174 lines
18 KiB
XML
174 lines
18 KiB
XML
<document id="A0C23DD12079B331764A45B72C488375" ID-DOI="10.11646/zootaxa.4755.1.6" ID-GBIF-Dataset="9f6752d6-7c9a-4756-acfe-badb2a0301d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3724291" ID-ZooBank="D0B3FFA8-C262-4CD6-98AB-B46C55FCEF6D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584968592865" checkinUser="plazi" docAuthor="Busanello, Dilson G. C., Irmler, Ulrich & Caron, Edilson" docDate="2020" docId="03C887D5FF8FFF8D83C6F9DFC3A9FE50" docLanguage="en" docName="zootaxa.4755.1.6.pdf" docOrigin="Zootaxa 4755 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudastenus Bernhauer 1933" docType="key" docVersion="5" lastPageNumber="142" masterDocId="FFF1FFADFF8DFF8E8351FF82C634FFE6" masterDocTitle="Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)" masterLastPageNumber="154" masterPageNumber="139" pageNumber="141" updateTime="1698730789478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B58B4E7A842C3B8069B553081B14FDAF">Revision of Pseudastenus Bernhauer, synonym with Xenasterides Newton and six new species (Coleoptera, Staphylinidae, Paederinae)</mods:title>
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<mods:namePart id="5BFF8C786B9082BF5A955D0785CFA381">Busanello, Dilson G. C.</mods:namePart>
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<mods:namePart id="41CEB5886C231599F36960DB9258721C">Irmler, Ulrich</mods:namePart>
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<mods:name id="C7EB2A5FBCE90BD7648D245437CA3CB9" type="personal">
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<mods:namePart id="C844CA09110243A22808858B558185C9">Caron, Edilson</mods:namePart>
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<mods:title id="FFB785F98C1831BFE6332256F8FB2CF9">Zootaxa</mods:title>
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<mods:date id="B9E75EDB79D1E849BF6457B5DE17C557">2020</mods:date>
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<mods:number id="082D4560FFD95A5B96DE4728A6B0F999">2020-03-23</mods:number>
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<mods:number id="9ABBD6BA21E0D2720237E8C88D79BACA">4755</mods:number>
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<mods:identifier id="9EA0F0D16B98907E7F2ACD5E7B3C551D" type="DOI">10.11646/zootaxa.4755.1.6</mods:identifier>
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<treatment id="03C887D5FF8FFF8D83C6F9DFC3A9FE50" ID-DOI="http://doi.org/10.5281/zenodo.3728591" ID-GBIF-Taxon="162696931" ID-Zenodo-Dep="3728591" LSID="urn:lsid:plazi:treatment:03C887D5FF8FFF8D83C6F9DFC3A9FE50" httpUri="http://treatment.plazi.org/id/03C887D5FF8FFF8D83C6F9DFC3A9FE50" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
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<subSubSection id="C37B6548FF8FFF8C83C6F9DFC495F991" box="[151,673,1629,1656]" pageId="2" pageNumber="141" type="nomenclature">
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<paragraph id="8BDE36C3FF8FFF8C83C6F9DFC495F991" blockId="2.[151,673,1629,1656]" box="[151,673,1629,1656]" pageId="2" pageNumber="141">
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<heading id="D09681AFFF8FFF8C83C6F9DFC495F991" bold="true" box="[151,673,1629,1656]" fontSize="11" level="1" pageId="2" pageNumber="141" reason="1">
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<emphasis id="B915EAD1FF8FFF8C83C6F9DFC495F991" bold="true" box="[151,673,1629,1656]" pageId="2" pageNumber="141">
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Key to species of
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<taxonomicName id="4C614D40FF8FFF8C823FF9DFC495F991" ID-CoL="9NM4T" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1933" box="[366,673,1629,1656]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD1FF8FFF8C823FF9DFC427F99E" bold="true" box="[366,531,1629,1656]" italics="true" pageId="2" pageNumber="141">Pseudastenus</emphasis>
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Bernhauer
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C37B6548FF8FFF8D83C6F920C3A9FE50" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141" type="key">
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<keyStep id="30952E66FF8FFF8C83C6F920C3A8F8E9" pageId="2" pageNumber="141">
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<paragraph id="8BDE36C3FF8FFF8C83C6F920C3A8F932" blockId="2.[151,1437,1697,2042]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F920C3A8F932" pageId="2" pageNumber="141">1. Elytra shorter than pronotum, 0.9 times as long as pronotum, sides divergent to posterior margin, widest at posterior margin. ................................................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8FFF8C83C6F95FC3A8F8E9" blockId="2.[151,1437,1697,2042]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F95FC3A8F8E9" pageId="2" pageNumber="141">- Elytra longer than pronotum, ranging between 1.1 and 1.4 times as long as pronotum, sides slightly curved, widest in front of posterior margin...................................................................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30952E66FF8FFF8C83C6F895C3A8F862" pageId="2" pageNumber="141">
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<paragraph id="8BDE36C3FF8FFF8C83C6F895C3A8F8AF" blockId="2.[151,1437,1697,2042]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F895C3A8F8AF" pageId="2" pageNumber="141">2. Eyes reduced, approximately only 0.20 times as long as total length of head; postocular length more the two times longer than eyes; elytra unicolored or not conspicuously bicolored........................................................ 3</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8FFF8C83C6F8D0C3A8F862" blockId="2.[151,1437,1697,2042]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F8D0C3A8F862" pageId="2" pageNumber="141">
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- Eyes less reduced, approximately 0.30 times as long as total length of head; postocular length only 1.6 times as long as eyes; elytra conspicuously bicolored, with distinct black anterior half.........................................
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<taxonomicName id="4C614D40FF8FFF8C8666F8F2C3A8F862" ID-CoL="9QFT9" authorityName="Bernhauer" authorityYear="1933" box="[1335,1436,1903,1925]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="barretoi">
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<emphasis id="B915EAD1FF8FFF8C8666F8F2C3A8F862" box="[1335,1436,1903,1925]" italics="true" pageId="2" pageNumber="141">P. barretoi</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30952E66FF8FFF8C83C6F80EC3A9F83A" pageId="2" pageNumber="141">
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<paragraph id="8BDE36C3FF8FFF8C83C6F80EC3A8F859" blockId="2.[151,1437,1697,2042]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F80EC3A8F859" pageId="2" pageNumber="141">
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3. Eyes slightly larger; postocular sides approximately parallel; elytra slightly bicolored, anterior half of elytra indistinctly darker than posterior half..........................................................................
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<taxonomicName id="4C614D40FF8FFF8C8657F828C3A8F859" ID-CoL="9V8WZ" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1286,1436,1962,1983]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="ribeirocostae">
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<emphasis id="B915EAD1FF8FFF8C8657F828C3A8F859" box="[1286,1436,1962,1983]" italics="true" pageId="2" pageNumber="141">P. ribeirocostae</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8FFF8C83C6F845C3A9F83A" blockId="2.[151,1437,1697,2042]" box="[151,1437,1991,2013]" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8C83C6F845C3A9F83A" box="[151,1437,1991,2013]" pageId="2" pageNumber="141">
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- Eyes slightly smaller; postocular sides divergent; elytra unicolored yellow................................
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<taxonomicName id="4C614D40FF8FFF8C867BF84AC3A9F83A" ID-CoL="9V8WW" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1322,1437,1991,2013]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="schubarti">
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<emphasis id="B915EAD1FF8FFF8C867BF84AC3A9F83A" box="[1322,1437,1991,2013]" italics="true" pageId="2" pageNumber="141">P. schubarti</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<paragraph id="8BDE36C3FF8FFF8D83C6F866C3A8FF2B" blockId="2.[151,1437,1697,2042]" lastBlockId="3.[151,1437,153,439]" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
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<keyStep id="30952E66FF8FFF8D83C6F866C3A8FF2B" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
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<keyLead id="309095F6FF8FFF8D83C6F866C3A8FF2B" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
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4. Eyes large, 0.33 times as long as total length of head; postocular length equal to eye length...................
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<taxonomicName id="4C614D40FF8FFF8C8662F867C3A8F81F" ID-CoL="9V8WX" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1331,1436,2020,2042]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="oculatus">
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<emphasis id="B915EAD1FF8FFF8C8662F867C3A8F81F" box="[1331,1436,2020,2042]" italics="true" pageId="2" pageNumber="141">P. oculatus</emphasis>
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</taxonomicName>
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- Eyes smaller, between 0.24 and 0.31 times as long as total length of head; postocular length distinctly longer than eye length ................................................................................................... 5
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</keyLead>
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</keyStep>
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</paragraph>
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<keyStep id="30952E66FF8EFF8D83C6FF56C3A9FEE1" pageId="3" pageNumber="142">
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<paragraph id="8BDE36C3FF8EFF8D83C6FF56C3A8FF0C" blockId="3.[151,1437,153,439]" box="[151,1436,212,234]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FF56C3A8FF0C" box="[151,1436,212,234]" pageId="3" pageNumber="142">5. Pronotum light brown; elytra with basal half darker than apical half (in some specimens not so conspicuous)............. 6</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8EFF8D83C6FF70C3A9FEE1" blockId="3.[151,1437,153,439]" box="[151,1437,241,263]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FF70C3A9FEE1" box="[151,1437,241,263]" pageId="3" pageNumber="142">
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- Pronotum rust brown to reddish; elytra with basal two-thirds darker than apical onethird...................
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<taxonomicName id="4C614D40FF8EFF8D8642FF70C3A9FEE1" ID-CoL="9V8WV" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1299,1437,242,263]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="ferrugineus">
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<emphasis id="B915EAD1FF8EFF8D8642FF70C316FEE1" box="[1299,1314,242,263]" italics="true" pageId="3" pageNumber="142">P</emphasis>
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.
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<emphasis id="B915EAD1FF8EFF8D867FFF70C3A9FEE1" box="[1326,1437,242,263]" italics="true" pageId="3" pageNumber="142">ferrugineus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30952E66FF8EFF8D83C6FE8DC3A8FE9A" pageId="3" pageNumber="142">
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<paragraph id="8BDE36C3FF8EFF8D83C6FE8DC3A8FEA4" blockId="3.[151,1437,153,439]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FE8DC3A8FEA4" pageId="3" pageNumber="142">6. Larger, at least 3.00 mm long; elytra bicolored, dark brown to black in anterior half and light brown in posterior half; aedeagus with apical sclerites of internal sac not elongate............................................................. 7</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8EFF8D83C6FEC8C3A8FE9A" blockId="3.[151,1437,153,439]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FEC8C3A8FE9A" pageId="3" pageNumber="142">
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- Smaller,
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<quantity id="4C999B26FF8EFF8D8269FEC8C7B9FEB9" box="[312,397,330,351]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="142" unit="mm" value="2.2">2.20 mm</quantity>
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long; elytra unicolored light brown; aedeagus with apical sclerites of internal sac elongate and straight............................................................................................
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<taxonomicName id="4C614D40FF8EFF8D8642FEE5C3A8FE9A" ID-CoL="9V8X2" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1299,1436,359,380]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="amazonicus">
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<emphasis id="B915EAD1FF8EFF8D8642FEE5C3A8FE9A" box="[1299,1436,359,380]" italics="true" pageId="3" pageNumber="142">P. amazonicus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30952E66FF8EFF8D83C6FE07C3A9FE50" pageId="3" pageNumber="142">
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<paragraph id="8BDE36C3FF8EFF8D83C6FE07C3A9FE7F" blockId="3.[151,1437,153,439]" box="[151,1437,388,410]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FE07C3A9FE7F" box="[151,1437,388,410]" pageId="3" pageNumber="142">
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7. Head light brown; median lobe of aedeagus with apical sclerites entirely narrow and S-shaped (
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<figureCitation id="135A2A46FF8EFF8D8727FE06C2DEFE7C" box="[1142,1258,388,410]" captionStart="FIGURES 46–47" captionStartId="12.[151,264,1161,1186]" captionTargetBox="[247,1380,261,1071]" captionTargetId="figure@12.[151,1436,181,1135]" captionTargetPageId="12" captionText="FIGURES 46–47. Aedeagus of Pseudastenus plaumanni. 46. Dissected from the lectotype (FMNH), dorsal view. 47. Dissected from a specimen collected in the type locality, dorsal and lateral view, respectively. Details: a, basal sclerite; b, apical sclerite; c, basal sclerite of the ejaculatory duct; d, apical sclerite of the ejaculatory duct. Scale bars: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.3724305" httpUri="https://zenodo.org/record/3724305/files/figure.png" pageId="3" pageNumber="142">Figs. 46–47</figureCitation>
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)....
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<taxonomicName id="4C614D40FF8EFF8D864FFE07C3A9FE7F" ID-CoL="9RCDB" authorityName="Bierig" authorityYear="1939" box="[1310,1437,388,410]" class="Insecta" family="Staphylinidae" genus="Xenaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="plaumanni">
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<emphasis id="B915EAD1FF8EFF8D864FFE07C3A9FE7F" box="[1310,1437,388,410]" italics="true" pageId="3" pageNumber="142">P. plaumanni</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BDE36C3FF8EFF8D83C6FE20C3A9FE50" blockId="3.[151,1437,153,439]" box="[151,1437,417,439]" pageId="3" pageNumber="142">
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<keyLead id="309095F6FF8EFF8D83C6FE20C3A9FE50" box="[151,1437,417,439]" pageId="3" pageNumber="142">
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- Head dark brown; median lobe of aedeagus with apical sclerites broad on the basal half and not S-shaped (
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<figureCitation id="135A2A46FF8EFF8D87B0FE23C313FE51" box="[1249,1319,417,439]" captionStart="FIGURES 48–54" captionStartId="13.[151,264,1895,1920]" captionTargetBox="[232,1346,220,1805]" captionTargetId="figure@13.[164,1424,181,1869]" captionTargetPageId="13" captionText="FIGURES 48–54. Aedeagus of Pseudastenus, dorsal and lateral view, respectively, figures 51–54 including microscopic detail of basal slerites. 48. P. ribeirocostae sp. nov.; 49. P. ferrugineus sp. nov.; 50. P. latus sp. nov.; 51. P. barretoi; 52. P. oculatus sp. nov.; 53. P. amazonicus sp. nov.; 54. P. schubarti sp. nov. Scale bar: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3724309" httpUri="https://zenodo.org/record/3724309/files/figure.png" pageId="3" pageNumber="142">Fig. 50</figureCitation>
|
||
)...
|
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<taxonomicName id="4C614D40FF8EFF8D8602FE20C3A9FE50" ID-CoL="9V8WT" authorityName="Busanello & Irmler & Caron" authorityYear="2020" box="[1363,1437,417,439]" class="Insecta" family="Staphylinidae" genus="Pseudastenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="B915EAD1FF8EFF8D8602FE20C356FE51" box="[1363,1378,418,439]" italics="true" pageId="3" pageNumber="142">P</emphasis>
|
||
.
|
||
<emphasis id="B915EAD1FF8EFF8D863FFE23C3A9FE50" box="[1390,1437,417,438]" italics="true" pageId="3" pageNumber="142">latus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |