444 lines
50 KiB
XML
444 lines
50 KiB
XML
<document id="E1D46D0D23CC2218DBF23A9A14E3BA6B" ID-DOI="10.11646/zootaxa.3683.2.2" ID-GBIF-Dataset="edc5aac3-f1f9-4ff7-b583-616556bf1407" ID-ISSN="1175-5326" ID-Zenodo-Dep="223681" ID-ZooBank="B393CCFE-94EE-438F-958D-3712BA3D4C18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460644097812" checkinUser="plazi" docAuthor="Lobo, Fernando, Laspiur, Alejandro & Acosta, Juan Carlos" docDate="2013" docId="03C97438FFDC6A7A0CEBFDE5A7D1DFF6" docLanguage="en" docName="zt03683p132.pdf" docOrigin="Zootaxa 3683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phymaturus aguanegra Lobo, Laspiur & Acosta, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="122" masterDocId="FFF00C40FFDF6A7F0C7CFF8CA53AD972" masterDocTitle="Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina" masterLastPageNumber="132" masterPageNumber="117" pageNumber="120" updateTime="1698336511002" updateUser="plazi">
|
||
<mods:mods id="CAAB70956BF78D061A9AAD3615D3415F" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="63029362C3BA74BCFC775C470C6CAA0B">
|
||
<mods:title id="6AD54B0B5468111D73B564BDF1B7DF06">Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="964EDBEF861FC52731E482A6C57D7538" type="personal">
|
||
<mods:role id="56769B35D6FFABA6D79321574CCB3BA3">
|
||
<mods:roleTerm id="4C93119319D11B5C1135011E3807E146">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="86C23CB7CD95978C0E0A9D8B74B536C1">Lobo, Fernando</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CEABD042F36697170D86302DBD03B5E5" type="personal">
|
||
<mods:role id="D470DF484B5AEE8C14F7F813EE5D3FC8">
|
||
<mods:roleTerm id="6924A7B78C5113CC9C5B8266CF92D012">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4D715142FDC6E41F26AB0CE2B6603A87">Laspiur, Alejandro</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="52425181D6300FE74705BF1BEB64E616" type="personal">
|
||
<mods:role id="AA81CF7DE377D148897E63B3C2BBDEE6">
|
||
<mods:roleTerm id="5658D0DD4E851D6C13917F0E1EE0872A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="817F335EBE09124995621D187A9BA4BA">Acosta, Juan Carlos</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8771754FB7E3FCFE52537425A5F187B7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="11AD183BE3DCF7399CA0871219788CBA" type="host">
|
||
<mods:titleInfo id="6D50F902BD95C4421D03451E22904B48">
|
||
<mods:title id="C6AF50F956D7487FA496422A324B57A1">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="72C033EA06B2B0B31469065148069D13">
|
||
<mods:date id="CF6F46F1451120321B93200FC80FFC68">2013</mods:date>
|
||
<mods:detail id="8D3E1CAA7DAB87D029A398670DD30829" type="volume">
|
||
<mods:number id="65C81B1758801959F8C524B5B0636EAF">3683</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5BF4103FD149A5B27FFC7626BF5AE054" type="issue">
|
||
<mods:number id="A40FA2687A891616F33D52FF356F0640">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="3B05996E74201B02B45ABA327D376C89" unit="page">
|
||
<mods:start id="056C969BF0951DB1BD002DE42BEF18BF">117</mods:start>
|
||
<mods:end id="083130F99F787B5A173A65DF0547E40B">132</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="EE029415F68E13826F361720F282967A">journal article</mods:classification>
|
||
<mods:identifier id="95CDAF921EA63D05A64831F8D6712F2A" type="DOI">10.11646/zootaxa.3683.2.2</mods:identifier>
|
||
<mods:identifier id="D7F5BE203C5447C99463F4640F1CBB5B" type="GBIF-Dataset">edc5aac3-f1f9-4ff7-b583-616556bf1407</mods:identifier>
|
||
<mods:identifier id="090D6A98E7DDCB23D283729D05129CAC" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="023D2269EB2823667DBEC40B5FBB2501" type="Zenodo-Dep">223681</mods:identifier>
|
||
<mods:identifier id="A0A23493C770AB335FFDD70F552AFCD5" type="ZooBank">B393CCFE-94EE-438F-958D-3712BA3D4C18</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C97438FFDC6A7A0CEBFDE5A7D1DFF6" ID-DOI="http://doi.org/10.5281/zenodo.5696280" ID-GBIF-Taxon="119571811" ID-Zenodo-Dep="5696280" LSID="urn:lsid:plazi:treatment:03C97438FFDC6A7A0CEBFDE5A7D1DFF6" httpUri="http://treatment.plazi.org/id/03C97438FFDC6A7A0CEBFDE5A7D1DFF6" lastPageId="5" lastPageNumber="122" pageId="3" pageNumber="120">
|
||
<subSubSection id="C37A96A5FFDC6A7C0CEBFDE5A720DBF1" box="[151,538,616,643]" pageId="3" pageNumber="120" type="nomenclature">
|
||
<paragraph id="8BDFC52EFFDC6A7C0CEBFDE5A720DBF1" blockId="3.[151,538,616,643]" box="[151,538,616,643]" pageId="3" pageNumber="120">
|
||
<heading id="D0977242FFDC6A7C0CEBFDE5A720DBF1" bold="true" box="[151,538,616,643]" fontSize="11" level="1" pageId="3" pageNumber="120" reason="1">
|
||
<emphasis id="B914193CFFDC6A7C0CEBFDE5A720DBF1" bold="true" box="[151,538,616,643]" pageId="3" pageNumber="120">
|
||
<taxonomicName id="4C60BEADFFDC6A7C0CEBFDE5A48EDBF1" box="[151,436,617,643]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="aguanegra" status="sp. nov.">
|
||
<emphasis id="B914193CFFDC6A7C0CEBFDE5A48EDBF1" bold="true" box="[151,436,617,643]" italics="true" pageId="3" pageNumber="120">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227A447FFDC6A7C0DC0FDE4A720DBF1" box="[444,538,616,643]" pageId="3" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFDC6A7C0CEBFD3CA6EEDA46" pageId="3" pageNumber="120" type="materials_examined">
|
||
<paragraph id="8BDFC52EFFDC6A7C0CEBFD3CA104DB9E" blockId="3.[151,1435,687,820]" pageId="3" pageNumber="120">
|
||
<typeStatus id="54DB7B8CFFDC6A7C0CEBFD3CA43EDBBA" box="[151,260,688,712]" pageId="3" pageNumber="120" type="holotype">
|
||
<emphasis id="B914193CFFDC6A7C0CEBFD3CA43EDBBA" bold="true" box="[151,260,688,712]" pageId="3" pageNumber="120">Holotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<figureCitation id="135BD9ABFFDC6A7C0D6EFD23A477DBB5" box="[274,333,687,711]" captionStart="FIGURE 1. A" captionStartId="3.[151,250,1817,1840]" captionTargetBox="[151,1436,873,1795]" captionTargetId="figure@3.[151,1436,873,1797]" captionTargetPageId="3" captionText="FIGURE 1. A. Dorsal view of the holotype specimen of Phymaturus aguanegra sp. nov. (MCN 975) B. Ventral view of the same specimen. SVL: 95.3 mm." httpUri="https://zenodo.org/record/223682/files/figure.png" pageId="3" pageNumber="120">Fig 1</figureCitation>
|
||
.): MCN 975. Male. Paso Agua Negra, Iglesia department, San Juan province,
|
||
<collectingCountry id="F37785BEFFDC6A7C08CBFD23A010DBB5" box="[1207,1322,687,711]" name="Argentina" pageId="3" pageNumber="120">Argentina</collectingCountry>
|
||
.
|
||
<geoCoordinate id="EE54A3E9FFDC6A7C0948FD23A0ADDBB5" box="[1332,1431,687,711]" direction="south" orientation="latitude" pageId="3" pageNumber="120" precision="925" value="-30.383333">30º23’ S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE54A3E9FFDC6A7C0CEBFD58A43EDB9E" box="[151,260,724,748]" direction="west" orientation="longitude" pageId="3" pageNumber="120" precision="925" value="-69.566666">69º34’ W</geoCoordinate>
|
||
.
|
||
<quantity id="4C9868CBFFDC6A7C0D72FD58A458DB9E" box="[270,354,724,748]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="120" unit="m" value="2900.0">2900 m</quantity>
|
||
asl.
|
||
<date id="FFDEE3EEFFDC6A7C0DEAFD58A72EDB9E" box="[406,532,724,748]" pageId="3" pageNumber="120" value="2007-04">April 2007</date>
|
||
. A. Laspiur, E. Sanabria & L. Quiroga collectors.
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFDC6A7C0CB5FD74A6EEDA46" blockId="3.[151,1435,687,820]" pageId="3" pageNumber="120">
|
||
<emphasis id="B914193CFFDC6A7C0CB5FD74A470DA62" bold="true" box="[201,330,760,784]" pageId="3" pageNumber="120">
|
||
<typeStatus id="54DB7B8CFFDC6A7C0CB5FD74A47FDA62" box="[201,325,760,784]" pageId="3" pageNumber="120" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
MCN (969, 973-975, 977, 979, 982, 984, 988, 990-991, 995 and 971-972, 976, 978, 980, 981, 983, 986-987, 989, 992-993).
|
||
<specimenCount id="9D660EA7FFDC6A7C0DD3FC90A728DA46" box="[431,530,796,820]" pageId="3" pageNumber="120" type="male">13 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D660EA7FFDC6A7C0E35FC90A7FDDA46" box="[585,711,796,820]" pageId="3" pageNumber="120" type="female">14 females</specimenCount>
|
||
. Same data as
|
||
<typeStatus id="54DB7B8CFFDC6A7C0F17FC90A6EBDA46" box="[875,977,796,820]" pageId="3" pageNumber="120" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1F95A6FFDC6A7C0CEBF895A4E6DE3D" httpUri="https://zenodo.org/record/223682/files/figure.png" pageId="3" pageNumber="120" targetBox="[151,1436,873,1795]" targetPageId="3">
|
||
<paragraph id="8BDFC52EFFDC6A7C0CEBF895A4E6DE3D" blockId="3.[151,1436,1817,1871]" pageId="3" pageNumber="120">
|
||
<emphasis id="B914193CFFDC6A7C0CEBF895A42EDE5D" bold="true" box="[151,276,1817,1840]" pageId="3" pageNumber="120">FIGURE 1.</emphasis>
|
||
A. Dorsal view of the holotype specimen of
|
||
<taxonomicName id="4C60BEADFFDC6A7C0E97F896A6E1DE42" box="[747,987,1818,1840]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="aguanegra" status="sp. nov.">
|
||
<emphasis id="B914193CFFDC6A7C0E97F896A6E1DE42" box="[747,987,1818,1840]" italics="true" pageId="3" pageNumber="120">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227A447FFDC6A7C0F9FF896A10FDE42" box="[995,1077,1818,1840]" pageId="3" pageNumber="120" rank="species">
|
||
<emphasis id="B914193CFFDC6A7C0F9FF896A10FDE42" bold="true" box="[995,1077,1818,1840]" pageId="3" pageNumber="120">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MCN 975) B.Ventral view of the same specimen. SVL: 95.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37A96A5FFDC6A7B0CBBF8F9A71BDDED" lastPageId="4" lastPageNumber="121" pageId="3" pageNumber="120" type="diagnosis">
|
||
<paragraph id="8BDFC52EFFDC6A7B0CBBF8F9A7F0D9A6" blockId="3.[151,1436,1909,2006]" lastBlockId="4.[151,1437,151,2012]" lastPageId="4" lastPageNumber="121" pageId="3" pageNumber="120">
|
||
<emphasis id="B914193CFFDC6A7C0CBBF8F9A47BDEFC" bold="true" box="[199,321,1909,1934]" pageId="3" pageNumber="120">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C60BEADFFDC6A7C0D3BF8FAA774DEFC" box="[327,590,1910,1934]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDC6A7C0D3BF8FAA774DEFC" box="[327,590,1910,1934]" italics="true" pageId="3" pageNumber="120">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C60BEADFFDC6A7C0E84F8FAA66EDEFC" box="[760,852,1910,1934]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="120" phylum="Chordata" rank="species" species="palluma">
|
||
<emphasis id="B914193CFFDC6A7C0E84F8FAA66EDEFC" box="[760,852,1910,1934]" italics="true" pageId="3" pageNumber="120">palluma</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<emphasis id="B914193CFFDC6A7C0FD7F8FBA6D3DEFC" box="[939,1001,1911,1934]" italics="true" pageId="3" pageNumber="120">sensu</emphasis>
|
||
<bibRefCitation id="EFF1B8DFFFDC6A7C0F8CF8FAA1A6DEFC" author="Etheridge" box="[1008,1180,1910,1934]" pageId="3" pageNumber="120" refString="Etheridge, R. E. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995</bibRefCitation>
|
||
) because it has square, non-imbricate superciliaries, scales of the tail markedly spinose, usually a fragmented subocular, and the subocular-supralabials are separated by two or more scale rows. This new taxon belongs to the Puna subclade of the
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CBCFF14A426D9C2" box="[192,284,152,176]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="palluma">
|
||
<emphasis id="B914193CFFDB6A7B0CBCFF14A426D9C2" box="[192,284,152,176]" italics="true" pageId="4" pageNumber="121">palluma</emphasis>
|
||
</taxonomicName>
|
||
group (Lobo & Quinteros 2005) because it shows the typical dorsal “spray” pattern (southern
|
||
<taxonomicName id="4C60BEADFFDB6A7B093CFF14A0A6D9C2" box="[1344,1436,152,176]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="palluma">
|
||
<emphasis id="B914193CFFDB6A7B093CFF14A0A6D9C2" box="[1344,1436,152,176]" italics="true" pageId="4" pageNumber="121">palluma</emphasis>
|
||
</taxonomicName>
|
||
group members show a dorsal reticulated pattern).
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFDB6A7B0CB5FF53A71BDDED" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="121">
|
||
Within the Puna subclade,
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E7DFF6CA631D98A" box="[513,779,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0E7DFF6CA631D98A" box="[513,779,224,248]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C60BEADFFDB6A7B0FD4FF6DA14FD98A" box="[936,1141,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0FD4FF6DA14FD98A" box="[936,1141,224,248]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFDB6A7B08FAFF6DA1D3D98A" box="[1158,1257,225,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B08FAFF6DA1D3D98A" box="[1158,1257,225,248]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B095AFF6DA0A6D98A" box="[1318,1436,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B095AFF6DA0A6D98A" box="[1318,1436,224,248]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
because it presents a complete melanism over the dorsum of neck, whereas the other three species exhibit a clear interruption in the mid dorsal line (
|
||
<figureCitation id="135BD9ABFFDB6A7B0E5FFEABA74AD84D" box="[547,624,295,319]" captionStart="FIGURE 2" captionStartId="5.[151,250,1114,1137]" captionTargetBox="[158,1430,198,1089]" captionTargetId="figure@5.[151,1436,193,1093]" captionTargetPageId="5" captionText="FIGURE 2. Patterns of coloration in life of Phymaturus aguanegra sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223683/files/figure.png" pageId="4" pageNumber="121">Figs. 2</figureCitation>
|
||
A and B).
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E9BFEA4A6D5D832" box="[743,1007,296,320]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0E9BFEA4A6D5D832" box="[743,1007,296,320]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C60BEADFFDB6A7B08F9FEA5A06AD832" box="[1157,1360,296,320]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B08F9FEA5A06AD832" box="[1157,1360,296,320]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B09FBFEA5A5E6D816" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B09FBFEA5A5E6D816" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
in the presence of enlarged scales at the base of tail in males, which are absent in these two species (see this character in Fig. 15B of Lobo
|
||
<emphasis id="B914193CFFDB6A7B0D9BFEFDA72DD8FA" box="[487,535,368,392]" italics="true" pageId="4" pageNumber="121">et al</emphasis>
|
||
. 2010a).
|
||
<taxonomicName id="4C60BEADFFDB6A7B0EFCFEFCA6BCD8FA" box="[640,902,368,392]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0EFCFEFCA6BCD8FA" box="[640,902,368,392]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
has strongly keeled tarsal scales, differing from
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CEBFE19A437D8DE" box="[151,269,404,428]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0CEBFE19A437D8DE" box="[151,269,404,428]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
(slightly keeled). Tail of males and females of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F4AFE18A17BD8DE" box="[822,1089,404,428]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0F4AFE18A17BD8DE" box="[822,1089,404,428]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
lacks a pattern, whereas in
|
||
<taxonomicName id="4C60BEADFFDB6A7B09FBFE19A47FD8A2" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B09FBFE19A47FD8A2" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFDB6A7B0D28FE35A48ED8A2" box="[340,436,441,464]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B0D28FE35A48ED8A2" box="[340,436,441,464]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DBFFE35A70DD8A2" box="[451,567,440,464]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0DBFFE35A70DD8A2" box="[451,567,440,464]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E09FE35A62CD8A2" box="[629,790,440,464]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0E09FE35A62CD8A2" box="[629,790,440,464]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
the tail of females is always ringed, and the tail of males of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CC9FE51A46FD886" box="[181,341,476,500]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0CC9FE51A46FD886" box="[181,341,476,500]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DF7FE51A4D1D886" box="[395,491,477,500]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B0DF7FE51A4D1D886" box="[395,491,477,500]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
is ringed (not as marked as in females); however, in males of
|
||
<taxonomicName id="4C60BEADFFDB6A7B08DFFE51A057D886" box="[1187,1389,476,500]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B08DFFE51A057D886" box="[1187,1389,476,500]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B914193CFFDB6A7B0CEBFD8DA43CDB6A" box="[151,262,512,536]" italics="true" pageId="4" pageNumber="121">P. l a u re n t i</emphasis>
|
||
the ringed pattern may be present or absent (see this character in Fig. 10C of
|
||
<bibRefCitation id="EFF1B8DFFFDB6A7B080DFE73A0AADB65" author="Lobo" box="[1137,1424,511,535]" pageId="4" pageNumber="121" refString="Lobo, F. & Quinteros, S. (2005 b) A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia, 45, 143 - 177. http: // dx. doi. org / 10.1590 / S 0031 - 10492005001300001" type="journal article" year="2005" yearSuffix="b">Lobo & Quinteros 2005b</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CEBFDA8A4A4DB4E" box="[151,414,548,572]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0CEBFDA8A4A4DB4E" box="[151,414,548,572]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
lacks enlarged scales on the anterior margin of the antehumeral fold and centre of chest, as in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CC8FDC5A444DB12" box="[180,382,584,608]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0CC8FDC5A444DB12" box="[180,382,584,608]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DC9FDC5A713DB12" box="[437,553,584,608]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0DC9FDC5A713DB12" box="[437,553,584,608]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135BD9ABFFDB6A7B0E44FDCBA749DB2D" box="[568,627,583,607]" captionStart="FIGURE 2" captionStartId="5.[151,250,1114,1137]" captionTargetBox="[158,1430,198,1089]" captionTargetId="figure@5.[151,1436,193,1093]" captionTargetPageId="5" captionText="FIGURE 2. Patterns of coloration in life of Phymaturus aguanegra sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223683/files/figure.png" pageId="4" pageNumber="121">Fig.2</figureCitation>
|
||
C and D). Females of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F08FDC5A130DB12" box="[884,1034,585,608]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0F08FDC5A130DB12" box="[884,1034,585,608]" italics="true" pageId="4" pageNumber="121">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
lack flank coloration, whereas in
|
||
<taxonomicName id="4C60BEADFFDB6A7B09FBFDC5A47FDBF6" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B09FBFDC5A47FDBF6" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
(yellow),
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DC6FDE1A760DBF6" box="[442,602,620,644]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0DC6FDE1A760DBF6" box="[442,602,620,644]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0EEFFDE1A63CDBF6" box="[659,774,620,644]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0EEFFDE1A63CDBF6" box="[659,774,620,644]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
(orange), coloration is conspicuous. A vertebral dark gray stripe is usually present on the dorsum of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E0CFD1CA642DBDA" box="[624,888,656,680]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0E0CFD1CA642DBDA" box="[624,888,656,680]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
, whereas in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0874FD1DA1E8DBDA" box="[1032,1234,656,680]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0874FD1DA1E8DBDA" box="[1032,1234,656,680]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0976FD1DA044DBDA" box="[1290,1406,656,680]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0976FD1DA044DBDA" box="[1290,1406,656,680]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
, it is always absent. In
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DEEFD39A4C3DBBE" box="[402,505,693,716]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B0DEEFD39A4C3DBBE" box="[402,505,693,716]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E47FD39A7DBDBBE" box="[571,737,692,716]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0E47FD39A7DBDBBE" box="[571,737,692,716]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
this vertebral stripe is light gray. Females of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0968FD38A42ADB82" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0968FD38A42ADB82" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
lack transversal white stripes on the dorsal pattern, as females of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F82FD55A1F3DB82" box="[1022,1225,728,752]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0F82FD55A1F3DB82" box="[1022,1225,728,752]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and some females of
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CC4FD71A414DA66" box="[184,302,764,788]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B0CC4FD71A414DA66" box="[184,302,764,788]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
(see this character in Fig. 10A of Lobo
|
||
<emphasis id="B914193CFFDB6A7B0F79FD71A603DA66" box="[773,825,764,788]" italics="true" pageId="4" pageNumber="121">et al</emphasis>
|
||
. 2010a). Females and, to a lesser extent, males of
|
||
<taxonomicName id="4C60BEADFFDB6A7B09FBFD71A42BDA4A" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B09FBFD71A42BDA4A" italics="true" pageId="4" pageNumber="121">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
exhibit a tricolor dorsal pattern, with two shades of brown and scattered ferriferous oxide spots; this pattern is not found in any other species of the group, except the species we describe below and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0896FCC9A0AEDA2E" box="[1258,1428,836,860]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFDB6A7B0896FCC9A0AEDA2E" box="[1258,1428,836,860]" italics="true" pageId="4" pageNumber="121">P. paihuanense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C60BEADFFDB6A7B0CEBFCE4A49BDAF2" box="[151,417,872,896]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0CEBFCE4A49BDAF2" box="[151,417,872,896]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
lacks a scapular spot, present in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F5EFCE5A6F9DAF2" box="[802,963,872,896]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0F5EFCE5A6F9DAF2" box="[802,963,872,896]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F83FCE5A1BCDAF2" box="[1023,1158,872,896]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="extrilidus">
|
||
<emphasis id="B914193CFFDB6A7B0F83FCE5A1BCDAF2" box="[1023,1158,872,896]" italics="true" pageId="4" pageNumber="121">P. extrilidus</emphasis>
|
||
</taxonomicName>
|
||
. Rostral scale is always undivided in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0D4BFC00A77FDAD6" box="[311,581,908,932]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0D4BFC00A77FDAD6" box="[311,581,908,932]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
, whereas it can be divided in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0FBDFC01A1ABDAD6" box="[961,1169,908,932]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0FBDFC01A1ABDAD6" box="[961,1169,908,932]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFDB6A7B08A9FC01A001DAD6" box="[1237,1339,909,932]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B08A9FC01A001DAD6" box="[1237,1339,909,932]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
. Sexual dichromatism is more pronounced in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0E2BFC3DA786DABA" box="[599,700,945,968]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFDB6A7B0E2BFC3DA786DABA" box="[599,700,945,968]" italics="true" pageId="4" pageNumber="121">P. punae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFDB6A7B0EACFC3DA69ADABA" box="[720,928,944,968]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFDB6A7B0EACFC3DA69ADABA" box="[720,928,944,968]" italics="true" pageId="4" pageNumber="121">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFDB6A7B0FC8FC3DA163DABA" box="[948,1113,944,968]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFDB6A7B0FC8FC3DA163DABA" box="[948,1113,944,968]" italics="true" pageId="4" pageNumber="121">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C60BEADFFDB6A7B08DEFC3DA020DABA" box="[1186,1306,944,968]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFDB6A7B08DEFC3DA020DABA" box="[1186,1306,944,968]" italics="true" pageId="4" pageNumber="121">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C60BEADFFDB6A7B09FBFC3DA42ADA9E" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B09FBFC3DA42ADA9E" italics="true" pageId="4" pageNumber="121">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
, in which the dorsal pattern of males is slightly yellow or inconspicuous.
|
||
<taxonomicName id="4C60BEADFFDB6A7B0839FC58A077DA9E" box="[1093,1357,980,1004]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFDB6A7B0839FC58A077DA9E" box="[1093,1357,980,1004]" italics="true" pageId="4" pageNumber="121">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
differs from the recently described Chilean species
|
||
<taxonomicName id="4C60BEADFFDB6A7B0EF8FC75A129DD7D" authority="Nunez et al., 2010" authorityName="Nunez et al." authorityYear="2010" box="[644,1043,1015,1040]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFDB6A7B0EF8FC75A616DD62" box="[644,812,1016,1040]" italics="true" pageId="4" pageNumber="121">P. paihuanense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF1B8DFFFDB6A7B0F46FC7BA131DD7D" author="Nunez" box="[826,1035,1015,1040]" pageId="4" pageNumber="121" refString="Nunez, H., Veloso, A., Espejo, P., Veloso, C., Cortes, A. & Araya, S. (2010) Nuevas especies de Phymaturus (grupo " palluma ") para la zona cordillerana central de Chile (Reptilia, Sauria, Liolaemidae). Boletin del Museo Nacional de Historia Natural, 59, 41 - 74." type="journal article" year="2010">
|
||
Núñez
|
||
<emphasis id="B914193CFFDB6A7B0FF0FC75A687DD62" box="[908,957,1016,1040]" italics="true" pageId="4" pageNumber="121">et al</emphasis>
|
||
., 2010
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the presence of dorsal melanism of head and neck (absent in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0D87FB91A791DD46" box="[507,683,1052,1076]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFDB6A7B0D87FB91A791DD46" box="[507,683,1052,1076]" italics="true" pageId="4" pageNumber="121">P. paihuanense</emphasis>
|
||
</taxonomicName>
|
||
), in having scales not enlarged on the anterior margin of the antehumeral fold (present in
|
||
<taxonomicName id="4C60BEADFFDB6A7B0DA3FBCDA7B0DD2A" box="[479,650,1088,1112]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFDB6A7B0DA3FBCDA7B0DD2A" box="[479,650,1088,1112]" italics="true" pageId="4" pageNumber="121">P. paihuanense</emphasis>
|
||
</taxonomicName>
|
||
) and differs from
|
||
<taxonomicName id="4C60BEADFFDB6A7B0F27FBCDA044DD25" authority="Troncoso-Palacios & Lobo 2012" authorityName="Troncoso-Palacios & Lobo" authorityYear="2012" box="[859,1406,1087,1112]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="damasense">
|
||
<emphasis id="B914193CFFDB6A7B0F27FBCDA6C8DD2A" box="[859,1010,1088,1112]" italics="true" pageId="4" pageNumber="121">P. damasense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF1B8DFFFDB6A7B087EFBB3A04DDD25" author="Troncoso-Palacios" box="[1026,1399,1087,1111]" pageId="4" pageNumber="121" refString="Troncoso-Palacios, J. & Lobo, F. (2012) A new species of Phymaturus (Iguania: Liolaemidae) of the palluma group from Central Chile. Cuadernos de Herpetologia, 26, 69 - 78." type="journal article" year="2012">Troncoso-Palacios & Lobo 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the presence of non-projected scales on the anterior border of auditory meatus (present in
|
||
<taxonomicName id="4C60BEADFFDB6A7B08FAFBE9A027DD0E" box="[1158,1309,1124,1148]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="121" phylum="Chordata" rank="species" species="damasense">
|
||
<emphasis id="B914193CFFDB6A7B08FAFBE9A027DD0E" box="[1158,1309,1124,1148]" italics="true" pageId="4" pageNumber="121">P. damasense</emphasis>
|
||
</taxonomicName>
|
||
) and in the color pattern of males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFDB6A7A0CBBFB27A030DF6A" lastPageId="5" lastPageNumber="122" pageId="4" pageNumber="121" type="description">
|
||
<paragraph id="8BDFC52EFFDB6A7B0CBBFB27A042DEAE" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="121">
|
||
<emphasis id="B914193CFFDB6A7B0CBBFB27A4D1DDB6" bold="true" box="[199,491,1195,1220]" pageId="4" pageNumber="121">
|
||
Description of
|
||
<typeStatus id="54DB7B8CFFDB6A7B0D01FB20A4DCDDB6" box="[381,486,1196,1220]" pageId="4" pageNumber="121" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Snout-vent length
|
||
<quantity id="4C9868CBFFDB6A7B0F6AFB20A6BBDDB6" box="[790,897,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="9.53" pageId="4" pageNumber="121" unit="mm" value="95.3">95.3 mm</quantity>
|
||
. Head length
|
||
<quantity id="4C9868CBFFDB6A7B0859FB20A1AADDB6" box="[1061,1168,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="4" pageNumber="121" unit="mm" value="17.6">17.6 mm</quantity>
|
||
. Head width
|
||
<quantity id="4C9868CBFFDB6A7B0952FB20A0A2DDB6" box="[1326,1432,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="4" pageNumber="121" unit="mm" value="16.7">16.7 mm</quantity>
|
||
. Head height (at parietal)
|
||
<quantity id="4C9868CBFFDB6A7B0DCFFB5CA736DD9A" box="[435,524,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="121" unit="mm" value="9.8">9.8 mm</quantity>
|
||
. Axilla-groin
|
||
<quantity id="4C9868CBFFDB6A7B0ED7FB5CA62ADD9A" box="[683,784,1232,1256]" metricMagnitude="-2" metricUnit="m" metricValue="5.14" pageId="4" pageNumber="121" unit="mm" value="51.4">51.4 mm</quantity>
|
||
(53.9 % of Snout-vent length). Tail length (excluding the regenerated part)
|
||
<quantity id="4C9868CBFFDB6A7B0D27FB78A4F8DC7E" box="[347,450,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" pageId="4" pageNumber="121" unit="mm" value="61.5">61.5 mm</quantity>
|
||
. Body slightly wide, trunk width:
|
||
<quantity id="4C9868CBFFDB6A7B0F42FB78A698DC7E" box="[830,930,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="4" pageNumber="121" unit="mm" value="36.4">36.4 mm</quantity>
|
||
(38.2 % of SVL). Twenty-two smooth dorsal head scales. Three scale organs in one postrostral and none in the other three postrostrals. Nasal bordered by nine scales, not in contact with rostral. Canthal separated from nasal by three scales. Loreal region slightly concave. Ten enlarged supralabial scales, none in contact with subocular. Ten enlarged infralabials. Auditory meatus oval, without enlarged scales in its anterior border. Auricular scale absent. Eleven convex, juxtaposed temporals. Rostral undivided. Mental subpentagonal, in contact with six scales. Interparietal bordered by eight scales. Frontal region without an azygous scale. Supraorbital semicircles inconspicuous. No distinctly enlarged supraoculars. Eleven imbricate flat superciliaries. Subocular unfragmented separated from supralabials by one to two rows of lorilabials. Seventeen lorilabials, the last subocular. Preocular separated from lorilabial row by four scales. Scales of throat round, flat, and juxtaposed.
|
||
<collectingCountry id="F37785BEFFDB6A7B0DAEF9B4A733DF22" box="[466,521,1592,1616]" name="China" pageId="4" pageNumber="121">Chin</collectingCountry>
|
||
shields undifferentiated. Sixty-six gulars between auditory meata. Lateral nuchal folds well developed, with granular scales over longitudinal fold. Antehumeral pocket well developed. Fifty-eight scales between auditory meatus and shoulder. In ventral view, gular fold not developed and posterior gular folds present with their anterior margins without enlarged scales on their borders. Lack of enlarged scales in the centre of chest. Dorsal scales round, smooth, juxtaposed. Forty-four dorsal scales along midline of the trunk in a distance equivalent to head length. Scales around midbody: 198. Mid-dorsal scales not enlarged compared with those on flanks. Ventral scales larger than dorsals. Ventral scales between mental and precloacal pores: 182. Eight precloacal pores in an undivided row plus one supernumerary pore. Brachial and antebrachial scales smooth with rounded posterior margins. Supracarpals laminar, round, smooth. Subdigital lamellae of fingers with three keels. Number of subdigital lamellae of fingers I: 8; II: 13; III: 15; IV: 20; V: 13. Claws moderately long. Supradigital lamellae convex, imbricate. Infracarpals and infratarsals with round margins and 2-3 obtuse mucrons. Supracarpals and supratarsals smooth, with round posterior margins. Subdigital lamellae of toes I: 9; II: 13; III: 17; IV: 23; V: 17.
|
||
</paragraph>
|
||
<caption id="DF1F95A6FFDA6A7A0CEBFBD6A034DD03" box="[151,1294,1114,1137]" httpUri="https://zenodo.org/record/223683/files/figure.png" pageId="5" pageNumber="122" targetBox="[158,1430,198,1089]" targetPageId="5">
|
||
<paragraph id="8BDFC52EFFDA6A7A0CEBFBD6A034DD03" blockId="5.[151,1294,1114,1137]" box="[151,1294,1114,1137]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CEBFBD6A436DD02" bold="true" box="[151,268,1114,1137]" pageId="5" pageNumber="122">FIGURE 2</emphasis>
|
||
. Patterns of coloration in life of
|
||
<taxonomicName id="4C60BEADFFDA6A7A0E2AFBD7A67EDD03" box="[598,836,1115,1137]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="122" phylum="Chordata" rank="species" species="aguanegra" status="sp. nov.">
|
||
<emphasis id="B914193CFFDA6A7A0E2AFBD7A67EDD03" box="[598,836,1115,1137]" italics="true" pageId="5" pageNumber="122">Phymaturus aguanegra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A227A447FFDA6A7A0F37FBD7A6A6DD03" box="[843,924,1115,1137]" pageId="5" pageNumber="122" rank="species">
|
||
<emphasis id="B914193CFFDA6A7A0F37FBD7A6A6DD03" bold="true" box="[843,924,1115,1137]" pageId="5" pageNumber="122">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A and B: Males; C and D: Females.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDFC52EFFDA6A7A0CB5FB15A0A3DD8A" blockId="5.[151,1436,1176,1669]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CB5FB15A4C6DDC2" box="[201,508,1176,1200]" italics="true" pageId="5" pageNumber="122">Pattern of body and limbs:</emphasis>
|
||
body trunk and dorsal surfaces of fore and hindlimbs covered by thinly scattered brown spots. No conspicuous pattern is present, except dorsum of neck and head, which becomes dark, almost melanic (
|
||
<figureCitation id="135BD9ABFFDA6A7A0C83FB6CA46FDD8A" box="[255,341,1248,1272]" captionStart="FIGURE 1. A" captionStartId="3.[151,250,1817,1840]" captionTargetBox="[151,1436,873,1795]" captionTargetId="figure@3.[151,1436,873,1797]" captionTargetPageId="3" captionText="FIGURE 1. A. Dorsal view of the holotype specimen of Phymaturus aguanegra sp. nov. (MCN 975) B. Ventral view of the same specimen. SVL: 95.3 mm." httpUri="https://zenodo.org/record/223682/files/figure.png" pageId="5" pageNumber="122">Fig. 1A</figureCitation>
|
||
&B). This dark color is extended over the sides of head and throat. Melanism can reach the chest.
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFDA6A7A0CAEFA88A103DC32" blockId="5.[151,1436,1176,1669]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CAEFA88A49ADC6E" box="[210,416,1284,1308]" italics="true" pageId="5" pageNumber="122">Coloration in life:</emphasis>
|
||
the general background coloration is brown-orange, giving an appearance of ferric oxide coloration (
|
||
<figureCitation id="135BD9ABFFDA6A7A0D64FAA4A467DC32" box="[280,349,1320,1344]" captionStart="FIGURE 2" captionStartId="5.[151,250,1114,1137]" captionTargetBox="[158,1430,198,1089]" captionTargetId="figure@5.[151,1436,193,1093]" captionTargetPageId="5" captionText="FIGURE 2. Patterns of coloration in life of Phymaturus aguanegra sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223683/files/figure.png" pageId="5" pageNumber="122">Fig. 2</figureCitation>
|
||
). In a few specimens a light yellow dorsal color can be observed.
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFDA6A7A0CB5FAC1A030DF6A" blockId="5.[151,1436,1176,1669]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CB5FAC1A400DC16" box="[201,314,1357,1380]" italics="true" pageId="5" pageNumber="122">Variation:</emphasis>
|
||
Based on
|
||
<specimenCount id="9D660EA7FFDA6A7A0DCFFAC0A735DC16" box="[435,527,1356,1380]" pageId="5" pageNumber="122" type="adult">28 adult</specimenCount>
|
||
specimens (
|
||
<specimenCount id="9D660EA7FFDA6A7A0EE1FAC0A639DC16" box="[669,771,1356,1380]" pageId="5" pageNumber="122" type="male">13 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D660EA7FFDA6A7A0F40FAC0A686DC16" box="[828,956,1356,1380]" pageId="5" pageNumber="122" type="female">15 females</specimenCount>
|
||
). SVL
|
||
<quantity id="4C9868CBFFDA6A7A0870FAC0A196DC16" box="[1036,1196,1356,1380]" metricMagnitude="-2" metricUnit="m" metricValue="8.47" metricValueMax="9.85" metricValueMin="7.09" pageId="5" pageNumber="122" unit="mm" value="84.7" valueMax="98.5" valueMin="70.9">70.9-98.5 mm</quantity>
|
||
(x = 85.6; SD = 9.1) for adult specimens only. Head length 15.8–20.6% (x = 18.8%; SD = 1.1) of SVL. Tail length 0.85–1.06 (x = 0.93; SD = 0.06) times SVL. Scales around midbody 165–220 (x = 191.7; SD = 13.3). Dorsal head scales 20–27 (x = 23.4; SD = 1.9). Ventrals 163–194 (x = 181.1; SD = 9.2). Scales surrounding interparietal 7–11 (x = 9; SD = 0.9). Scales of neck along longitudinal fold from posterior border of auditory meatus to shoulder 54–70 (x = 63.2; SD = 4.4). Gulars 56–98 (x = 74.3; SD = 11.4). Scales between rostral and frontal 8–12 (x = 9.5; SD = 1.3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFDA6A7A0CB5F9A9A61CDF12" pageId="5" pageNumber="122" type="etymology">
|
||
<paragraph id="8BDFC52EFFDA6A7A0CB5F9A9A61CDF12" blockId="5.[151,1436,1176,1669]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CB5F9A9A475DF4F" bold="true" box="[201,335,1573,1597]" pageId="5" pageNumber="122">Etymology.</emphasis>
|
||
The specific epithet is given after its
|
||
<typeStatus id="54DB7B8CFFDA6A7A0E82F9A9A614DF4F" box="[766,814,1573,1597]" pageId="5" pageNumber="122">type</typeStatus>
|
||
locality. [“Agua Negra”] toponym, which refers to the dark water aspect of Andean streams near the
|
||
<typeStatus id="54DB7B8CFFDA6A7A0EE7F9C5A7F1DF13" box="[667,715,1609,1633]" pageId="5" pageNumber="122">type</typeStatus>
|
||
locality
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFDA6A7A0CB5F9E0A7D1DFF6" box="[201,747,1644,1669]" pageId="5" pageNumber="122" type="distribution">
|
||
<paragraph id="8BDFC52EFFDA6A7A0CB5F9E0A7D1DFF6" blockId="5.[151,1436,1176,1669]" box="[201,747,1644,1669]" pageId="5" pageNumber="122">
|
||
<emphasis id="B914193CFFDA6A7A0CB5F9E0A458DFF7" bold="true" box="[201,354,1644,1669]" pageId="5" pageNumber="122">Distribution.</emphasis>
|
||
Only known from its
|
||
<typeStatus id="54DB7B8CFFDA6A7A0E26F9E1A7B0DFF7" box="[602,650,1645,1669]" pageId="5" pageNumber="122">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |