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<document id="3642BCEDBDAAD2855280A6C8699E5F05" ID-CLB-Dataset="29397" ID-DOI="10.11646/zootaxa.4462.3.1" ID-GBIF-Dataset="ee3428d1-9e3b-495d-99a5-e61ea679b5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441611" ID-ZooBank="75D121A7-F718-4F69-9E33-E82AB371403A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538403914940" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2018" docId="03C887DB60748C13FF67D2BB1A48FCDE" docLanguage="en" docName="zootaxa.4462.3.1.pdf" docOrigin="Zootaxa 4462 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asiagomphus pacificus" docType="treatment" docVersion="8" lastPageNumber="318" masterDocId="FFF1FFA360658C01FFF0D13A1956FF82" masterDocTitle="The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)" masterLastPageNumber="330" masterPageNumber="301" pageNumber="318" updateTime="1698561160019" updateUser="ExternalLinkService">
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<mods:title id="8575B31C939EC2A41123E284CC36CF77">The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)</mods:title>
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<mods:date id="B507353633739E5F3CBC27A3077B33AF">2018</mods:date>
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<treatment id="03C887DB60748C13FF67D2BB1A48FCDE" ID-DOI="http://doi.org/10.5281/zenodo.5974771" ID-GBIF-Taxon="148424958" ID-Zenodo-Dep="5974771" LSID="urn:lsid:plazi:treatment:03C887DB60748C13FF67D2BB1A48FCDE" httpUri="http://treatment.plazi.org/id/03C887DB60748C13FF67D2BB1A48FCDE" lastPageId="18" lastPageNumber="318" pageId="17" pageNumber="318">
<subSubSection id="C37B654660748C10FF67D2BB1B07FC3F" pageId="17" pageNumber="318" type="nomenclature">
<paragraph id="8BDE36CD60748C10FF67D2BB1B00FC19" blockId="17.[151,598,897,957]" box="[151,598,897,923]" pageId="17" pageNumber="318">
<heading id="D09681A160748C10FF67D2BB1B00FC19" bold="true" box="[151,598,897,923]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<taxonomicName id="4C614D4E60748C10FF67D2BB1B00FC19" ID-CoL="H9SQ" authority="(Chao, 1953)" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[151,598,897,923]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FF67D2BB1B00FC19" bold="true" box="[151,598,897,923]" pageId="17" pageNumber="318">
<emphasis id="B915EADF60748C10FF67D2BB18E6FC19" bold="true" box="[151,432,897,923]" italics="true" pageId="17" pageNumber="318">Asiagomphus pacificus</emphasis>
(Chao, 1953)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDE36CD60748C10FF67D29E1B07FC3F" blockId="17.[151,598,897,957]" box="[151,593,932,957]" pageId="17" pageNumber="318">
(
<figureCitation id="135A2A4860748C10FF6FD29E184EFC3F" box="[159,280,932,957]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[175,1416,193,1930]" captionTargetId="figure@9.[175,1416,193,1931]" captionTargetPageId="9" captionText="FIGURE 4. Anterior and posterior hamuli [a,c,e,g] lateral view, [b,d,f,h] antero-ventral lateral view. (ab) Asiagomphus auricolor male used for description; (cd) A. pacificus Cao Bang Prov., 19-V-2015; (ef) A. monticola sp. nov. holotype; (gh) A. superciliaris sp. nov. holotype." httpUri="https://zenodo.org/record/1441619/files/figure.png" pageId="17" pageNumber="318">Figs. 4cd</figureCitation>
,
<figureCitation id="135A2A4860748C10FED3D29E1817FC3F" box="[291,321,932,957]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="17" pageNumber="318">5b</figureCitation>
,
<figureCitation id="135A2A4860748C10FEBDD29E183DFC3F" box="[333,363,932,957]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="17" pageNumber="318">6b</figureCitation>
,
<figureCitation id="135A2A4860748C10FE87D29E18C3FC3F" box="[375,405,932,957]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="17" pageNumber="318">7b</figureCitation>
,
<figureCitation id="135A2A4860748C10FE51D29E18EBFC3E" box="[417,445,932,956]" captionStart="FIGURE 8" captionStartId="14.[151,250,1755,1777]" captionTargetBox="[174,1436,447,1721]" captionTargetId="figure@14.[151,1436,447,1733]" captionTargetPageId="14" captionText="FIGURE 8. Right branch of male epiproct in dorso-lateral view. (a) Asiagomphus auricolor male used for description; (b) A. monticola holotype; (c) A. pacificus Cao Bang Prov., 19-V-2015; (d) A. kosterini holotype; (e) A. superciliaris holotype." httpUri="https://zenodo.org/record/1441627/files/figure.png" pageId="17" pageNumber="318">8c</figureCitation>
,
<figureCitation id="135A2A4860748C10FE39D29E1B44FC3F" box="[457,530,932,957]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [ad, g] male, [ef, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="17" pageNumber="318">
<quantity id="4C999B2860748C10FE39D29E18A5FC3F" box="[457,499,932,957]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="17" pageNumber="318" unit="g" value="10.0">10g</quantity>
h
</figureCitation>
,
<figureCitation id="135A2A4860748C10FDEDD29E1B11FC3F" box="[541,583,932,957]" captionStart="FIGURE 13" captionStartId="22.[151,250,1766,1788]" captionTargetBox="[151,1437,481,1745]" captionTargetId="figure@22.[151,1437,481,1745]" captionTargetPageId="22" captionText="FIGURE 13. Asiagomphus spp. males in nature. (a) A. acco, Xuan Son NP, 8-VI-2014; (b) A. acco, Pia Oac NR, 8-VII-2014; (c) A. kosterini, Lam Dong Prov., 17-IV-2016; (d) A. pacificus, Cao Bang Prov., 19-V-2015; (e) A. reinhardti, Gia Lai Prov., 20- VI-2016; (f) A. reinhardti, Gia Lai Prov., 13-V-2017." httpUri="https://zenodo.org/record/1441637/files/figure.png" pageId="17" pageNumber="318">13d</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BDE36CD60748C10FF67D2DC1B85FC7E" blockId="17.[151,1436,998,1174]" box="[151,723,998,1020]" pageId="17" pageNumber="318">
<taxonomicName id="4C614D4E60748C10FF67D2DC1876FC7E" authorityName="Asahina" authorityYear="1985" box="[151,288,998,1020]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60748C10FF67D2DC1876FC7E" box="[151,288,998,1020]" italics="true" pageId="17" pageNumber="318">Asiagomphus</emphasis>
</taxonomicName>
sp.:
<bibRefCitation id="EFF04B3C60748C10FEA0D2DC18B7FC7E" box="[336,481,998,1020]" pageId="17" pageNumber="318" refId="ref17777" refString="Wilson, K. D. P. (1999) Dragonflies (Odonata) of Dinghu Shan Biosphere Reserve, Guangdong Province, China. International Journal of Odonatology, 2, 23 - 53. https: // doi. org / 10.1080 / 13887890.1999.9748111" type="journal article">Wilson (1999)</bibRefCitation>
pp. 3436, figs. 1920;
</paragraph>
</subSubSection>
<subSubSection id="C37B654660748C10FF67D53F184FFB14" pageId="17" pageNumber="318" type="reference_group">
<paragraph id="8BDE36CD60748C10FF67D53F1B6CFBDA" blockId="17.[151,1436,998,1174]" pageId="17" pageNumber="318">
<taxonomicName id="4C614D4E60748C10FF67D53F1B74FB99" ID-CoL="H9SQ" authority=": Wilson (2005)" authorityName=": Wilson" authorityYear="2005" box="[151,546,1029,1051]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FF67D53F18D7FB99" box="[151,385,1029,1051]" italics="true" pageId="17" pageNumber="318">Asiagomphus pacificus</emphasis>
:
<bibRefCitation id="EFF04B3C60748C10FE7FD53F1B74FB99" box="[399,546,1029,1051]" pageId="17" pageNumber="318" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson (2005)</bibRefCitation>
</taxonomicName>
pp. 131132, figs. 14ad [records from Guangxi, China];
<bibRefCitation id="EFF04B3C60748C10FB86D53F1C1EFB99" box="[1142,1352,1029,1051]" pageId="17" pageNumber="318" refId="ref17870" refString="Wilson, K. D. P. &amp; Xu, Z. F. (2009) Gomphidae of Guangdong &amp; Hong Kong, China (Odonata: Anisoptera). Zootaxa, 2177, 1 - 62." type="journal article">Wilson &amp; Xu (2009)</bibRefCitation>
pp. 12 13, figs. 2ad [comparison with
<taxonomicName id="4C614D4E60748C10FDE0D51E1BF3FBB8" ID-CoL="H9SF" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[528,677,1060,1082]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FDE0D51E1BF3FBB8" box="[528,677,1060,1082]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
, reappraisal of historic records of
<taxonomicName id="4C614D4E60748C10FBF3D51E1DCEFBB8" ID-CoL="H9SF" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[1027,1176,1060,1082]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FBF3D51E1DCEFBB8" box="[1027,1176,1060,1082]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
from Guangdong, China, overview of records from literature].
</paragraph>
<paragraph id="8BDE36CD60748C10FF67D558184FFB14" blockId="17.[151,1436,998,1174]" pageId="17" pageNumber="318">
[For extensive overview of literature with records of this species, see
<bibRefCitation id="EFF04B3C60748C10FC82D55B1AA1FBF5" box="[882,1015,1121,1143]" pageId="17" pageNumber="318" refId="ref17086" refString="Chao, H. F. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou, Fujian, 14 + 486 pp." type="book">Chao (1990)</bibRefCitation>
,
<bibRefCitation id="EFF04B3C60748C10FBF3D55B1DC7FBF5" box="[1027,1169,1121,1143]" pageId="17" pageNumber="318" refId="ref17777" refString="Wilson, K. D. P. (1999) Dragonflies (Odonata) of Dinghu Shan Biosphere Reserve, Guangdong Province, China. International Journal of Odonatology, 2, 23 - 53. https: // doi. org / 10.1080 / 13887890.1999.9748111" type="journal article">Wilson (1999</bibRefCitation>
,
<bibRefCitation id="EFF04B3C60748C10FB6ED55B1D84FBF5" box="[1182,1234,1121,1143]" pageId="17" pageNumber="318" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">2005</bibRefCitation>
) and
<bibRefCitation id="EFF04B3C60748C10FAE0D55B185AFB14" pageId="17" pageNumber="318" refId="ref17870" refString="Wilson, K. D. P. &amp; Xu, Z. F. (2009) Gomphidae of Guangdong &amp; Hong Kong, China (Odonata: Anisoptera). Zootaxa, 2177, 1 - 62." type="journal article">Wilson &amp; Xu (2009)</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C37B654660748C10FF67D5F81CC3FAC5" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="8BDE36CD60748C10FF67D5F81DCBFB7D" blockId="17.[151,1436,1218,2035]" pageId="17" pageNumber="318">
<materialsCitation id="3B093C9060748C10FF67D5F81D68FB59" ID-GBIF-Occurrence="1913893013" box="[151,1086,1218,1243]" collectingDate="2015-05-19" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
<emphasis id="B915EADF60748C10FF67D5F818D2FB59" bold="true" box="[151,388,1218,1243]" pageId="17" pageNumber="318">Material examined:</emphasis>
<specimenCount id="9D67FD4460748C10FE7DD5F918EFFB58" box="[397,441,1219,1243]" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
<collectingRegion id="49A5F82F60748C10FE32D5F91B65FB59" box="[450,563,1219,1243]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60748C10FD1DD5F81A19FB59" box="[749,847,1218,1243]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560748C10FCACD5F81A81FB59" box="[860,983,1218,1243]" pageId="17" pageNumber="318" value="2015-05-19">
<date id="FFDF100D60748C10FCACD5F81A81FB59" box="[860,983,1218,1243]" pageId="17" pageNumber="318" value="2015-05-19">19-V-2015</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060748C10FBBCD5F918BAFB7D" ID-GBIF-Occurrence="1913893031" collectingDate="2014-06-24" country="Vietnam" location="Bac Kan" pageId="17" pageNumber="318" specimenCount="1" specimenCount-female="1" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460748C10FBBCD5F91D2EFB58" box="[1100,1144,1219,1243]" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49A5F82F60748C10FB78D5F91DBDFB58" box="[1160,1259,1219,1242]" country="Vietnam" name="Bac Kan" pageId="17" pageNumber="318">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60748C10FF67D5DC19AFFB7D" box="[151,249,1254,1279]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560748C10FEF4D5DD18DCFB7D" box="[260,394,1255,1279]" pageId="17" pageNumber="318" value="2014-06-24">
<date id="FFDF100D60748C10FEF4D5DD18DCFB7D" box="[260,394,1255,1279]" pageId="17" pageNumber="318" value="2014-06-24">24-VI-2014</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060748C10FE0AD5DD1DCBFB7D" ID-GBIF-Occurrence="1913893015" box="[506,1181,1254,1279]" collectingDate="2018-06-06" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="1" specimenCount-female="1" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460748C10FE0AD5DD1B72FB7C" box="[506,548,1255,1279]" pageId="17" pageNumber="318" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49A5F82F60748C10FDC2D5DD1BF7FB7D" box="[562,673,1255,1279]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60748C10FCA6D5DC1AEEFB7D" box="[854,952,1254,1279]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560748C10FC33D5DC1D6DFB7D" box="[963,1083,1254,1279]" pageId="17" pageNumber="318" value="2018-06-06">
<date id="FFDF100D60748C10FC33D5DC1D6DFB7D" box="[963,1083,1254,1279]" pageId="17" pageNumber="318" value="2018-06-06">6-VI-2018</date>
</collectingDate>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="8BDE36CD60748C10FF37D4301CC3FAC5" blockId="17.[151,1436,1218,2035]" pageId="17" pageNumber="318">
<materialsCitation id="3B093C9060748C10FF37D4301985FAC5" ID-GBIF-Occurrence="1913893030" collectingDate="2015-05-19" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="4" specimenCount-male="4" stateProvince="Cao Bang">
<emphasis id="B915EADF60748C10FF37D4301A37FAA1" bold="true" box="[199,865,1290,1315]" pageId="17" pageNumber="318">Additional records based on field observations by TK:</emphasis>
<specimenCount id="9D67FD4460748C10FC9BD4311AF9FAA0" box="[875,943,1291,1315]" pageId="17" pageNumber="318" type="male">4 ♂♂</specimenCount>
,
<collectingRegion id="49A5F82F60748C10FC4DD4311D79FAA1" box="[957,1071,1291,1315]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60748C10FB1CD4301C18FAA1" box="[1260,1358,1290,1315]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560748C10FAACD4311985FAC5" pageId="17" pageNumber="318" value="2015-05-19">
<date id="FFDF100D60748C10FAACD4311985FAC5" pageId="17" pageNumber="318" value="2015-05-19">19-V- 2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060748C10FF2DD4151B19FAC5" ID-GBIF-Occurrence="1913893020" box="[221,591,1326,1351]" collectingDate="2015-05-20" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="3" specimenCount-male="3" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460748C10FF2DD4151849FAC4" box="[221,287,1327,1351]" pageId="17" pageNumber="318" type="male">3 ♂♂</specimenCount>
, same location,
<collectingDate id="EF9BE9E560748C10FE23D4141B19FAC5" box="[467,591,1326,1351]" pageId="17" pageNumber="318" value="2015-05-20">
<date id="FFDF100D60748C10FE23D4141B19FAC5" box="[467,591,1326,1351]" pageId="17" pageNumber="318" value="2015-05-20">20-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060748C10FDAAD4151A95FAC5" ID-GBIF-Occurrence="1913893023" box="[602,963,1326,1351]" collectingDate="2015-06-18" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460748C10FDAAD4151BD1FAC4" box="[602,647,1327,1351]" pageId="17" pageNumber="318" type="male">1 ♂</specimenCount>
, same location,
<collectingDate id="EF9BE9E560748C10FCCBD4141A95FAC5" box="[827,963,1326,1351]" pageId="17" pageNumber="318" value="2015-06-18">
<date id="FFDF100D60748C10FCCBD4141A95FAC5" box="[827,963,1326,1351]" pageId="17" pageNumber="318" value="2015-06-18">18-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060748C10FC3ED4151CC4FAC5" ID-GBIF-Occurrence="1913893029" box="[974,1426,1326,1351]" collectingDate="2018-06-06" country="Vietnam" location="Cao Bang" pageId="17" pageNumber="318" specimenCount="23" specimenCount-female="3" specimenCount-male="20" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460748C10FC3ED4151D4DFAC4" box="[974,1051,1327,1351]" pageId="17" pageNumber="318" type="male">20 ♂♂</specimenCount>
<specimenCount id="9D67FD4460748C10FBD2D4151D34FAC4" box="[1058,1122,1327,1351]" pageId="17" pageNumber="318" type="female">3 ♀♀</specimenCount>
, same location,
<collectingDate id="EF9BE9E560748C10FAE8D4141CC4FAC5" box="[1304,1426,1326,1351]" pageId="17" pageNumber="318" value="2018-06-06">
<date id="FFDF100D60748C10FAE8D4141CC4FAC5" box="[1304,1426,1326,1351]" pageId="17" pageNumber="318" value="2018-06-06">6-VI-2018</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B654660748C10FF37D4681850F92C" pageId="17" pageNumber="318" type="diagnosis">
<paragraph id="8BDE36CD60748C10FF37D4681B73FA31" blockId="17.[151,1436,1218,2035]" pageId="17" pageNumber="318">
<emphasis id="B915EADF60748C10FF37D4681898FAE9" bold="true" box="[199,462,1362,1387]" pageId="17" pageNumber="318">Differential diagnosis.</emphasis>
For differences between
<taxonomicName id="4C614D4E60748C10FD1BD4681AB9FAE9" authorityName=": Wilson" authorityYear="2005" box="[747,1007,1362,1387]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FD1BD4681AB9FAE9" box="[747,1007,1362,1387]" italics="true" pageId="17" pageNumber="318">Asiagomphus pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D4E60748C10FBD6D4691D9DFAE9" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[1062,1227,1362,1387]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FBD6D4691D9DFAE9" box="[1062,1227,1362,1387]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
see the discussion below.
<taxonomicName id="4C614D4E60748C10FF1AD44C18A6FA0D" authorityName=": Wilson" authorityYear="2005" box="[234,496,1398,1423]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FF1AD44C18A6FA0D" box="[234,496,1398,1423]" italics="true" pageId="17" pageNumber="318">Asiagomphus pacificus</emphasis>
</taxonomicName>
can be separated from all other Vietnamese species by the longitudinal markings along the dorsal carina of the S26.
</paragraph>
<paragraph id="8BDE36CD60748C10FF37D4841850F92C" blockId="17.[151,1436,1218,2035]" pageId="17" pageNumber="318">
<emphasis id="B915EADF60748C10FF37D4841894FA55" bold="true" box="[199,450,1470,1495]" pageId="17" pageNumber="318">Habitat and ecology.</emphasis>
In
<collectingRegion id="49A5F82F60748C10FE1DD4851B98FA55" box="[493,718,1470,1495]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang Province</collectingRegion>
the species occurred at a medium-sized river, with sandy and pebbly substrates and larger rocks in a forested valley close to
<collectingRegion id="49A5F82F60748C10FCAFD4D91A99FA79" box="[863,975,1507,1531]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang</collectingRegion>
City. Males perched in the sun on larger rocks in the stream or on pebble beaches by the side of the river, whereas females where found perched on vegetation a little removed from the stream. The female in
<collectingRegion id="49A5F82F60748C10FCDAD7111AA0F9C1" box="[810,1014,1578,1603]" country="Vietnam" name="Bac Kan" pageId="17" pageNumber="318">Bac Kan Province</collectingRegion>
was found at a shallow rocky stream, over a pebble substrate in patchy degraded forest between cleared hills close to Ba Be NP, perched over the stream in bushes at head height. In
<collectingRegion id="49A5F82F60748C10FE1DD7491B37F909" box="[493,609,1651,1675]" country="Vietnam" name="Cao Bang" pageId="17" pageNumber="318">Cao Bang</collectingRegion>
it occurred together with
<taxonomicName id="4C614D4E60748C10FC6BD7481D25F909" box="[923,1139,1650,1675]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60748C10FC6BD7481D25F909" box="[923,1139,1650,1675]" italics="true" pageId="17" pageNumber="318">Asiagomphus acco</emphasis>
</taxonomicName>
and in
<collectingRegion id="49A5F82F60748C10FB24D7491C6CF908" box="[1236,1338,1651,1674]" country="Vietnam" name="Bac Kan" pageId="17" pageNumber="318">Bac Kan</collectingRegion>
with
<taxonomicName id="4C614D4E60748C10FA74D74919A9F92D" authorityName=": Fraser" authorityYear="1926" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60748C10FA74D74919A9F92D" italics="true" pageId="17" pageNumber="318">A. auricolor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B654660748C13FF37D7801A48FCDE" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318" type="discussion">
<paragraph id="8BDE36CD60748C13FF37D7801A48FCDE" blockId="17.[151,1436,1218,2035]" lastBlockId="18.[151,1437,151,861]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
<emphasis id="B915EADF60748C10FF37D780181AF951" bold="true" box="[199,332,1722,1747]" pageId="17" pageNumber="318">Discussion.</emphasis>
There has been considerable confusion regarding the identity of two very similar
<taxonomicName id="4C614D4E60748C10FAF5D7801CCAF951" authorityName="Asahina" authorityYear="1985" box="[1285,1436,1722,1747]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60748C10FAF5D7801CCAF951" box="[1285,1436,1722,1747]" italics="true" pageId="17" pageNumber="318">Asiagomphus</emphasis>
</taxonomicName>
species with longitudinal markings along the dorsum of the abdomen that occur in southern
<collectingCountry id="F376765D60748C10FB68D7E41D89F975" box="[1176,1247,1758,1783]" name="China" pageId="17" pageNumber="318">China</collectingCountry>
:
<taxonomicName id="4C614D4E60748C10FB19D7E51C3BF975" authorityName=": Wilson" authorityYear="2005" box="[1257,1389,1759,1783]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FB19D7E51C3BF975" box="[1257,1389,1759,1783]" italics="true" pageId="17" pageNumber="318">A. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D4E60748C10FF67D639186AF899" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[151,316,1794,1819]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FF67D639186AF899" box="[151,316,1794,1819]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF04B3C60748C10FEBBD63818BAF899" box="[331,492,1794,1819]" pageId="17" pageNumber="318" refId="ref17777" refString="Wilson, K. D. P. (1999) Dragonflies (Odonata) of Dinghu Shan Biosphere Reserve, Guangdong Province, China. International Journal of Odonatology, 2, 23 - 53. https: // doi. org / 10.1080 / 13887890.1999.9748111" type="journal article">Wilson (1999)</bibRefCitation>
recounts the confusion surrounding the identity of
<taxonomicName id="4C614D4E60748C10FBDDD6391D85F899" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[1069,1235,1794,1819]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FBDDD6391D85F899" box="[1069,1235,1794,1819]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F376765D60748C10FAE8D6391CCAF899" box="[1304,1436,1795,1819]" name="Hong Kong" pageId="17" pageNumber="318">Hong Kong</collectingCountry>
described by Asahina, first (1966) as
<taxonomicName id="4C614D4E60748C10FDCBD61D1BB6F8BD" baseAuthorityName="Chao" baseAuthorityYear="1953" box="[571,736,1830,1855]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FDCBD61D1BB6F8BD" box="[571,736,1830,1855]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
and later (1988) reidentified as
<taxonomicName id="4C614D4E60748C10FBB8D61D1CCEF8BD" authority="(Needham, 1930)" baseAuthorityName="Needham" baseAuthorityYear="1930" box="[1096,1432,1830,1855]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="septimus">
<emphasis id="B915EADF60748C10FBB8D61D1D9CF8BC" box="[1096,1226,1831,1854]" italics="true" pageId="17" pageNumber="318">A. septimus</emphasis>
(Needham, 1930)
</taxonomicName>
.
<bibRefCitation id="EFF04B3C60748C10FF67D670186DF8E1" box="[151,315,1866,1891]" pageId="17" pageNumber="318" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson (2005)</bibRefCitation>
and
<bibRefCitation id="EFF04B3C60748C10FE88D6701B3EF8E1" box="[376,616,1866,1891]" pageId="17" pageNumber="318" refId="ref17870" refString="Wilson, K. D. P. &amp; Xu, Z. F. (2009) Gomphidae of Guangdong &amp; Hong Kong, China (Odonata: Anisoptera). Zootaxa, 2177, 1 - 62." type="journal article">Wilson &amp; Xu (2009)</bibRefCitation>
discussed the differences between the very similar
<taxonomicName id="4C614D4E60748C10FB30D6711C11F8E1" authorityName=": Wilson" authorityYear="2005" box="[1216,1351,1867,1891]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FB30D6711C11F8E1" box="[1216,1351,1867,1891]" italics="true" pageId="17" pageNumber="318">A. pacificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D4E60748C10FA74D671184BF805" baseAuthorityName="Chao" baseAuthorityYear="1953" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FA74D671184BF805" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
</taxonomicName>
, focusing on differences of body structure and patterning: “[
<taxonomicName id="4C614D4E60748C10FBF2D6551DDAF805" authorityName=": Wilson" authorityYear="2005" box="[1026,1164,1903,1927]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FBF2D6551DDAF805" box="[1026,1164,1903,1927]" italics="true" pageId="17" pageNumber="318">A. pacificus</emphasis>
</taxonomicName>
is] very similar to
<taxonomicName id="4C614D4E60748C10FA74D6551832F829" authority="Chao" authorityName="Chao" authorityYear="1953" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FA74D655184BF829" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
Chao
</taxonomicName>
, which has more extensive yellow coloration on the sides of the synthorax, a pair of quadrate spots on the labrum and is generally a less robust insect. The labrum of
<taxonomicName id="4C614D4E60748C10FC42D68C1D41F84D" authorityName=": Wilson" authorityYear="2005" box="[946,1047,1974,1999]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">pacificus</taxonomicName>
is entirely black” (
<bibRefCitation id="EFF04B3C60748C10FB1AD68C1994F871" pageId="17" pageNumber="318" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson 2005, p. 132</bibRefCitation>
) and “
<taxonomicName id="4C614D4E60748C10FEFED6E01B44F871" authorityName=": Wilson" authorityYear="2005" box="[270,530,2010,2035]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60748C10FEFED6E01B44F871" box="[270,530,2010,2035]" italics="true" pageId="17" pageNumber="318">Asiagomphus pacificus</emphasis>
</taxonomicName>
is significantly larger and slightly bulkier species than
<taxonomicName id="4C614D4E60748C10FB74D6E11C3AF871" authority="Chao" authorityName="Chao" authorityYear="1953" box="[1156,1388,2010,2035]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60748C10FB74D6E11C7FF871" box="[1156,1321,2010,2035]" italics="true" pageId="17" pageNumber="318">A. hainanensis</emphasis>
Chao
</taxonomicName>
and its males feature a uniformly straight occipital margin without any small spines. Its antehumeral stripe may be complete or interrupted in the middle […] and its labrum is entirely black” (
<bibRefCitation id="EFF04B3C60778C13FC0DD1861D8DFF56" box="[1021,1243,188,213]" pageId="18" pageNumber="319" refId="ref17870" refString="Wilson, K. D. P. &amp; Xu, Z. F. (2009) Gomphidae of Guangdong &amp; Hong Kong, China (Odonata: Anisoptera). Zootaxa, 2177, 1 - 62." type="journal article">Wilson 2009, p. 13</bibRefCitation>
). Based on these differences, Wilson identified specimens from
<collectingRegion id="49A5F82F60778C13FD55D1E51A53FF7A" box="[677,773,223,248]" country="China" name="Guangxi" pageId="18" pageNumber="319">Guangxi</collectingRegion>
and
<collectingRegion id="49A5F82F60778C13FCCCD1E51AE9FF7A" box="[828,959,223,248]" country="China" name="Guangdong" pageId="18" pageNumber="319">Guangdong</collectingRegion>
as
<taxonomicName id="4C614D4E60778C13FC15D1DA1D3FFF7A" authorityName=": Wilson" authorityYear="2005" box="[997,1129,224,248]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60778C13FC15D1DA1D3FFF7A" box="[997,1129,224,248]" italics="true" pageId="18" pageNumber="319">A. pacificus</emphasis>
</taxonomicName>
. Mr. Wen-Chi Yeh (
<emphasis id="B915EADF60778C13FABFD1DB1CCDFF7A" box="[1359,1435,223,248]" italics="true" pageId="18" pageNumber="319">in litt.)</emphasis>
pointed out that male specimens from
<collectingCountry id="F376765D60778C13FDB7D03E1BCFFE9F" box="[583,665,260,285]" name="Taiwan" pageId="18" pageNumber="319">Taiwan</collectingCountry>
of these two species are often difficult to reliably separate, although dorsal stripes in
<taxonomicName id="4C614D4E60778C13FEBFD0121885FEC2" authorityName=": Wilson" authorityYear="2005" box="[335,467,296,320]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60778C13FEBFD0121885FEC2" box="[335,467,296,320]" italics="true" pageId="18" pageNumber="319">A. pacificus</emphasis>
</taxonomicName>
tend to be more parallel-sided and more weakly connected to the collar, whereas in
<taxonomicName id="4C614D4E60778C13FA74D012184BFEE7" authorityName="Chao" authorityYear="1953" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B915EADF60778C13FA74D012184BFEE7" italics="true" pageId="18" pageNumber="319">A. hainanensis</emphasis>
</taxonomicName>
the dorsal stripes are more broadly connected to the collar stripes and attenuate upwards. Their caudal appendages and secondary genitalia are not distinctly different. My observations of other
<taxonomicName id="4C614D4E60778C13FB5AD0551C17FE0A" authorityName="Asahina" authorityYear="1985" box="[1194,1345,367,392]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60778C13FB5AD0551C17FE0A" box="[1194,1345,367,392]" italics="true" pageId="18" pageNumber="319">Asiagomphus</emphasis>
</taxonomicName>
species show that the presence or absence of small quadrate spots on the labrum may vary within a species, and so does size. The observed males in
<collectingRegion id="49A5F82F60778C13FE23D0821BFBFE52" box="[467,685,439,464]" country="Vietnam" name="Cao Bang" pageId="18" pageNumber="319">Cao Bang Province</collectingRegion>
moreover displayed both parallel-sided dorsal stripes more weakly connected to the collar and more upwardly attenuated dorsal stripes broadly connected to the collar. However, the identity of my specimens, which also have all black labrum, is further supported by the spines adjacent to the lateral ocelli in the females. Mr. Wen-Chi Yeh (
<emphasis id="B915EADF60778C13FD36D31F1A5AFDBF" box="[710,780,548,573]" italics="true" pageId="18" pageNumber="319">in litt.</emphasis>
) commented that although the differences between their vulvar laminae do not provide a reliable help to identification, at least in
<collectingCountry id="F376765D60778C13FC12D37D1D62FDE2" box="[994,1076,583,608]" name="Taiwan" pageId="18" pageNumber="319">Taiwan</collectingCountry>
females of both species can be reliably separated on the basis of their head structures. Like my specimens,
<taxonomicName id="4C614D4E60778C13FC06D3561D2DFD06" authorityName=": Wilson" authorityYear="2005" box="[1014,1147,620,644]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60778C13FC06D3561D2DFD06" box="[1014,1147,620,644]" italics="true" pageId="18" pageNumber="319">A. pacificus</emphasis>
</taxonomicName>
on
<collectingCountry id="F376765D60778C13FB58D3561DACFD07" box="[1192,1274,620,645]" name="Taiwan" pageId="18" pageNumber="319">Taiwan</collectingCountry>
has one spine between each lateral ocellus and the seam of the compound eye and another at the posterior end of the ridges shielding the lateral ocelli posteriorly. It has therefore four prominent spines around the ocelli. Even though my specimens have a small spine on the occipital ridge not seen in the Taiwanese specimens, this strongly supports the identification of the Vietnamese material as
<taxonomicName id="4C614D4E60778C13FD7AD3C61A59FC96" authorityName=": Wilson" authorityYear="2005" box="[650,783,764,788]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60778C13FD7AD3C61A59FC96" box="[650,783,764,788]" italics="true" pageId="18" pageNumber="319">A. pacificus</emphasis>
</taxonomicName>
. This is in line with the conclusion by
<bibRefCitation id="EFF04B3C60778C13FB39D3C61C3DFC97" box="[1225,1387,764,789]" pageId="18" pageNumber="319" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson (2005)</bibRefCitation>
and
<bibRefCitation id="EFF04B3C60778C13FF67D22518DCFCBA" box="[151,394,799,824]" pageId="18" pageNumber="319" refId="ref17870" refString="Wilson, K. D. P. &amp; Xu, Z. F. (2009) Gomphidae of Guangdong &amp; Hong Kong, China (Odonata: Anisoptera). Zootaxa, 2177, 1 - 62." type="journal article">Wilson &amp; Xu (2009)</bibRefCitation>
, although they used different characters to support their conclusion, that Chinese material from
<collectingRegion id="49A5F82F60778C13FF24D27E1862FCDF" box="[212,308,836,861]" country="China" name="Guangxi" pageId="18" pageNumber="319">Guangxi</collectingRegion>
and
<collectingRegion id="49A5F82F60778C13FE9BD27E18B8FCDF" box="[363,494,836,861]" country="China" name="Guangdong" pageId="18" pageNumber="319">Guangdong</collectingRegion>
also concerns
<taxonomicName id="4C614D4E60778C13FD63D27E1A41FCDE" authorityName=": Wilson" authorityYear="2005" box="[659,791,836,860]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B915EADF60778C13FD63D27E1A41FCDE" box="[659,791,836,860]" italics="true" pageId="18" pageNumber="319">A. pacificus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>