794 lines
91 KiB
XML
794 lines
91 KiB
XML
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<mods:title id="8EC68B1CA3EB4629D32EB7F66861495E">Fetilinia dentator gen. et sp. n. from Pakistan (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="48AF6B75389054DA0486BF648786AC13">Lowe, Graeme</mods:namePart>
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<mods:namePart id="806E30F6B16316953ABE1D06F71F7925">Kovařík, František</mods:namePart>
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<mods:title id="95D3A699C7D61664F45E66EA481D6429">Euscorpius</mods:title>
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<mods:date id="D6FABF1F471BBE5ADCA6FA9383689775">2020</mods:date>
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<taxonomicName id="4C6039BCFFC8C519B7243D2DFB30FB95" ID-CoL="8LYXG" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1049,1153,1031,1058]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">Fetilinia</taxonomicName>
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.
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(
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<figureCitation id="135B5EBAFFC8C519B7383D03FB24FBF5" box="[1029,1173,1065,1090]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–9" captionStart-2="Figures 10–23" captionStart-3="Figures 24–29" captionStart-4="Figures 30–31" captionStartId-0="3.[113,190,2005,2027]" captionStartId-1="6.[115,192,1967,1989]" captionStartId-2="7.[113,190,1916,1938]" captionStartId-3="8.[113,190,1172,1194]" captionStartId-4="9.[113,190,2010,2032]" captionTargetBox-0="[103,1465,335,1870]" captionTargetBox-1="[113,1474,244,1881]" captionTargetBox-2="[113,1454,234,1891]" captionTargetBox-3="[113,1467,245,1156]" captionTargetBox-4="[113,1474,282,1949]" captionTargetId-0="figure-35@3.[83,1483,319,1878]" captionTargetId-1="figure-70@6.[113,1474,244,1895]" captionTargetId-2="figure-99@7.[113,1474,234,1895]" captionTargetId-3="figure-498@8.[113,1474,234,1156]" captionTargetId-4="figure-34@9.[113,1474,282,1949]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Figures 1–2. Fetilinia dentator gen. et sp. n., subadult male holotype in dorsal (1) and ventral (2) views. Scale bar: 10 mm." captionText-1="Figures 3–9. Fetilinia dentator gen. et sp. n., subadult male holotype, metasoma IV–V and telson ventral view (3), metasoma and telson dorsal (4, 7), ventral (5, 8), and lateral (6) views, metasoma V and telson lateral view (9). Figures 3–5 are under UV light. Scale bar: 10 mm (4–8)." captionText-2="Figures 10–23: Fetilinia dentator gen. et sp. n., subadult male holotype. Figures 10–12. Carapace and tergites (10) and sternopectinal area and sternites (11–12) Figures 13–23. Pedipalp segments, chela dorsal (14), eXternal (15), ventral (16) views, patella dorsal (17), dorsoeXternal (18), and ventral (19) views, femur and trochanter dorsoeXternal (20), internal (21), and ventral (22) views, movable (13) and fiXed (23) finger dentition. Trichobothrial pattern is indicated by white circles in Figures 15–18 and 20–21. Figures 10–11 under UV light." captionText-3="Figures 24–29: Fetilinia dentator gen. et sp. n., subadult male holotype. Figures 24–25. Chelicerae ventral (24) and chelicerae dorsal with anterior part of carapace with median and lateral eyes (25). Figures 26–29. Left legs I–IV, retrolateral aspect (respectively)." captionText-4="Figures 30–31. Fetilinia dentator gen. et sp. n., subadult male holotype in dorsal (30) and ventral (31) views, under UV light." figureDoi-0="http://doi.org/10.5281/zenodo.4648978" figureDoi-1="http://doi.org/10.5281/zenodo.4648980" figureDoi-2="http://doi.org/10.5281/zenodo.4648982" figureDoi-3="http://doi.org/10.5281/zenodo.4648984" figureDoi-4="http://doi.org/10.5281/zenodo.4648986" httpUri-0="https://zenodo.org/record/4648978/files/figure.png" httpUri-1="https://zenodo.org/record/4648980/files/figure.png" httpUri-2="https://zenodo.org/record/4648982/files/figure.png" httpUri-3="https://zenodo.org/record/4648984/files/figure.png" httpUri-4="https://zenodo.org/record/4648986/files/figure.png" pageId="2" pageNumber="1">Figures 1–31</figureCitation>
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,
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<tableCitation id="C6E27784FFC8C519B7A23D03FB5CFBF5" box="[1183,1261,1065,1090]" captionStart="Table 1" captionStartId="4.[115,171,977,998]" captionTargetId="graphics-630@4.[118,1476,227,965]" captionTargetPageId="4" captionText="Table 1. Comparative measurements of types of Fetilinia dentator gen. et sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/DF1F12B7FFCEC51FB34E3AFBFE24FBB3" pageId="2" pageNumber="1" tableUuid="DF1F12B7FFCEC51FB34E3AFBFE24FBB3">Table 1</tableCitation>
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)
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</paragraph>
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<emphasis id="B9149E2DFFC8C519B0033D60FCDCFBD5" bold="true" box="[830,877,1098,1122]" pageId="2" pageNumber="1">http</emphasis>
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:
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.
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:
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:
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<emphasis id="B9149E2DFFC8C519B7DD3D60FAB6FBD5" bold="true" box="[1248,1287,1098,1122]" pageId="2" pageNumber="1">org</emphasis>
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:
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:
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<paragraph id="8BDF423FFFC8C519B00F3D80FB79FB76" blockId="2.[818,1224,1193,1218]" box="[818,1224,1193,1218]" pageId="2" pageNumber="1">
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.
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<taxonomicName id="4C6039BCFFC8C519B0F43D83FB34FB76" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[969,1157,1193,1217]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="dentator" status="sp. nov.">
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<emphasis id="B9149E2DFFC8C519B0F43D83FB34FB76" box="[969,1157,1193,1217]" italics="true" pageId="2" pageNumber="1">Fetilinia dentator</emphasis>
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.
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<emphasis id="B9149E2DFFC8C519B78E3D80FB73FB75" bold="true" box="[1203,1218,1194,1218]" pageId="2" pageNumber="1">n</emphasis>
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.
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<subSubSection id="C37A11B4FFC8C519B00F3DDAFB53F930" pageId="2" pageNumber="1" type="etymology">
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<paragraph id="8BDF423FFFC8C519B00F3DDAFB53F930" blockId="2.[818,1476,1264,1672]" pageId="2" pageNumber="1">
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ETYMOLOGY. The generic epithet
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<taxonomicName id="4C6039BCFFC8C519B7E93DDAFA81FABF" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1236,1328,1264,1288]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B9149E2DFFC8C519B7E93DDAFA81FABF" box="[1236,1328,1264,1288]" italics="true" pageId="2" pageNumber="1">Fetilinia</emphasis>
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</taxonomicName>
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(masculine; constructed to rhyme with
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<taxonomicName id="4C6039BCFFC8C519B75F3C3AFB50FA9F" authorityName="Vachon" authorityYear="1974" box="[1122,1249,1296,1320]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B9149E2DFFC8C519B75F3C3AFB50FA9F" box="[1122,1249,1296,1320]" italics="true" pageId="2" pageNumber="1">Kraepelinia</emphasis>
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</taxonomicName>
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, a similar genus) is a patronym honoring Victor Fet (
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<collectingCountry id="F37702AFFFC8C519B7A13C1AFB65FAFF" box="[1180,1236,1328,1352]" name="United States of America" pageId="2" pageNumber="1">USA</collectingCountry>
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||
) in recognition of his many important contributions to the knowledge of scorpions and his generous support of many scorpiologists.
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<taxonomicName id="4C6039BCFFC8C519B6793C5AFA72FA3F" authorityName="Vachon" authorityYear="1974" box="[1348,1475,1392,1416]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B9149E2DFFC8C519B6793C5AFA72FA3F" box="[1348,1475,1392,1416]" italics="true" pageId="2" pageNumber="1">Kraepelinia</emphasis>
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</taxonomicName>
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is a very special genus for VF who published its first record from
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<collectingCountry id="F37702AFFFC8C519B0513C9AFC4EFA7F" box="[876,1023,1456,1480]" name="Turkmenistan" pageId="2" pageNumber="1">Turkmenistan</collectingCountry>
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where he started his scorpiological career (
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<bibRefCitation id="EFF13FCEFFC8C519B0043CFAFC1BFA5F" author="FET" box="[825,938,1488,1512]" pageId="2" pageNumber="1" pagination="37 - 43" refId="ref4521" refString="FET, V. 1984. [New for the USSR genus and species of scorpions from Badhyz: Kraepelinia palpator (Birula, 1903) (Scorpiones, Buthidae)]. Izvestiia Akademii Nauk Turkmenskoi SSR, ser. biol. nauk [Proceedings of the Academy of Sciences of the Turkmen SSR, Ser. Biol. Sci.], 4: 37 - 43 (in Russian)." type="journal article" year="1984">Fet, 1984</bibRefCitation>
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). At the time, the only known specimen of
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<taxonomicName id="4C6039BCFFC8C519B6963CFBFB80F9BF" authority="(Birula, 1903)" baseAuthorityName="Birula" baseAuthorityYear="1903" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="palpator">
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<emphasis id="B9149E2DFFC8C519B6963CFBFC3FF9BF" italics="true" pageId="2" pageNumber="1">K. palpator</emphasis>
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(
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<bibRefCitation id="EFF13FCEFFC8C519B0A33CDAFB98F9BF" author="BIRULA" box="[926,1065,1520,1544]" pageId="2" pageNumber="1" pagination="67 - 80" refId="ref4476" refString="BIRULA, A. A. 1903. Beitrage zur Kenntniss der Scorpionenfauna Ost-Persiens. (2 Beitrag). Bulletin de l'Academie Imperiale des Sciences de Saint. - Petersbourg, ser. 5, 19 (2): 67 - 80." type="journal article" year="1903">Birula, 1903</bibRefCitation>
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)
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</taxonomicName>
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from
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<collectingCountry id="F37702AFFFC8C519B74B3CDAFAB8F9BF" box="[1142,1289,1520,1544]" name="Turkmenistan" pageId="2" pageNumber="1">Turkmenistan</collectingCountry>
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was collected in 1972 by Yuri Gorelov (1933–2018), an outstanding naturalist, whose life and work in Central Asia (Badghyz Natural Reserve) has been an inspiration and role model for many young Russian zoologists, including VF.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A11B4FFC8C51FB00F3F9CFA18F84E" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1" type="diagnosis">
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<paragraph id="8BDF423FFFC8C51FB00F3F9CFEA6F8EE" blockId="2.[818,1477,1718,2030]" lastBlockId="4.[113,773,1057,1881]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
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DIAGNOSIS.Adult size small, total length of subadult male
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<quantity id="4C98EFDAFFC8C519B64B3F9CFA71F979" box="[1398,1472,1718,1742]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="1" unit="mm" value="22.0">22 mm</quantity>
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. Carapace trapezoidal, densely granulate, without carinae, anterior margin almost straight; median eyes large, ocular tubercle raised; five pairs of lateral eyes present, two reduced in size. Sternum
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<typeStatus id="54DBFC9DFFC8C519B00F3E1DFCEFF8F8" box="[818,862,1847,1871]" pageId="2" pageNumber="1">type</typeStatus>
|
||
1, relatively small, and triangular to subpentagonal in shape; posterior depression very large and deep. Pectines large, with 23–23 teeth in subadult male and 19–19 teeth in juvenile female; fulcra present. Tergites I–VI tricarinate, all tergites densely granulate. Sternites with slit-like spiracles; sternite V without a well-defined smooth patch; sternite VII with weak carinae. Metasomal segments relatively short and stout, nearly uniform in width, with carination well developed; metasoma I–III with 8–10 carinae; metasoma II–III with enlarged dentition on ventrolateral and ventromedian carinae; metasoma V with irregular, enlarged lobate dentition on ventrolateral carinae; posterior margins of tergite VII and metasoma I–III with fine fringes of microsetae. Telson with elongate vesicle, aculeus stout, about same length as vesicle, subaculear tubercle absent. Cheliceral dentition follows typical buthid pattern (Vachon, 1963), fiXed finger with two denticles on ventral surface. Pedipalps slender, chelae narrower than patella; trichobothrial pattern neobothriotaXic
|
||
<typeStatus id="54DBFC9DFFCEC51FB34C3CABFF2CFA2E" box="[113,157,1409,1433]" pageId="4" pageNumber="3">type</typeStatus>
|
||
C, femur with trichobothrium
|
||
<emphasis id="B9149E2DFFCEC51FB2D83CABFE4BFA28" box="[485,506,1409,1439]" italics="true" pageId="4" pageNumber="3">
|
||
d
|
||
<subScript id="17E4407AFFCEC51FB2CF3CBBFE4BFA28" attach="left" box="[498,506,1425,1439]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
</emphasis>
|
||
absent,
|
||
<emphasis id="B9149E2DFFCEC51FB16F3CABFDEEFA2E" box="[594,607,1409,1433]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB1623CBBFDD7FA28" attach="left" box="[607,614,1425,1439]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB1523CABFDCDFA2E" box="[623,636,1409,1433]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB1413CBBFD32FA28" attach="left" box="[636,643,1425,1439]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB1B13CABFD28FA2E" box="[652,665,1409,1433]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB1A43CBBFD10FA28" attach="left" box="[665,673,1425,1439]" fontSize="6" pageId="4" pageNumber="3">4</subScript>
|
||
arranged in b- configuration (non-refleX angle opening internally), patella
|
||
<emphasis id="B9149E2DFFCEC51FB34C3CEBFF37FA68" box="[113,134,1473,1503]" italics="true" pageId="4" pageNumber="3">
|
||
d
|
||
<subScript id="17E4407AFFCEC51FB3433CFBFF37FA68" attach="left" box="[126,134,1489,1503]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
</emphasis>
|
||
located between dorsomedian and dorsointernal carinae, chela manus
|
||
<emphasis id="B9149E2DFFCEC51FB3C13CCBFEA8FA4E" box="[252,281,1505,1529]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB2253CDBFEAEFA48" attach="left" box="[280,287,1521,1535]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB20C3CCBFEFFFA4E" box="[305,334,1505,1529]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB2703CDBFEE5FA48" attach="left" box="[333,340,1521,1535]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB25B3CCBFE32FA4E" box="[358,387,1505,1529]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<quantity id="4C98EFDAFFCEC51FB2BE3CDBFE12FA4E" box="[387,419,1505,1535]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="3" unit="in" value="3.0">
|
||
<subScript id="17E4407AFFCEC51FB2BE3CDBFE3AFA48" attach="left" box="[387,395,1521,1535]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
in
|
||
</quantity>
|
||
δ-configuration (non-refleX angle opening distally),
|
||
<emphasis id="B9149E2DFFCEC51FB20C3F2BFEF0F9AF" box="[305,321,1537,1560]" italics="true" pageId="4" pageNumber="3">V</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB27C3F3BFEF9F9A8" attach="left" box="[321,328,1553,1567]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB26C3F2BFED0F9AF" box="[337,353,1537,1560]" italics="true" pageId="4" pageNumber="3">V</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB25C3F3BFED8F9A8" attach="left" box="[353,361,1553,1567]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
aXis slightly inclined internally,
|
||
<emphasis id="B9149E2DFFCEC51FB1FA3F2BFD51F9AE" box="[711,736,1537,1561]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
|
||
on distal manus (not fiXed finger), fiXed finger with
|
||
<emphasis id="B9149E2DFFCEC51FB15C3F0BFDCAF98E" box="[609,635,1569,1593]" italics="true" pageId="4" pageNumber="3">db</emphasis>
|
||
situated near base of finger and distal to
|
||
<emphasis id="B9149E2DFFCEC51FB2B83F6BFE10F9EF" box="[389,417,1601,1624]" italics="true" pageId="4" pageNumber="3">est</emphasis>
|
||
,
|
||
<emphasis id="B9149E2DFFCEC51FB2933F6BFE73F9EE" box="[430,450,1601,1625]" italics="true" pageId="4" pageNumber="3">dt</emphasis>
|
||
at mid-finger and level with
|
||
<emphasis id="B9149E2DFFCEC51FB1D43F6BFD4AF9EF" box="[745,763,1601,1624]" italics="true" pageId="4" pageNumber="3">et</emphasis>
|
||
; dentate margins of pedipalp fingers straight, without lobe/ notch combination, equipped with 8 rows of median denticles arranged nearly linearly, non-imbricated, each flanked by a single eXternal and internal accessory granule; 5 subterminal granules. Legs I– III with tibia and tarsi short, curved, flat, with setation modified into bristlecombs on basitarsi only, telotarsi with two rows of long setae on ventral surface; tibial spurs moderate, tarsal spurs well-developed.
|
||
</paragraph>
|
||
<caption id="DF1F12B7FFC9C518B34C3EFFFAA7F85C" ID-DOI="http://doi.org/10.5281/zenodo.4648978" ID-Zenodo-Dep="4648978" box="[113,1302,2005,2027]" httpUri="https://zenodo.org/record/4648978/files/figure.png" pageId="3" pageNumber="2" startId="3.[113,190,2005,2027]" targetBox="[103,1465,335,1870]" targetPageId="3">
|
||
<paragraph id="8BDF423FFFC9C518B34C3EFFFAA7F85C" blockId="3.[113,1302,2005,2027]" box="[113,1302,2005,2027]" pageId="3" pageNumber="2">
|
||
<emphasis id="B9149E2DFFC9C518B34C3EFFFF59F85D" bold="true" box="[113,232,2005,2027]" pageId="3" pageNumber="2">Figures 1–2</emphasis>
|
||
.
|
||
<taxonomicName id="4C6039BCFFC9C518B3C93EFFFE2FF85D" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[244,414,2005,2026]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="dentator" status="gen. et sp. n.">
|
||
<emphasis id="B9149E2DFFC9C518B3C93EFFFE2FF85D" box="[244,414,2005,2026]" italics="true" pageId="3" pageNumber="2">Fetilinia dentator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFC9C518B2993EFFFD90F85C" box="[420,545,2005,2027]" pageId="3" pageNumber="2" rank="species">
|
||
<emphasis id="B9149E2DFFC9C518B2993EFFFE79F85D" bold="true" box="[420,456,2005,2026]" pageId="3" pageNumber="2">gen</emphasis>
|
||
. et
|
||
<emphasis id="B9149E2DFFC9C518B2D13EFFFDB2F85D" bold="true" box="[492,515,2005,2026]" pageId="3" pageNumber="2">sp</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFC9C518B1333EFFFDAAF85D" bold="true" box="[526,539,2005,2026]" pageId="3" pageNumber="2">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, subadult male holotype in dorsal (1) and ventral (2) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDF423FFFCEC51FB1E439C7FA90FC0A" pageId="4" pageNumber="3">
|
||
<table id="F960B09FFFCE3AE4B34139C7FA01FC77" box="[124,1456,237,960]" gridcols="4" gridrows="19" pageId="4" pageNumber="3">
|
||
<tr id="3550407DFFCE3AE4B34139C7FA01FEB2" box="[124,1456,237,261]" gridrow="0" pageId="4" pageNumber="3" rowspan-0="1" rowspan-1="1">
|
||
<th id="76812901FFCE3AE4B1E439C7FB81FEB2" box="[729,1072,237,261]" gridcol="2" gridrow="0" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C6039BCFFCEC51FB1E439C7FC29FEB2" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[729,920,237,261]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="dentator" status="gen. et sp. n.">
|
||
<emphasis id="B9149E2DFFCEC51FB1E439C7FC29FEB2" bold="true" box="[729,920,237,261]" italics="true" pageId="4" pageNumber="3">Fetilinia dentator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB09939C7FB81FEB2" box="[932,1072,237,261]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B9149E2DFFCEC51FB09939C7FBBCFEB2" bold="true" box="[932,1037,237,261]" pageId="4" pageNumber="3">gen. et sp</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB72739C7FB98FEB2" bold="true" box="[1050,1065,237,261]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
<th id="76812901FFCE3AE4B76439C7FA01FEB2" box="[1113,1456,237,261]" gridcol="3" gridrow="0" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C6039BCFFCEC51FB76439C7FAA9FEB2" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1113,1304,237,261]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="dentator" status="gen. et sp. n.">
|
||
<emphasis id="B9149E2DFFCEC51FB76439C7FAA9FEB2" bold="true" box="[1113,1304,237,261]" italics="true" pageId="4" pageNumber="3">Fetilinia dentator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB61939C7FA01FEB2" box="[1316,1456,237,261]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B9149E2DFFCEC51FB61939C7FA3CFEB2" bold="true" box="[1316,1421,237,261]" pageId="4" pageNumber="3">gen. et sp</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB6A739C7FA18FEB2" bold="true" box="[1434,1449,237,261]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B341383FFA01FE99" box="[124,1456,277,302]" gridrow="1" pageId="4" pageNumber="3" rowspan-1="1">
|
||
<th id="76812901FFCE3AE4B341383FFEF7FE99" box="[124,326,277,302]" gridcol="0" gridrow="1" pageId="4" pageNumber="3">
|
||
<emphasis id="B9149E2DFFCEC51FB341383FFEF1FE9A" bold="true" box="[124,320,277,301]" pageId="4" pageNumber="3">Dimensions (mm)</emphasis>
|
||
</th>
|
||
<td id="76812901FFCE3AE4B1E4383FFB81FE99" box="[729,1072,277,302]" gridcol="2" gridrow="1" pageId="4" pageNumber="3">♂ subadult, holotype</td>
|
||
<td id="76812901FFCE3AE4B764383FFA01FE99" box="[1113,1456,277,302]" gridcol="3" gridrow="1" pageId="4" pageNumber="3">♀ juvenile, paratype</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413810FA01FEE2" box="[124,1456,314,341]" gridrow="2" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413810FEF7FEE2" box="[124,326,314,341]" gridcol="0" gridrow="2" pageId="4" pageNumber="3">Carapace</th>
|
||
<td id="76812901FFCE3AE4B1113810FD20FEE2" box="[556,657,314,341]" gridcol="1" gridrow="2" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76812901FFCE3AE4B1E43810FB81FEE2" box="[729,1072,314,341]" gridcol="2" gridrow="2" pageId="4" pageNumber="3">2.41 / 3.03</td>
|
||
<td id="76812901FFCE3AE4B7643810FA01FEE2" box="[1113,1456,314,341]" gridcol="3" gridrow="2" pageId="4" pageNumber="3">2.46 / 3.18</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B341384BFA01FECC" box="[124,1456,353,379]" gridrow="3" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B341384BFEF7FECC" box="[124,326,353,379]" gridcol="0" gridrow="3" pageId="4" pageNumber="3">Mesosoma</th>
|
||
<td id="76812901FFCE3AE4B111384BFD20FECC" box="[556,657,353,379]" gridcol="1" gridrow="3" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E4384BFB81FECC" box="[729,1072,353,379]" gridcol="2" gridrow="3" pageId="4" pageNumber="3">7.50</td>
|
||
<td id="76812901FFCE3AE4B764384BFA01FECC" box="[1113,1456,353,379]" gridcol="3" gridrow="3" pageId="4" pageNumber="3">7.56</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B34138A2FA01FE15" box="[124,1456,392,418]" gridrow="4" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B34138A2FEF7FE15" box="[124,326,392,418]" gridcol="0" gridrow="4" pageId="4" pageNumber="3">Tergite VII</th>
|
||
<td id="76812901FFCE3AE4B11138A2FD20FE15" box="[556,657,392,418]" gridcol="1" gridrow="4" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76812901FFCE3AE4B1E438A2FB81FE15" box="[729,1072,392,418]" gridcol="2" gridrow="4" pageId="4" pageNumber="3">1.64 / 3.00</td>
|
||
<td id="76812901FFCE3AE4B76438A2FA01FE15" box="[1113,1456,392,418]" gridcol="3" gridrow="4" pageId="4" pageNumber="3">1.57 / 3.04</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413885FA01FE7E" box="[124,1456,431,457]" gridrow="5" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413885FEF7FE7E" box="[124,326,431,457]" gridcol="0" gridrow="5" pageId="4" pageNumber="3">Metasoma + telson</th>
|
||
<td id="76812901FFCE3AE4B1113885FD20FE7E" box="[556,657,431,457]" gridcol="1" gridrow="5" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E43885FB81FE7E" box="[729,1072,431,457]" gridcol="2" gridrow="5" pageId="4" pageNumber="3">12.05</td>
|
||
<td id="76812901FFCE3AE4B7643885FA01FE7E" box="[1113,1456,431,457]" gridcol="3" gridrow="5" pageId="4" pageNumber="3">12.30</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B34138FFFA01FE58" box="[124,1456,469,495]" gridrow="6" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B34138FFFEF7FE58" box="[124,326,469,495]" gridcol="0" gridrow="6" pageId="4" pageNumber="3">Segment I</th>
|
||
<td id="76812901FFCE3AE4B11138FFFD20FE58" box="[556,657,469,495]" gridcol="1" gridrow="6" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E438FFFB81FE58" box="[729,1072,469,495]" gridcol="2" gridrow="6" pageId="4" pageNumber="3">1.45 / 1.75 / 1.43</td>
|
||
<td id="76812901FFCE3AE4B76438FFFA01FE58" box="[1113,1456,469,495]" gridcol="3" gridrow="6" pageId="4" pageNumber="3">1.47 / 1.77 / 1.49</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B34138D6FA01FDA1" box="[124,1456,508,534]" gridrow="7" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B34138D6FEF7FDA1" box="[124,326,508,534]" gridcol="0" gridrow="7" pageId="4" pageNumber="3">Segment II</th>
|
||
<td id="76812901FFCE3AE4B11138D6FD20FDA1" box="[556,657,508,534]" gridcol="1" gridrow="7" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E438D6FB81FDA1" box="[729,1072,508,534]" gridcol="2" gridrow="7" pageId="4" pageNumber="3">1.68 / 1.70 / 1.50</td>
|
||
<td id="76812901FFCE3AE4B76438D6FA01FDA1" box="[1113,1456,508,534]" gridcol="3" gridrow="7" pageId="4" pageNumber="3">1.73 / 1.73 / 1.50</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413B09FA01FD8A" box="[124,1456,547,573]" gridrow="8" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413B09FEF7FD8A" box="[124,326,547,573]" gridcol="0" gridrow="8" pageId="4" pageNumber="3">Segment III</th>
|
||
<td id="76812901FFCE3AE4B1113B09FD20FD8A" box="[556,657,547,573]" gridcol="1" gridrow="8" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E43B09FB81FD8A" box="[729,1072,547,573]" gridcol="2" gridrow="8" pageId="4" pageNumber="3">1.80 / 1.74 / 1.53</td>
|
||
<td id="76812901FFCE3AE4B7643B09FA01FD8A" box="[1113,1456,547,573]" gridcol="3" gridrow="8" pageId="4" pageNumber="3">1.88 / 1.74 / 1.55</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413B63FA01FDD3" box="[124,1456,585,612]" gridrow="9" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413B63FEF7FDD3" box="[124,326,585,612]" gridcol="0" gridrow="9" pageId="4" pageNumber="3">Segment IV</th>
|
||
<td id="76812901FFCE3AE4B1113B63FD20FDD3" box="[556,657,585,612]" gridcol="1" gridrow="9" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E43B63FB81FDD3" box="[729,1072,585,612]" gridcol="2" gridrow="9" pageId="4" pageNumber="3">2.23 / 1.73 / 1.42</td>
|
||
<td id="76812901FFCE3AE4B7643B63FA01FDD3" box="[1113,1456,585,612]" gridcol="3" gridrow="9" pageId="4" pageNumber="3">2.25 / 1.75 / 1.48</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413B5AFA01FD3D" box="[124,1456,624,650]" gridrow="10" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413B5AFEF7FD3D" box="[124,326,624,650]" gridcol="0" gridrow="10" pageId="4" pageNumber="3">Segment V</th>
|
||
<td id="76812901FFCE3AE4B1113B5AFD20FD3D" box="[556,657,624,650]" gridcol="1" gridrow="10" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E43B5AFB81FD3D" box="[729,1072,624,650]" gridcol="2" gridrow="10" pageId="4" pageNumber="3">2.51 / 1.67 / 1.15</td>
|
||
<td id="76812901FFCE3AE4B7643B5AFA01FD3D" box="[1113,1456,624,650]" gridcol="3" gridrow="10" pageId="4" pageNumber="3">2.60 / 1.68 / 1.20</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413BBDFA01FD06" box="[124,1456,663,689]" gridrow="11" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413BBDFEF7FD06" box="[124,326,663,689]" gridcol="0" gridrow="11" pageId="4" pageNumber="3">Telson</th>
|
||
<td id="76812901FFCE3AE4B1113BBDFD20FD06" box="[556,657,663,689]" gridcol="1" gridrow="11" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76812901FFCE3AE4B1E43BBDFB81FD06" box="[729,1072,663,689]" gridcol="2" gridrow="11" pageId="4" pageNumber="3">2.38 / 0.96 / 0.73</td>
|
||
<td id="76812901FFCE3AE4B7643BBDFA01FD06" box="[1113,1456,663,689]" gridcol="3" gridrow="11" pageId="4" pageNumber="3">2.37 / 0.97 / 0.81</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413B94FA01FD6F" box="[124,1456,702,728]" gridrow="12" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413B94FEF7FD6F" box="[124,326,702,728]" gridcol="0" gridrow="12" pageId="4" pageNumber="3">Pedipalp</th>
|
||
<td id="76812901FFCE3AE4B1113B94FD20FD6F" box="[556,657,702,728]" gridcol="1" gridrow="12" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E43B94FB81FD6F" box="[729,1072,702,728]" gridcol="2" gridrow="12" pageId="4" pageNumber="3">6.60</td>
|
||
<td id="76812901FFCE3AE4B7643B94FA01FD6F" box="[1113,1456,702,728]" gridcol="3" gridrow="12" pageId="4" pageNumber="3">6.69</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413BCEFA01FD49" box="[124,1456,740,766]" gridrow="13" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413BCEFEF7FD49" box="[124,326,740,766]" gridcol="0" gridrow="13" pageId="4" pageNumber="3">Femur</th>
|
||
<td id="76812901FFCE3AE4B1113BCEFD20FD49" box="[556,657,740,766]" gridcol="1" gridrow="13" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76812901FFCE3AE4B1E43BCEFB81FD49" box="[729,1072,740,766]" gridcol="2" gridrow="13" pageId="4" pageNumber="3">1.67 / 0.58</td>
|
||
<td id="76812901FFCE3AE4B7643BCEFA01FD49" box="[1113,1456,740,766]" gridcol="3" gridrow="13" pageId="4" pageNumber="3">1.74 / 0.62</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413A21FA01FC92" box="[124,1456,779,805]" gridrow="14" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413A21FEF7FC92" box="[124,326,779,805]" gridcol="0" gridrow="14" pageId="4" pageNumber="3">Patella</th>
|
||
<td id="76812901FFCE3AE4B1113A21FD20FC92" box="[556,657,779,805]" gridcol="1" gridrow="14" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76812901FFCE3AE4B1E43A21FB81FC92" box="[729,1072,779,805]" gridcol="2" gridrow="14" pageId="4" pageNumber="3">2.10 / 0.78</td>
|
||
<td id="76812901FFCE3AE4B7643A21FA01FC92" box="[1113,1456,779,805]" gridcol="3" gridrow="14" pageId="4" pageNumber="3">2.08 / 0.78</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413A18FA01FCFB" box="[124,1456,818,844]" gridrow="15" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413A18FEF7FCFB" box="[124,326,818,844]" gridcol="0" gridrow="15" pageId="4" pageNumber="3">Chela</th>
|
||
<td id="76812901FFCE3AE4B1113A18FD20FCFB" box="[556,657,818,844]" gridcol="1" gridrow="15" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E43A18FB81FCFB" box="[729,1072,818,844]" gridcol="2" gridrow="15" pageId="4" pageNumber="3">2.83</td>
|
||
<td id="76812901FFCE3AE4B7643A18FA01FCFB" box="[1113,1456,818,844]" gridcol="3" gridrow="15" pageId="4" pageNumber="3">2.87</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413A72FA01FCC4" box="[124,1456,856,883]" gridrow="16" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413A72FEF7FCC4" box="[124,326,856,883]" gridcol="0" gridrow="16" pageId="4" pageNumber="3">Manus</th>
|
||
<td id="76812901FFCE3AE4B1113A72FD20FCC4" box="[556,657,856,883]" gridcol="1" gridrow="16" pageId="4" pageNumber="3">W / D</td>
|
||
<td id="76812901FFCE3AE4B1E43A72FB81FCC4" box="[729,1072,856,883]" gridcol="2" gridrow="16" pageId="4" pageNumber="3">0.56 / 0.54</td>
|
||
<td id="76812901FFCE3AE4B7643A72FA01FCC4" box="[1113,1456,856,883]" gridcol="3" gridrow="16" pageId="4" pageNumber="3">0.60 / 0.57</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413A55FA01FC2E" box="[124,1456,895,921]" gridrow="17" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413A55FEF7FC2E" box="[124,326,895,921]" gridcol="0" gridrow="17" pageId="4" pageNumber="3">Movable finger</th>
|
||
<td id="76812901FFCE3AE4B1113A55FD20FC2E" box="[556,657,895,921]" gridcol="1" gridrow="17" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E43A55FB81FC2E" box="[729,1072,895,921]" gridcol="2" gridrow="17" pageId="4" pageNumber="3">1.75</td>
|
||
<td id="76812901FFCE3AE4B7643A55FA01FC2E" box="[1113,1456,895,921]" gridcol="3" gridrow="17" pageId="4" pageNumber="3">1.85</td>
|
||
</tr>
|
||
<tr id="3550407DFFCE3AE4B3413A8FFA01FC77" box="[124,1456,933,960]" gridrow="18" pageId="4" pageNumber="3">
|
||
<th id="76812901FFCE3AE4B3413A8FFEF7FC77" box="[124,326,933,960]" gridcol="0" gridrow="18" pageId="4" pageNumber="3">Total</th>
|
||
<td id="76812901FFCE3AE4B1113A8FFD20FC77" box="[556,657,933,960]" gridcol="1" gridrow="18" pageId="4" pageNumber="3">L</td>
|
||
<td id="76812901FFCE3AE4B1E43A8FFB81FC77" box="[729,1072,933,960]" gridcol="2" gridrow="18" pageId="4" pageNumber="3">21.96</td>
|
||
<td id="76812901FFCE3AE4B7643A8FFA01FC77" box="[1113,1456,933,960]" gridcol="3" gridrow="18" pageId="4" pageNumber="3">22.32</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF1F12B7FFCEC51FB34E3AFBFE24FBB3" ID-Table-UUID="DF1F12B7FFCEC51FB34E3AFBFE24FBB3" httpUri="http://table.plazi.org/id/DF1F12B7FFCEC51FB34E3AFBFE24FBB3" pageId="4" pageNumber="3" startId="4.[115,171,977,998]" targetBox="[124,1456,237,960]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BDF423FFFCEC51FB34E3AFBFE24FBB3" blockId="4.[115,1473,977,1028]" pageId="4" pageNumber="3">
|
||
<emphasis id="B9149E2DFFCEC51FB34E3AFBFF0DFC51" bold="true" box="[115,188,977,998]" pageId="4" pageNumber="3">Table 1</emphasis>
|
||
. Comparative measurements of types of
|
||
<taxonomicName id="4C6039BCFFCEC51FB1033AFBFD56FC51" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[574,743,977,998]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="dentator" status="gen. et sp. n.">
|
||
<emphasis id="B9149E2DFFCEC51FB1033AFBFD56FC51" box="[574,743,977,998]" italics="true" pageId="4" pageNumber="3">Fetilinia dentator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB1D13AFBFCD7FC50" box="[748,870,977,999]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B9149E2DFFCEC51FB1D13AFBFCA1FC51" bold="true" box="[748,784,977,998]" pageId="4" pageNumber="3">gen</emphasis>
|
||
. et
|
||
<emphasis id="B9149E2DFFCEC51FB00C3AFBFCF9FC51" bold="true" box="[817,840,977,998]" pageId="4" pageNumber="3">sp</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB06F3AFBFCEEFC51" bold="true" box="[850,863,977,998]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDF423FFFCEC51FB34C3EABFA18F84E" blockId="4.[113,773,1921,2047]" lastBlockId="4.[816,1476,1057,2041]" pageId="4" pageNumber="3">
|
||
AFFINITIES.
|
||
<taxonomicName id="4C6039BCFFCEC51FB3D13EABFEF7F82E" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[236,326,1921,1945]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB3D13EABFEF7F82E" box="[236,326,1921,1945]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB2713EABFE24F82F" box="[332,405,1921,1945]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB2713EABFEC2F82E" bold="true" box="[332,371,1921,1945]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB2BD3EABFE3EF82E" bold="true" box="[384,399,1921,1945]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
belongs to the Palaearctic ‘
|
||
<taxonomicName id="4C6039BCFFCEC51FB1893EABFD4DF82E" authorityName="Leach" authorityYear="1815" box="[692,764,1921,1945]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB1893EABFD4DF82E" box="[692,764,1921,1945]" italics="true" pageId="4" pageNumber="3">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group of
|
||
<bibRefCitation id="EFF13FCEFFCEC51FB3EC3E8BFEDEF80E" author="FET" box="[209,367,1953,1977]" pageId="4" pageNumber="3" pagination="1 - 40" refId="ref4595" refString="FET, V., M. E. SOLEGLAD & G. LOWE. 2005. A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1 - 40." type="journal article" year="2005">Fet et al., 2005</bibRefCitation>
|
||
, according to the following characters: trichobothrial pattern typeA-β; patella trichobothrium
|
||
<emphasis id="B9149E2DFFCEC51FB1A63EEBFD19F86E" box="[667,680,1985,2009]" italics="true" pageId="4" pageNumber="3">d</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB1953EFBFD01F868" attach="left" box="[680,688,2001,2015]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
internal to dorsomedian carina, manus
|
||
<emphasis id="B9149E2DFFCEC51FB28F3ECBFE7EF84E" box="[434,463,2017,2041]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB2F23EDBFE66F848" attach="left" box="[463,471,2033,2047]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB2DB3ECBFDB2F84E" box="[486,515,2017,2041]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<subScript id="17E4407AFFCEC51FB13E3EDBFDBAF848" attach="left" box="[515,523,2033,2047]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
-
|
||
<emphasis id="B9149E2DFFCEC51FB1273ECBFD86F84E" box="[538,567,2017,2041]" italics="true" pageId="4" pageNumber="3">Eb</emphasis>
|
||
<quantity id="4C98EFDAFFCEC51FB10A3EDBFDE7F84E" box="[567,598,2017,2047]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="3" unit="in" value="3.0">
|
||
<subScript id="17E4407AFFCEC51FB10A3EDBFD8EF848" attach="left" box="[567,575,2033,2047]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
in
|
||
</quantity>
|
||
δ configuration; tibial spurs present on legs III–IV; pedipalp chela finger median denticle rows non-imbricated; and posterior margins of tergite VII & metasoma I–III bearing fringes of microsetae. Within this group, it is similar to another small monotypic genus,
|
||
<taxonomicName id="4C6039BCFFCEC51FB00D3D8BFBF8FB0E" authority="Vachon, 1974" authorityName="Vachon" authorityYear="1974" box="[816,1097,1185,1209]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB00D3D8BFC1CFB0E" box="[816,941,1185,1209]" italics="true" pageId="4" pageNumber="3">Kraepelinia</emphasis>
|
||
<bibRefCitation id="EFF13FCEFFCEC51FB0883D8BFBF8FB0E" author="VACHON" box="[949,1097,1185,1209]" pageId="4" pageNumber="3" pagination="857 - 958" refId="ref5029" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaXie en arachnologie. Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle (Paris), 3 e serie, 140, Zoologie 104: 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
, which also has: enlarged dentition on ventrolateral and ventromedian carinae of metasoma II–III; irregular, enlarged lobate dentition on ventrolateral carinae of metasoma V; and trichobothrium
|
||
<emphasis id="B9149E2DFFCEC51FB7A73C2BFB02FAAE" box="[1178,1203,1281,1305]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
|
||
located on distal manus. However,
|
||
<taxonomicName id="4C6039BCFFCEC51FB0A33C0BFBAAFA8E" authorityName="Vachon" authorityYear="1974" box="[926,1051,1313,1337]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0A33C0BFBAAFA8E" box="[926,1051,1313,1337]" italics="true" pageId="4" pageNumber="3">Kraepelinia</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C6039BCFFCEC51FB7953C0BFAB3FA8E" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1192,1282,1313,1337]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB7953C0BFAB3FA8E" box="[1192,1282,1313,1337]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB6343C0BFAE5FA8F" box="[1289,1364,1313,1337]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB6343C0BFA81FA8E" bold="true" box="[1289,1328,1313,1337]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB6023C0BFAFFFA8E" bold="true" box="[1343,1358,1313,1337]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in several other characters: thickened pedipalp fingers that are atypical for buthids; smooth carapace and tergites; metasoma V with large, lobate denticles on ventral surface; telson bulbous and granulate.
|
||
<taxonomicName id="4C6039BCFFCEC51FB00D3C8BFC3BFA0E" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[816,906,1441,1465]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB00D3C8BFC3BFA0E" box="[816,906,1441,1465]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB0A83C8BFC55FA0F" box="[917,996,1441,1465]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0A83C8BFC0DFA0E" bold="true" box="[917,956,1441,1465]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB0F33C8BFC6CFA0E" bold="true" box="[974,989,1441,1465]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is also similar to five other small ‘
|
||
<taxonomicName id="4C6039BCFFCEC51FB64E3C8BFA0AFA0E" authorityName="Leach" authorityYear="1815" box="[1395,1467,1441,1465]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB64E3C8BFA0AFA0E" box="[1395,1467,1441,1465]" italics="true" pageId="4" pageNumber="3">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group genera:
|
||
<taxonomicName id="4C6039BCFFCEC51FB0EE3CEBFB67FA6E" authority="Simon, 1882" authorityName="Simon" authorityYear="1882" box="[979,1238,1473,1497]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0EE3CEBFB8DFA6E" box="[979,1084,1473,1497]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
|
||
Simon, 1882
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB7DB3CEBFCD6FA4E" authority="Karsch, 1891" authorityName="Karsch" authorityYear="1891" class="Arachnida" family="Buthidae" genus="Orthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7DB3CEBFAD7FA6E" box="[1254,1382,1473,1497]" italics="true" pageId="4" pageNumber="3">Orthochirus</emphasis>
|
||
Karsch, 1891
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB04D3CCBFB08FA4E" authority="KovarIk, 1996" authorityName="KovarIk" authorityYear="1996" box="[880,1209,1505,1531]" class="Arachnida" family="Buthidae" genus="Baloorthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB04D3CCBFBAAFA4E" box="[880,1051,1505,1529]" italics="true" pageId="4" pageNumber="3">Baloorthochirus</emphasis>
|
||
KovařÍk, 1996
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB7FF3CCBFCD6F9AE" authority="KovarIk, 1998" authorityName="KovarIk" authorityYear="1998" class="Arachnida" family="Buthidae" genus="Orthochiroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7FF3CCBFAD0FA4E" box="[1218,1377,1505,1529]" italics="true" pageId="4" pageNumber="3">Orthochiroides</emphasis>
|
||
KovařÍk, 1998
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6039BCFFCEC51FB0953F2BFB05F9AE" authority="Lowe, 2018" authorityName="Lowe" authorityYear="2018" box="[936,1204,1537,1561]" class="Arachnida" family="Buthidae" genus="Xenobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0953F2BFB95F9AE" box="[936,1060,1537,1561]" italics="true" pageId="4" pageNumber="3">Xenobuthus</emphasis>
|
||
Lowe, 2018
|
||
</taxonomicName>
|
||
. These display a similar habitus, with a trapezoidal carapace and small, short pedipalps.
|
||
<taxonomicName id="4C6039BCFFCEC51FB00D3F6BFC01F9EE" authorityName="Karsch" authorityYear="1891" box="[816,944,1601,1625]" class="Arachnida" family="Buthidae" genus="Orthochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB00D3F6BFC01F9EE" box="[816,944,1601,1625]" italics="true" pageId="4" pageNumber="3">Orthochirus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6039BCFFCEC51FB0E33F6BFBCCF9EE" authorityName="KovarIk" authorityYear="1998" box="[990,1149,1601,1625]" class="Arachnida" family="Buthidae" genus="Orthochiroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0E33F6BFBCCF9EE" box="[990,1149,1601,1625]" italics="true" pageId="4" pageNumber="3">Orthochiroides</emphasis>
|
||
</taxonomicName>
|
||
have telson shapes quite similar to that of
|
||
<taxonomicName id="4C6039BCFFCEC51FB0AB3F4BFC41F9CE" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[918,1008,1633,1657]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB0AB3F4BFC41F9CE" box="[918,1008,1633,1657]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB0CB3F4BFB8EF9CF" box="[1014,1087,1633,1657]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB0CB3F4BFBACF9CE" bold="true" box="[1014,1053,1633,1657]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB7163F4BFB8BF9CE" bold="true" box="[1067,1082,1633,1657]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, but they differ in having metasomal segments IV–V dilated and punctate.
|
||
<taxonomicName id="4C6039BCFFCEC51FB7963FABFAA2F92E" authorityName="Simon" authorityYear="1882" box="[1195,1299,1665,1689]" class="Arachnida" family="Buthidae" genus="Butheolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7963FABFAA2F92E" box="[1195,1299,1665,1689]" italics="true" pageId="4" pageNumber="3">Butheolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB6203FABFA28F92E" authorityName="Lowe" authorityYear="2018" box="[1309,1433,1665,1689]" class="Arachnida" family="Buthidae" genus="Xenobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB6203FABFA28F92E" box="[1309,1433,1665,1689]" italics="true" pageId="4" pageNumber="3">Xenobuthus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6039BCFFCEC51FB00D3F8BFC7EF90E" authorityName="KovarIk" authorityYear="1998" box="[816,975,1697,1721]" class="Arachnida" family="Buthidae" genus="Orthochiroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB00D3F8BFC7EF90E" box="[816,975,1697,1721]" italics="true" pageId="4" pageNumber="3">Orthochiroides</emphasis>
|
||
</taxonomicName>
|
||
differ in having bulbous telsons.All five of these genera further differ in lacking enlarged or lobate metasomal dentition. It is possible that
|
||
<taxonomicName id="4C6039BCFFCEC51FB7723FCBFB18F94E" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1103,1193,1761,1785]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB7723FCBFB18F94E" box="[1103,1193,1761,1785]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB7933FCBFB49F94F" box="[1198,1272,1761,1785]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7933FCBFB64F94E" bold="true" box="[1198,1237,1761,1785]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB7DF3FCBFB40F94E" bold="true" box="[1250,1265,1761,1785]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is phylogenetically associated with this ‘orthochiroid’ compleX, but has evolved its own specialized metasomal structure. The pattern of enlarged metasomal dentition in
|
||
<taxonomicName id="4C6039BCFFCEC51FB7103E6BFB1BF8EE" authorityName="Vachon" authorityYear="1974" box="[1069,1194,1857,1881]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7103E6BFB1BF8EE" box="[1069,1194,1857,1881]" italics="true" pageId="4" pageNumber="3">Kraepelinia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6039BCFFCEC51FB7DC3E6BFA8AF8EE" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1249,1339,1857,1881]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCEC51FB7DC3E6BFA8AF8EE" box="[1249,1339,1857,1881]" italics="true" pageId="4" pageNumber="3">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCEC51FB67E3E6BFA21F8EF" box="[1347,1424,1857,1881]" pageId="4" pageNumber="3" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB67E3E6BFADBF8EE" bold="true" box="[1347,1386,1857,1881]" pageId="4" pageNumber="3">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCEC51FB6473E6BFA38F8EE" bold="true" box="[1402,1417,1857,1881]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
also occurs in several other ‘
|
||
<taxonomicName id="4C6039BCFFCEC51FB70B3E4BFBCFF8CE" authorityName="Leach" authorityYear="1815" box="[1078,1150,1889,1913]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB70B3E4BFBCFF8CE" box="[1078,1150,1889,1913]" italics="true" pageId="4" pageNumber="3">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group genera:
|
||
<taxonomicName id="4C6039BCFFCEC51FB6173E4BFCD6F82E" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB6173E4BFAC2F8CE" box="[1322,1395,1889,1913]" italics="true" pageId="4" pageNumber="3">Buthus</emphasis>
|
||
Leach, 1815
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB04E3EABFB35F82E" authority="Lowe, 2010" authorityName="Lowe" authorityYear="2010" box="[883,1156,1921,1945]" class="Arachnida" family="Buthidae" genus="Femtobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB04E3EABFC4DF82E" box="[883,1020,1921,1945]" italics="true" pageId="4" pageNumber="3">Femtobuthus</emphasis>
|
||
Lowe, 2010
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB7B23EABFA0FF82E" authority="Vachon, 1950" authorityName="Vachon" authorityYear="1950" box="[1167,1470,1921,1945]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB7B23EABFA92F82E" box="[1167,1315,1921,1945]" italics="true" pageId="4" pageNumber="3">Odontobuthus</emphasis>
|
||
Vachon, 1950
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCEC51FB00D3E8BFB57F80E" authority="Lourenco & Duhem, 2009" authorityName="Lourenco & Duhem" authorityYear="2009" box="[816,1254,1953,1977]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB00D3E8BFC05F80E" box="[816,948,1953,1977]" italics="true" pageId="4" pageNumber="3">Pantobuthus</emphasis>
|
||
Lourenço & Duhem, 2009
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6039BCFFCEC51FB61A3E8BFA9EF86E" authority="Lowe, KovarIk, Stockmann & Stahlavsky, 2019" authorityName="Lowe, KovarIk, Stockmann & Stahlavsky" authorityYear="2019" class="Arachnida" family="Buthidae" genus="Trypanothacus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCEC51FB61A3E8BFA73F80E" box="[1319,1474,1953,1977]" italics="true" pageId="4" pageNumber="3">Trypanothacus</emphasis>
|
||
Lowe, KovařÍk, Stockmann & Šťáhlavský, 2019
|
||
</taxonomicName>
|
||
, and seems to be an adaptation of burrowing scorpions (
|
||
<bibRefCitation id="EFF13FCEFFCEC51FB7DF3ECBFA2CF84E" author="LOWE" box="[1250,1437,2017,2041]" pageId="4" pageNumber="3" pagination="1 - 30" refId="ref4831" refString="LOWE, G., F. KOVARIK, M. STOCKMANN & F. STAHLAVSKY. 2019. Trypanothacus gen. n., a new genus of burrowing scorpion from the Arabian Peninsula (Scorpiones: Buthidae). Euscorpius, 277: 1 - 30." type="journal article" year="2019">Lowe et al., 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A11B4FFCFC51EB34C39CDFA0DFD28" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="8BDF423FFFCFC51EB34C39CDFA0DFD28" blockId="5.[113,772,231,2047]" lastBlockId="5.[816,1475,231,671]" pageId="5" pageNumber="4">
|
||
REMARKS. We based
|
||
<taxonomicName id="4C6039BCFFCFC51EB26C39CDFE1AFF48" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[337,427,231,255]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB26C39CDFE1AFF48" box="[337,427,231,255]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB28E39CDFE4EFF49" box="[435,511,231,255]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB28E39CDFE6BFF48" bold="true" box="[435,474,231,255]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB2D439CDFE49FF48" bold="true" box="[489,504,231,255]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
on two type specimens, an immature (subadult) male and a juvenile female. Immaturity of the male is evidenced by a laterally swollen mesosoma in a well fed individual, a condition typical of immatures of many other scorpions that we have reared. The characters supporting an affinity with the ‘orthochiroid’ compleX of small scorpions (most ca.
|
||
<quantity id="4C98EFDAFFCFC51EB3E7388DFEE1FE08" box="[218,336,423,447]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="5" pageNumber="4" unit="mm" value="32.5" valueMax="40.0" valueMin="25.0">25–40 mm</quantity>
|
||
), imply that the adult of
|
||
<taxonomicName id="4C6039BCFFCFC51EB16A388DFD00FE08" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[599,689,423,447]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB16A388DFD00FE08" box="[599,689,423,447]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB185388DFCB2FE09" box="[696,771,423,447]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB185388DFD6EFE08" bold="true" box="[696,735,423,447]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB1D3388DFD4CFE08" bold="true" box="[750,765,423,447]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is also likely to be small in size. The size, strongly granulated integument and pectinal tooth count of the male are consistent with a late instar subadult, one ecdysis before maturity. Even if it were an earlier instar, the characters of trapezoidal carapace and small, short pedipalps are also partially present in early juveniles of ‘orthochiroid’ compleX scorpions. However, caution is certainly advisable when diagnosing a new genus from immature specimens. We therefore considered the alternative interpretation, that the type specimens represent earlier instars of a larger scorpion belonging to an already eXisting genus in the ‘
|
||
<taxonomicName id="4C6039BCFFCFC51EB2623A2DFE16FCA8" authorityName="Leach" authorityYear="1815" box="[351,423,775,799]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2623A2DFE16FCA8" box="[351,423,775,799]" italics="true" pageId="5" pageNumber="4">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group. A conspicuously dentate metasoma is usually present in early instars of species that eXhibit this character in adults. This narrows the list of likely alternative taXa known from this geographic region to four genera:
|
||
<taxonomicName id="4C6039BCFFCFC51EB3F23AADFEFAFC28" authorityName="Vachon" authorityYear="1974" box="[207,331,903,927]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB3F23AADFEFAFC28" box="[207,331,903,927]" italics="true" pageId="5" pageNumber="4">Kraepelinia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCFC51EB25E3AADFE1AFC28" authorityName="Leach" authorityYear="1815" box="[355,427,903,927]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB25E3AADFE1AFC28" box="[355,427,903,927]" italics="true" pageId="5" pageNumber="4">Buthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCFC51EB2FE3AADFDF0FC28" authorityName="Vachon" authorityYear="1950" box="[451,577,903,927]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2FE3AADFDF0FC28" box="[451,577,903,927]" italics="true" pageId="5" pageNumber="4">Mesobuthus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C6039BCFFCFC51EB1443AADFD4CFC28" authorityName="Lourenco & Duhem" authorityYear="2009" box="[633,765,903,927]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB1443AADFD4CFC28" box="[633,765,903,927]" italics="true" pageId="5" pageNumber="4">Pantobuthus</emphasis>
|
||
</taxonomicName>
|
||
. Could
|
||
<taxonomicName id="4C6039BCFFCFC51EB3813A8DFEA7FC08" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[188,278,935,959]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB3813A8DFEA7FC08" box="[188,278,935,959]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB2223A8DFEDCFC09" box="[287,365,935,959]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2223A8DFEF7FC08" bold="true" box="[287,326,935,959]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB26A3A8DFED7FC08" bold="true" box="[343,358,935,959]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
be an early instar juvenile of one of those genera? The slender pedipalps and elongate telson vesicle of
|
||
<taxonomicName id="4C6039BCFFCFC51EB3B03ACDFF56FC48" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[141,231,999,1023]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB3B03ACDFF56FC48" box="[141,231,999,1023]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB3D03ACDFE87FC49" box="[237,310,999,1023]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB3D03ACDFEA5FC48" bold="true" box="[237,276,999,1023]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB21D3ACDFE9EFC48" bold="true" box="[288,303,999,1023]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
are quite different from the robust pedipalps and bulbous telson typical of those four genera. However, these structures can often be quite tenuous in early instars, and later develop to become robust in adults.
|
||
<taxonomicName id="4C6039BCFFCFC51EB2CB3D6DFD64FBE8" baseAuthorityName="Birula" baseAuthorityYear="1903" box="[502,725,1095,1119]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="palpator">
|
||
<emphasis id="B9149E2DFFCFC51EB2CB3D6DFD64FBE8" box="[502,725,1095,1119]" italics="true" pageId="5" pageNumber="4">Kraepelinia palpator</emphasis>
|
||
</taxonomicName>
|
||
is a relatively small scorpion, with adult carapace length ca.
|
||
<quantity id="4C98EFDAFFCFC51EB1F83D4DFCB2FBC9" box="[709,771,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="4" unit="mm" value="4.0">4 mm</quantity>
|
||
(
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB3443DADFF4EFB28" author="BIRULA" box="[121,255,1159,1183]" pageId="5" pageNumber="4" pagination="67 - 80" refId="ref4476" refString="BIRULA, A. A. 1903. Beitrage zur Kenntniss der Scorpionenfauna Ost-Persiens. (2 Beitrag). Bulletin de l'Academie Imperiale des Sciences de Saint. - Petersbourg, ser. 5, 19 (2): 67 - 80." type="journal article" year="1903">Birula, 1903</bibRefCitation>
|
||
; Lourenço & Leguin, 2010), which would predict an adult body length of ca.
|
||
<quantity id="4C98EFDAFFCFC51EB2B33D8DFE6AFB09" box="[398,475,1191,1215]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="4" unit="mm" value="35.0">35 mm</quantity>
|
||
. We studied a subadult male
|
||
<taxonomicName id="4C6039BCFFCFC51EB34C3DEDFF5FFB68" authorityName="Vachon" authorityYear="1974" box="[113,238,1223,1247]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB34C3DEDFF5FFB68" box="[113,238,1223,1247]" italics="true" pageId="5" pageNumber="4">Kraepelinia</emphasis>
|
||
</taxonomicName>
|
||
with carapace length
|
||
<quantity id="4C98EFDAFFCFC51EB2E73DEDFD9DFB69" box="[474,556,1223,1247]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="4" unit="mm" value="2.9">2.9 mm</quantity>
|
||
and body length
|
||
<quantity id="4C98EFDAFFCFC51EB1D43DEDFF2CFB49" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="4" unit="mm" value="24.0">24 mm</quantity>
|
||
, similar in size to the subadult male of
|
||
<taxonomicName id="4C6039BCFFCFC51EB16E3DCDFD1CFB48" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[595,685,1255,1279]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB16E3DCDFD1CFB48" box="[595,685,1255,1279]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB18B3DCDFCB2FB49" box="[694,771,1255,1279]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB18B3DCDFD6CFB48" bold="true" box="[694,733,1255,1279]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB1D33DCDFD4CFB48" bold="true" box="[750,765,1255,1279]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="135B5EBAFFCFC51EB3443C2DFEB0FAA8" box="[121,257,1287,1311]" captionStart-0="Figures 30–31" captionStart-1="Figures 32–35" captionStartId-0="9.[113,190,2010,2032]" captionStartId-1="10.[113,187,2007,2029]" captionTargetBox-0="[113,1474,282,1949]" captionTargetBox-1="[113,1474,227,2000]" captionTargetId-0="figure-34@9.[113,1474,282,1949]" captionTargetId-1="figure-17@10.[113,1474,224,2002]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figures 30–31. Fetilinia dentator gen. et sp. n., subadult male holotype in dorsal (30) and ventral (31) views, under UV light." captionText-1="Figures 32–35: Kraepelinia palpator (Birula, 1903), subadult male habitus in dorsal (32, 34) and ventral (33, 35) views, under white light (32–33) under UV light (34–35).Turkmenistan, Badghyz, Eroilanduz, 35°42'04"N 61°48'53"E, el. 348 m, 7.IV.2002,leg.A.Gromov, GLPC.Scale bar:4 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4648986" figureDoi-1="http://doi.org/10.5281/zenodo.4648988" httpUri-0="https://zenodo.org/record/4648986/files/figure.png" httpUri-1="https://zenodo.org/record/4648988/files/figure.png" pageId="5" pageNumber="4">Figs. 30–35</figureCitation>
|
||
). The thickened pedipalp fingers and bulbous telson that are unique diagnostic characters of
|
||
<taxonomicName id="4C6039BCFFCFC51EB1BB3C0DFCB2FA88" authorityName="Vachon" authorityYear="1974" box="[646,771,1319,1343]" class="Arachnida" family="Buthidae" genus="Kraepelinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB1BB3C0DFCB2FA88" box="[646,771,1319,1343]" italics="true" pageId="5" pageNumber="4">Kraepelinia</emphasis>
|
||
</taxonomicName>
|
||
are already strongly eXpressed in the subadult, showing that
|
||
<taxonomicName id="4C6039BCFFCFC51EB34C3C4DFF7AFAC8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[113,203,1383,1407]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB34C3C4DFF7AFAC8" box="[113,203,1383,1407]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB3ED3C4DFEA8FAC9" box="[208,281,1383,1407]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB3ED3C4DFF46FAC8" bold="true" box="[208,247,1383,1407]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB23E3C4DFEA3FAC8" bold="true" box="[259,274,1383,1407]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is not a juvenile of this genus. The other genera in question (
|
||
<taxonomicName id="4C6039BCFFCFC51EB23A3CADFEFEFA28" authorityName="Leach" authorityYear="1815" box="[263,335,1415,1439]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB23A3CADFEFEFA28" box="[263,335,1415,1439]" italics="true" pageId="5" pageNumber="4">Buthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCFC51EB25A3CADFE54FA28" authorityName="Vachon" authorityYear="1950" box="[359,485,1415,1439]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB25A3CADFE54FA28" box="[359,485,1415,1439]" italics="true" pageId="5" pageNumber="4">Mesobuthus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6039BCFFCFC51EB2C13CADFD31FA28" authorityName="Lourenco & Duhem" authorityYear="2009" box="[508,640,1415,1439]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2C13CADFD31FA28" box="[508,640,1415,1439]" italics="true" pageId="5" pageNumber="4">Pantobuthus</emphasis>
|
||
</taxonomicName>
|
||
) possess a subrectangular carapace, a shape that differs markedly from the strongly trapezoidal carapace of
|
||
<taxonomicName id="4C6039BCFFCFC51EB2803CEDFDA6FA68" authority="An" authorityName="Lowe & Kovařík" authorityYear="2020" box="[445,535,1479,1503]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB2803CEDFDA6FA68" box="[445,535,1479,1503]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB1263CEDFDD3FA69" box="[539,610,1479,1503]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB1263CEDFDF3FA68" bold="true" box="[539,578,1479,1503]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB1703CEDFDEDFA68" bold="true" box="[589,604,1479,1503]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
An ontogenetic change from trapezoidal in juveniles to subrectangular in adults is implausible, being the reverse of the predicted polarity of such transformations (i.e., from primitive to derived). Another argument against
|
||
<taxonomicName id="4C6039BCFFCFC51EB2163F6DFE34F9E8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[299,389,1607,1631]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB2163F6DFE34F9E8" box="[299,389,1607,1631]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB2B63F6DFE67F9E9" box="[395,470,1607,1631]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2B63F6DFE03F9E8" bold="true" box="[395,434,1607,1631]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB2FD3F6DFE7EF9E8" bold="true" box="[448,463,1607,1631]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
being a juvenile of the other genera stems from their differing trichobothriotaXy, which would violate the established rule that trichobothrial patterns are nearly always conserved ontogenetically (
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB1623F8DFD48F908" author="VACHON" box="[607,761,1703,1727]" pageId="5" pageNumber="4" pagination="857 - 958" refId="ref5029" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaXie en arachnologie. Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle (Paris), 3 e serie, 140, Zoologie 104: 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C6039BCFFCFC51EB3B33FEDFEA3F968" authorityName="Lourenco & Duhem" authorityYear="2009" box="[142,274,1735,1759]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB3B33FEDFEA3F968" box="[142,274,1735,1759]" italics="true" pageId="5" pageNumber="4">Pantobuthus</emphasis>
|
||
</taxonomicName>
|
||
, and most
|
||
<taxonomicName id="4C6039BCFFCFC51EB2B93FEDFE7CF968" authorityName="Leach" authorityYear="1815" box="[388,461,1735,1759]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2B93FEDFE7CF968" box="[388,461,1735,1759]" italics="true" pageId="5" pageNumber="4">Buthus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6039BCFFCFC51EB13D3FEDFDCFF968" authorityName="Vachon" authorityYear="1950" box="[512,638,1735,1759]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB13D3FEDFDCFF968" box="[512,638,1735,1759]" italics="true" pageId="5" pageNumber="4">Mesobuthus</emphasis>
|
||
</taxonomicName>
|
||
, fiXed finger trichobothrium
|
||
<emphasis id="B9149E2DFFCFC51EB2243FCDFE82F948" box="[281,307,1767,1791]" italics="true" pageId="5" pageNumber="4">db</emphasis>
|
||
is located distal to
|
||
<emphasis id="B9149E2DFFCFC51EB1283FC2FD99F948" box="[533,552,1768,1791]" italics="true" pageId="5" pageNumber="4">et</emphasis>
|
||
, whereas it is level with or proXimal to
|
||
<emphasis id="B9149E2DFFCFC51EB2743E2DFEECF8A8" box="[329,349,1799,1823]" italics="true" pageId="5" pageNumber="4">dt</emphasis>
|
||
in
|
||
<taxonomicName id="4C6039BCFFCFC51EB2BC3E2DFE6AF8A8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[385,475,1799,1823]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB2BC3E2DFE6AF8A8" box="[385,475,1799,1823]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB2DE3E2DFD9EF8A9" box="[483,559,1799,1823]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB2DE3E2DFDBBF8A8" bold="true" box="[483,522,1799,1823]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB1243E2DFD99F8A8" bold="true" box="[537,552,1799,1823]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Distal translocation of
|
||
<emphasis id="B9149E2DFFCFC51EB3AD3E0DFF1BF888" box="[144,170,1831,1855]" italics="true" pageId="5" pageNumber="4">db</emphasis>
|
||
is typically associated with stretching and elongation of chela fingers, a trend that opposes the hypothetical ontogenetic transformation of slender fingers in
|
||
<taxonomicName id="4C6039BCFFCFC51EB2CA3E4DFDE0F8C8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[503,593,1895,1919]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB2CA3E4DFDE0F8C8" box="[503,593,1895,1919]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB1663E4DFD19F8C9" box="[603,680,1895,1919]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB1663E4DFD33F8C8" bold="true" box="[603,642,1895,1919]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB1AF3E4DFD10F8C8" bold="true" box="[658,673,1895,1919]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
to short robust fingers in the other genera. Furthermore,
|
||
<taxonomicName id="4C6039BCFFCFC51EB1943EADFCB2F828" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[681,771,1927,1951]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB1943EADFCB2F828" box="[681,771,1927,1951]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB34C3E8DFF71F809" box="[113,192,1959,1983]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB34C3E8DFF29F808" bold="true" box="[113,152,1959,1983]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB3973E8DFF08F808" bold="true" box="[170,185,1959,1983]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is neobothriotaXic, with femur
|
||
<emphasis id="B9149E2DFFCFC51EB1233E8DFD9AF808" box="[542,555,1959,1983]" italics="true" pageId="5" pageNumber="4">d</emphasis>
|
||
<subScript id="17E4407AFFCFC51EB1163E9DFD82F872" attach="left" box="[555,563,1975,1989]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
absent, in contrast to the other genera which are orthobothriotaXic. In scorpions, there is only one documented precedent in which reductive neobothriotaXy in juveniles transforms into orthobothriotaXy in adults. In many, if not all buthids, femoral trichobothrium
|
||
<emphasis id="B9149E2DFFCFC51EB6A73822FA10FEA8" box="[1434,1441,264,287]" italics="true" pageId="5" pageNumber="4">i</emphasis>
|
||
<subScript id="17E4407AFFCFC51EB69C383DFA18FE92" attach="left" box="[1441,1449,279,293]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
is absent in the second instar (first nymph), and only appears in the third and later instars (
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB723386DFB62FEE8" author="ARMAS" box="[1054,1235,327,351]" pageId="5" pageNumber="4" pagination="1 - 27" refId="ref4445" refString="ARMAS, L. F. de. 1986. BiologIa y morfometrIa de Rhopalurus garridoi Armas (Scorpiones: Buthidae). Poeyana, 333: 1 - 27." type="journal article" year="1986">Armas, 1986: 22</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB7E2386DFA0FFEE8" author="LOURENCO" box="[1247,1470,327,351]" pageId="5" pageNumber="4" pagination="95 - 117" refId="ref4725" refString="LOURENCO, W. R., 1979. Le scorpion Buthidae: Tityus mattogrossensis Borelli, 1901 (Morphologie, ecologie, biologie et developpement postembryonnaire). Bulletin du Museum National d'Histoire naturelle de Paris, 4 eme serie, 1 A, 1: 95 - 117." type="journal article" year="1979">Lourenço, 1979: 100</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB00D384DFBAEFEC8" author="STOCKMANN" box="[816,1055,359,383]" pageId="5" pageNumber="4" pagination="405 - 420" refId="ref4984" refString="STOCKMANN, R. 1979. Developpement postembryonnaire et cycle d'intermue chez un scorpion Buthidae: Buthotus minax occidentalis (Vachon & Stockmann). Bulletin du Museum National d'Histoire Naturelle, Paris, 1, A (2): 405 - 420." type="journal article" year="1979">Stockmann, 1979: 408</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB717384DFB40FEC8" author="VACHON" box="[1066,1265,359,383]" pageId="5" pageNumber="4" pagination="857 - 958" refId="ref5029" refString="VACHON, M. 1974. Etudes des caracteres utilises pour classer les familles et les genres des scorpions (Arachnides). 1. La trichobothriotaXie en arachnologie. Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle (Paris), 3 e serie, 140, Zoologie 104: 857 - 958." type="journal article" year="1974">Vachon, 1974: 873</bibRefCitation>
|
||
; R. Teruel, personal communication). However, similar abrupt early developmental changes have not been reported for buthid petite trichobothria including femur
|
||
<emphasis id="B9149E2DFFCFC51EB0D438EDFC47FE68" box="[1001,1014,455,479]" italics="true" pageId="5" pageNumber="4">d</emphasis>
|
||
<subScript id="17E4407AFFCFC51EB0CB38FDFC4FFE52" attach="left" box="[1014,1022,471,485]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
, whose presence or absence is normally consistent across instars. The size and well developed carination and granulation of the
|
||
<typeStatus id="54DBFC9DFFCFC51EB7213B2DFBC6FDA8" box="[1052,1143,519,543]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
male
|
||
<taxonomicName id="4C6039BCFFCFC51EB78B3B2DFAA1FDA8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1206,1296,519,543]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB78B3B2DFAA1FDA8" box="[1206,1296,519,543]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9149E2DFFCFC51EB62B3B2DFAEEFDA8" bold="true" box="[1302,1375,519,543]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB62B3B2DFAEEFDA8" box="[1302,1375,519,543]" pageId="5" pageNumber="4" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
indicates that it is not a second instar nymph, and it also has femur
|
||
<emphasis id="B9149E2DFFCFC51EB68E3B02FA0BFD88" box="[1459,1466,552,575]" italics="true" pageId="5" pageNumber="4">i</emphasis>
|
||
<subScript id="17E4407AFFCFC51EB6873B1DFA73FDF2" attach="left" box="[1466,1474,567,581]" fontSize="6" pageId="5" pageNumber="4">2</subScript>
|
||
already eXpressed (
|
||
<figureCitation id="135B5EBAFFCFC51EB73C3B6DFBE2FDE8" box="[1025,1107,583,607]" captionStart="Figures 10–23" captionStartId="7.[113,190,1916,1938]" captionTargetBox="[113,1454,234,1891]" captionTargetId="figure-99@7.[113,1474,234,1895]" captionTargetPageId="7" captionText="Figures 10–23: Fetilinia dentator gen. et sp. n., subadult male holotype. Figures 10–12. Carapace and tergites (10) and sternopectinal area and sternites (11–12) Figures 13–23. Pedipalp segments, chela dorsal (14), eXternal (15), ventral (16) views, patella dorsal (17), dorsoeXternal (18), and ventral (19) views, femur and trochanter dorsoeXternal (20), internal (21), and ventral (22) views, movable (13) and fiXed (23) finger dentition. Trichobothrial pattern is indicated by white circles in Figures 15–18 and 20–21. Figures 10–11 under UV light." figureDoi="http://doi.org/10.5281/zenodo.4648982" httpUri="https://zenodo.org/record/4648982/files/figure.png" pageId="5" pageNumber="4">Fig. 21</figureCitation>
|
||
). In summary, available evidence supports our hypothesis that
|
||
<taxonomicName id="4C6039BCFFCFC51EB7553B4DFB73FDC8" authority="Lowe & Kovařík, 2020" authorityName="Lowe & Kovařík" authorityYear="2020" box="[1128,1218,615,639]" class="Arachnida" family="Buthidae" genus="Fetilinia" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9149E2DFFCFC51EB7553B4DFB73FDC8" box="[1128,1218,615,639]" italics="true" pageId="5" pageNumber="4">Fetilinia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2272356FFCFC51EB7F73B4DFAA6FDC9" box="[1226,1303,615,639]" pageId="5" pageNumber="4" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB7F73B4DFB40FDC8" bold="true" box="[1226,1265,615,639]" pageId="5" pageNumber="4">gen</emphasis>
|
||
.
|
||
<emphasis id="B9149E2DFFCFC51EB63F3B4DFAA0FDC8" bold="true" box="[1282,1297,615,639]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is a new genus that does not fit into any of the eXisting ‘
|
||
<taxonomicName id="4C6039BCFFCFC51EB7EB3BADFAAFFD28" authorityName="Leach" authorityYear="1815" box="[1238,1310,647,671]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9149E2DFFCFC51EB7EB3BADFAAFFD28" box="[1238,1310,647,671]" italics="true" pageId="5" pageNumber="4">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A11B4FFCFC51EB00D3BEDFADCFC28" pageId="5" pageNumber="4" type="distribution">
|
||
<paragraph id="8BDF423FFFCFC51EB00D3BEDFADCFC28" blockId="5.[816,1474,711,927]" pageId="5" pageNumber="4">
|
||
DISTRIBUTION. Known only from the
|
||
<typeStatus id="54DBFC9DFFCFC51EB7FD3BEDFB5FFD68" box="[1216,1262,711,735]" pageId="5" pageNumber="4">type</typeStatus>
|
||
locality in northern
|
||
<collectingCountry id="F37702AFFFCFC51EB00D3BCDFC3FFD48" box="[816,910,743,767]" name="Pakistan" pageId="5" pageNumber="4">Pakistan</collectingCountry>
|
||
. The collection site lies at the edge of the Kohat Plateau, northeast of the Bannu Basin, an intermontane depression in the tectonically active orogenic belt at the western end of the Himalayas. The local substrate consists of Quaternary alluvial fan deposits produced by erosion of surrounding high mountain ranges (
|
||
<bibRefCitation id="EFF13FCEFFCFC51EB7903AADFAD0FC28" author="ABIR, I. A." box="[1197,1377,903,927]" pageId="5" pageNumber="4" pagination="242 - 258" refId="ref4391" refString="ABIR, I. A., S. D. KHAN, G. M. AZIZ & S. TARIQ. 2017. Bannu Basin, fold-and-thrust belt of northern Pakistan: Subsurface imaging and its implications for hydrocarbon eXploration. Marine and Petroleum Geology, 85: 242 - 258." type="journal article" year="2017">Abir et al., 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |