type</typeStatus>
locality.
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<collectingCountry id="F374760CBA47DE14FEB0FA9971E6FAF7" box="[360,420,1358,1384]" name="United States of America" pageId="41" pageNumber="492">USA</collectingCountry>
:
<collectingRegion id="49A7F87EBA47DE14FE74FA9A7278FAF7" box="[428,570,1357,1384]" country="United States of America" name="Florida" pageId="41" pageNumber="492">Florida state</collectingRegion>
: off
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, USFC sta. 2415;
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<elevation id="004ED1AFBA47DE14FB58FA9A748AFAF7" box="[1152,1224,1357,1384]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="41" pageNumber="492" unit="m" value="805.0">805 m</elevation>
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;
<collectorName id="2696534ABA47DE14FB09FA9975DEFAF8" box="[1233,1436,1357,1384]" pageId="41" pageNumber="492">
Steamer
<emphasis id="B917EA8EBA47DE14FAEAFA9A75DEFAF8" box="[1330,1436,1357,1383]" italics="true" pageId="41" pageNumber="492">Albatross</emphasis>
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col.,
<date id="FFDD105CBA47DE14FF16FAA67109FA13" box="[206,331,1393,1420]" pageId="41" pageNumber="492" value="1885-04-01">
<collectingDate id="EF99E9B4BA47DE14FF16FAA67109FA13" box="[206,331,1393,1420]" pageId="41" pageNumber="492" value="1885-04-01">01/iv/1885</collectingDate>
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.
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<emphasis id="B917EA8EBA47DE14FF1FFA4271F3FA30" bold="true" box="[199,433,1429,1455]" pageId="41" pageNumber="492">Material examined.</emphasis>
The
<typeStatus id="54D8883EBA47DE14FE32FA427212FA30" box="[490,592,1429,1455]" pageId="41" pageNumber="492" type="holotype">holotype</typeStatus>
.
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<subSubSection id="C3796517BA47DE14FF1FFA6D7144F934" pageId="41" pageNumber="492" type="discussion">
<paragraph id="8BDC369CBA47DE14FF1FFA6D7418F984" blockId="41.[151,1437,1321,1707]" pageId="41" pageNumber="492">
<emphasis id="B917EA8EBA47DE14FF1FFA6D7179FA4B" bold="true" box="[199,315,1466,1492]" pageId="41" pageNumber="492">Remarks.</emphasis>
<bibRefCitation id="EFF24B6DBA47DE14FE9BFA6E7184FA4B" author="Dall, W. H." box="[323,454,1465,1492]" pageId="41" pageNumber="492" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall (1927)</bibRefCitation>
clearly stated that the shell of USNM 108389 is the type specimen. Thus, it should be considered the
<typeStatus id="54D8883EBA47DE14FE99FA0A71E1FA68" box="[321,419,1501,1527]" pageId="41" pageNumber="492" type="holotype">holotype</typeStatus>
and the type locality the USFC sta. 2415 (Off Fernandina), as written in the original label. The author also refers to several other specimens from USFC sta. 2668 (Off
<collectingCountry id="F374760CBA47DE14FC28F9D6740DF984" box="[1008,1103,1537,1563]" name="Georgia" pageId="41" pageNumber="492">Georgia</collectingCountry>
).
</paragraph>
<paragraph id="8BDC369CBA47DE14FF1FF9F27144F934" blockId="41.[151,1437,1321,1707]" pageId="41" pageNumber="492">
<bibRefCitation id="EFF24B6DBA47DE14FF1FF9F27133F9DF" author="Bartsch, P." box="[199,369,1573,1600]" pageId="41" pageNumber="492" pagination="1 - 29" refId="ref23800" refString="Bartsch, P. (1947) A monograph of the west Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections, 106 (20), 1 - 29." type="journal article" year="1947">Bartsch (1947)</bibRefCitation>
included the present species in
<taxonomicName id="4C634D1FBA47DE14FD3BF9F27309F9A0" authorityName="Bartsch" authorityYear="1947" box="[739,843,1573,1599]" class="Gastropoda" family="Eulimidae" genus="Costaclis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="genus">
<emphasis id="B917EA8EBA47DE14FD3BF9F27309F9A0" box="[739,843,1573,1599]" italics="true" pageId="41" pageNumber="492">Costaclis</emphasis>
</taxonomicName>
, which was not adopted later by
<bibRefCitation id="EFF24B6DBA47DE14FB16F9F270A7F9FC" author="Bouchet, P. &amp; Waren, A." pageId="41" pageNumber="492" pagination="299 - 576" refId="ref23963" refString="Bouchet, P. &amp; Waren, A. (1986) Revision of the Northeast Atlantic Bathyal and Abyssal Aclididae, Eulimidae, Epitoniidae (Mollusca, Gastropoda). Bollettino Malacologico, Supplemento 2, 299 - 576. https: // doi. org / 10.5962 / bhl. title. 140762" type="journal article" year="1986">Bouchet &amp; Warén (1986)</bibRefCitation>
. A new combination of the present species in
<taxonomicName id="4C634D1FBA47DE14FD24F99E731AF9FC" authorityName="Jeffreys" authorityYear="1867" box="[764,856,1609,1635]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="genus">
<emphasis id="B917EA8EBA47DE14FD24F99E731AF9FC" box="[764,856,1609,1635]" italics="true" pageId="41" pageNumber="492">Graphis</emphasis>
</taxonomicName>
is proposed here, similarly to the case of
<taxonomicName id="4C634D1FBA47DE14FA98F99E70ABF918" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Bartsch" baseAuthorityYear="1911" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="cubana">
<emphasis id="B917EA8EBA47DE14FA98F99E70ABF918" italics="true" pageId="41" pageNumber="492">Graphis cubana</emphasis>
</taxonomicName>
commented above.
<taxonomicName id="4C634D1FBA47DE14FE17F9BA734DF917" authority="(Dall, 1927)" authorityName="Souza &amp; Araújo &amp; Pimenta" authorityYear="2024" baseAuthorityName="Dall" baseAuthorityYear="1927" box="[463,783,1645,1672]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Littorinimorpha" pageId="41" pageNumber="492" phylum="Mollusca" rank="species" species="rhyssa" status="comb. nov.">
<emphasis id="B917EA8EBA47DE14FE17F9BA723FF918" box="[463,637,1645,1671]" italics="true" pageId="41" pageNumber="492">Graphis rhyssa</emphasis>
(
<bibRefCitation id="EFF24B6DBA47DE14FD55F9BA7345F917" author="Dall, W. H." box="[653,775,1645,1672]" pageId="41" pageNumber="492" pagination="1 - 134" refId="ref24604" refString="Dall, W. H. (1927) Small shells from dredgings off the southeast coast of the United States by the United States Fisheries Steamer ' Albatross' in 1885 and 1886. Proceedings of the United States National Museum, 70 (2667), 1 - 134. https: // doi. org / 10.5479 / si. 00963801.70 - 2667.1" type="journal article" year="1927">Dall, 1927</bibRefCitation>
)
</taxonomicName>
<emphasis id="B917EA8EBA47DE14FCC0F9B973DBF917" bold="true" box="[792,921,1646,1672]" pageId="41" pageNumber="492">
<taxonomicNameLabel id="A22457F5BA47DE14FCC0F9B973DBF917" box="[792,921,1646,1672]" pageId="41" pageNumber="492" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
also has strong axial ribs but lacks the spiral sculpture.
</paragraph>
</subSubSection>
</treatment>
</document>