347 lines
43 KiB
XML
347 lines
43 KiB
XML
<document id="A906972F5D4CFACBA423320B7E9B65F3" ID-DOI="10.11646/zootaxa.3741.1.2" ID-GBIF-Dataset="6a6ba78e-a9c5-4d11-997a-3d06e5e31520" ID-ISSN="1175-5326" ID-Zenodo-Dep="223773" ID-ZooBank="BF5EA9B8-C6F4-448A-BEF9-1976AB4EC308" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460702326177" checkinUser="plazi" docAuthor="Lazányi, Eszter & Vagalinski, Boyan" docDate="2013" docId="03C887D3FFFBFFB3FF349FC5FC5DAEFA" docLanguage="en" docName="zt03741p100.pdf" docOrigin="Zootaxa 3741 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum sensu subsp. stricto Verhoeff 1894, sensu stricto" docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="FFF1FFABFFF9FFB7FFA39926FFF2AD11" masterDocTitle="Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)" masterLastPageNumber="100" masterPageNumber="55" pageNumber="57" updateTime="1698356536038" updateUser="plazi">
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<mods:title id="941B0CEAC5C5D4DE177BA69BC00AC403">Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="02F042CBBE5AE0A64BF29627600751BC">Lazányi, Eszter</mods:namePart>
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<mods:name id="693F6AC697DA151F44451ADBAB7F6FDB" type="personal">
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<mods:roleTerm id="27F66287351E73A47220BEE22CF13CD2">Author</mods:roleTerm>
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<mods:namePart id="2DEE9FC3FF0A35E4C9D669F8E392C13F">Vagalinski, Boyan</mods:namePart>
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<mods:typeOfResource id="7C45CBABCCC1B0095489E394123BBEBD">text</mods:typeOfResource>
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<mods:title id="ABD25AE6F660704780FDF83CAFB59958">Zootaxa</mods:title>
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<mods:part id="B2AAD167E95C6E8FD5052539016DCDEF">
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<mods:date id="9B197308D2786D8C310A86D222B03D62">2013</mods:date>
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<mods:number id="129B2ED9F1393F1A1FF1DD68F9BD0481">3741</mods:number>
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<mods:identifier id="9357A58F0CAE4DBA25A93CE2B43732E5" type="DOI">10.11646/zootaxa.3741.1.2</mods:identifier>
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<mods:identifier id="3AC34EB2E2C01C4AAE54037A86187579" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="323CA1517060DEF4B712F3EF765EC907" type="Zenodo-Dep">223773</mods:identifier>
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<treatment id="03C887D3FFFBFFB3FF349FC5FC5DAEFA" ID-DOI="http://doi.org/10.5281/zenodo.6151701" ID-GBIF-Taxon="119583510" ID-Zenodo-Dep="6151701" LSID="urn:lsid:plazi:treatment:03C887D3FFFBFFB3FF349FC5FC5DAEFA" httpUri="http://treatment.plazi.org/id/03C887D3FFFBFFB3FF349FC5FC5DAEFA" lastPageId="4" lastPageNumber="59" pageId="2" pageNumber="57">
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<subSubSection id="C37B654EFFFBFFB5FF349FC5FDD8AA6C" pageId="2" pageNumber="57" type="nomenclature">
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<paragraph id="8BDE36C5FFFBFFB5FF349FC5FCEBABEC" blockId="2.[151,793,1763,1789]" box="[151,793,1763,1789]" pageId="2" pageNumber="57">
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<heading id="D09681A9FFFBFFB5FF349FC5FCEBABEC" bold="true" box="[151,793,1763,1789]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
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<emphasis id="B915EAD7FFFBFFB5FF349FC5FCEBABEC" bold="true" box="[151,793,1763,1789]" pageId="2" pageNumber="57">
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Subgenus
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<taxonomicName id="4C614D46FFFBFFB5FEB49FC5FCEBABEC" authority="Verhoeff, 1894" authorityName="Verhoeff" authorityYear="1894" box="[279,793,1763,1789]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="sensu" subSpecies="stricto">
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<emphasis id="B915EAD7FFFBFFB5FEB49FC5FDAEABEC" bold="true" box="[279,604,1763,1789]" italics="true" pageId="2" pageNumber="57">
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Megaphyllum
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<taxonomicNameLabel id="A22657ACFFFBFFB5FE659FC5FDAEABEC" box="[454,604,1763,1789]" pageId="2" pageNumber="57" sensu="stricto">sensu stricto</taxonomicNameLabel>
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</emphasis>
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Verhoeff, 1894
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDE36C5FFFBFFB5FF349E0CFDD8AA6C" blockId="2.[151,997,1834,1947]" pageId="2" pageNumber="57">
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<taxonomicName id="4C614D46FFFBFFB5FF349E0CFE10AA51" authority="Chromatoilulus" authorityName="Chromatoilulus" box="[151,482,1834,1856]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFBFFB5FF349E0CFE10AA51" box="[151,482,1834,1856]" italics="true" pageId="2" pageNumber="57">Chromatoiulus (Chromatoilulus)</emphasis>
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</taxonomicName>
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: Lohmander 1936: 104; Attems 1940: 306.
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<taxonomicName id="4C614D46FFFBFFB5FF349E6EFE4DAA4F" authority="(Chromatoiulus)" baseAuthorityName="Chromatoiulus" box="[151,447,1864,1886]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFBFFB5FF349E6EFE4DAA4F" box="[151,447,1864,1886]" italics="true" pageId="2" pageNumber="57">Brachyiulus (Chromatoiulus)</emphasis>
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</taxonomicName>
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: Verhoeff 1896a: 219; 1896b: 211; Attems 1899: 328.
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<taxonomicName id="4C614D46FFFBFFB5FF349E41FE50AA6C" box="[151,418,1895,1917]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFBFFB5FF349E41FE50AA6C" box="[151,418,1895,1917]" italics="true" pageId="2" pageNumber="57">Brachyiulus (Brachyiulus)</emphasis>
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</taxonomicName>
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: 1930: 1668.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFFBFFB4FF349EA3FCDEAC0D" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="description">
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<paragraph id="8BDE36C5FFFBFFB5FF349EA3FD6EAA8A" blockId="2.[151,997,1834,1947]" box="[151,668,1925,1947]" pageId="2" pageNumber="57">
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<taxonomicName id="4C614D46FFFBFFB5FF349EA3FE3DAA8A" authority="(Chromatoiulus)" baseAuthorityName="Chromatoiulus" box="[151,463,1925,1947]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFBFFB5FF349EA3FE3DAA8A" box="[151,463,1925,1947]" italics="true" pageId="2" pageNumber="57">Megaphyllum (Chromatoiulus)</emphasis>
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</taxonomicName>
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: Verhoeff 1895:240
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</paragraph>
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<paragraph id="8BDE36C5FFFBFFB4FF349EEFFB2DADE9" blockId="2.[151,1436,1992,2017]" lastBlockId="3.[151,1437,151,2018]" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
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A rough definition of the subgenus
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<emphasis id="B915EAD7FFFBFFB5FD859EEEFD0EAAF1" box="[550,764,1992,2016]" italics="true" pageId="2" pageNumber="57">
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<taxonomicName id="4C614D46FFFBFFB5FD859EEEFD32AAF1" box="[550,704,1992,2016]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus" sensu="stricto">Megaphyllum</taxonomicName>
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<taxonomicNameLabel id="A22657ACFFFBFFB5FD6B9EEFFD0EAAF1" box="[712,764,1993,2016]" pageId="2" pageNumber="57" sensu="stricto">s.str.</taxonomicNameLabel>
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</emphasis>
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was given by Lazányi
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<emphasis id="B915EAD7FFFBFFB5FC5C9EEFFBCAAAF1" box="[1023,1080,1992,2016]" italics="true" pageId="2" pageNumber="57">et al.</emphasis>
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(2012) on the basis of previous publications (Verhoeff 1894d, Attems 1927); prior to that it had never been defined under the subgenus name
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<taxonomicName id="4C614D46FFFAFFB4FF34999AFEC3ADC5" box="[151,305,188,212]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFAFFB4FF34999AFEC3ADC5" box="[151,305,188,212]" italics="true" pageId="3" pageNumber="58">Megaphyllum</emphasis>
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</taxonomicName>
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. Verhoeff (1896a, 1896b) focused on gonopod structure, occasionally supplemented with other characters of the head, eye, repugnatorial glands, and form and setation of telson (Verhoeff 1930).
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</paragraph>
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<paragraph id="8BDE36C5FFFAFFB4FF649823FCDEAC0D" blockId="3.[151,1437,151,2018]" box="[199,812,260,285]" pageId="3" pageNumber="58">
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<typeStatus id="54DA8867FFFAFFB4FF649823FF0DAC0C" box="[199,255,261,285]" pageId="3" pageNumber="58">Type</typeStatus>
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species:
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<taxonomicName id="4C614D46FFFAFFB4FEC69822FCDBAC0D" authority="Verhoeff, 1894" authorityName="Verhoeff" authorityYear="1894" box="[357,809,260,285]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="projectum">
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<emphasis id="B915EAD7FFFAFFB4FEC69822FD84AC0D" box="[357,630,260,284]" italics="true" pageId="3" pageNumber="58">Megaphyllum projectum</emphasis>
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Verhoeff, 1894
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFFAFFB4FF649801FC8FACE4" pageId="3" pageNumber="58" type="diagnosis">
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<paragraph id="8BDE36C5FFFAFFB4FF649801FC8FACE4" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
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<emphasis id="B915EAD7FFFAFFB4FF649801FEB3AC51" bold="true" box="[199,321,295,320]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
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A subgenus of the genus
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<taxonomicName id="4C614D46FFFAFFB4FD21980EFCEEAC51" box="[642,796,296,320]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B915EAD7FFFAFFB4FD21980EFCEEAC51" box="[642,796,296,320]" italics="true" pageId="3" pageNumber="58">Megaphyllum</emphasis>
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</taxonomicName>
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with considerably broader promere than opisthomere, tightly “embracing” the latter antero-laterally, with an oblique riffle on its caudal surface. Opisthomere always with a well-developed posterior process, joining the main opisthomere body at an acute angle; usually with an anterior process, this being vestigial or fully absent in some species; solenomere with two distinct processes—posterior and anterior, the latter positioned at the end of the flagellum-conducting lamella. Vulvae subcylindrical, with an oval, mostly apically positioned opening; operculum shorter than bursa.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B654EFFFAFFB3FF6498D9FA83AC99" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="description">
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<paragraph id="8BDE36C5FFFAFFB4FF6498D9FDBBAF95" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
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<emphasis id="B915EAD7FFFAFFB4FF6498D9FEA3AF09" bold="true" box="[199,337,511,536]" pageId="3" pageNumber="58">Description</emphasis>
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: Body length and height: males:
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<quantity id="4C999B20FFFAFFB4FD7598D9FC92AF09" box="[726,864,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.645" metricValueMax="5.39" metricValueMin="1.9" pageId="3" pageNumber="58" unit="mm" value="36.45" valueMax="53.9" valueMin="19.0">19–53.9mm</quantity>
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,
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<quantity id="4C999B20FFFAFFB4FCCD98D9FC03AF09" box="[878,1009,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.5" metricValueMin="1.4" pageId="3" pageNumber="58" unit="mm" value="2.45" valueMax="3.5" valueMin="1.4">1.4–3.5mm</quantity>
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; females:
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<quantity id="4C999B20FFFAFFB4FBCB98D9FAF5AF09" box="[1128,1287,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" metricValueMax="5.46" metricValueMin="2.02" pageId="3" pageNumber="58" unit="mm" value="37.4" valueMax="54.6" valueMin="20.2">20.2–54.6mm</quantity>
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,
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<quantity id="4C999B20FFFAFFB4FAB69B26FA6AAF09" box="[1301,1432,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="4.7" metricValueMin="2.2" pageId="3" pageNumber="58" unit="mm" value="3.45" valueMax="4.7" valueMin="2.2">2.2–4.7mm</quantity>
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; body ring number: 47–52 (+1–2 apodous+T). Females usually longer, higher and more brightly colored than males. Developmental stadia (identified after Enghoff
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<emphasis id="B915EAD7FFFAFFB4FD189B6FFD05AF71" box="[699,759,584,608]" italics="true" pageId="3" pageNumber="58">et al.</emphasis>
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(1993)): adult males: VIII–XII (exceptionally XIII); adult females: IX–XIII (exceptionally XIV).
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</paragraph>
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<paragraph id="8BDE36C5FFFAFFB4FF649BB5FB3CAE06" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
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2 frontal, 4 supralabral and 19–28 labral setae. Ratio of 5th and 4th antennal articles: 5:4=1–1.1 (Fig. 6h). Form of gnathochilarium as on Fig.
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<date id="FFDF1005FFFAFFB4FDA19B90FDEDAFDE" box="[514,543,694,719]" pageId="3" pageNumber="58">6i.</date>
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Metazonital setation considerably dense. Preanal process long and pointed, sometimes curved upwards (Fig. 6j). Anal valves densely setose. Subanal scale trapezoid, never pointed; apically mostly with 2–3 teeth (Fig. 6k). Preanal process blunter, subanal scale more rounded in females.
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</paragraph>
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<paragraph id="8BDE36C5FFFAFFB4FF649A04FD11A990" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
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Males: Gnathal lobe well-developed. First leg-pair in the form of a simple hook (Fig. 6l). Penis flattened from antero-posteriad, apically with very short lobes and with two small, wide, (usually) rounded lamellae running parallel (
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<emphasis id="B915EAD7FFFAFFB4FF5F9A4CFEF9AE92" bold="true" box="[252,267,874,899]" italics="true" pageId="3" pageNumber="58">ll</emphasis>
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for left and
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<emphasis id="B915EAD7FFFAFFB4FE399A4CFE5FAE92" bold="true" box="[410,429,874,899]" italics="true" pageId="3" pageNumber="58">rl</emphasis>
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for right lamella) and pointing posteriad in some species—these lamellae sometimes not visible. Penis occasionally bent along its longitudinal axis (e.g. in
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<taxonomicName id="4C614D46FFFAFFB4FC2E9AB6FAF7AEB6" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[909,1285,910,935]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="montivagum">
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<emphasis id="B915EAD7FFFAFFB4FC2E9AB6FBCCAEB6" box="[909,1086,912,935]" italics="true" pageId="3" pageNumber="58">M. montivagum</emphasis>
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(Verhoeff, 1901)
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</taxonomicName>
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, see
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<figureCitation id="135A2A40FFFAFFB4FA9C9AA8FF02AEDA" captionStart="FIGURES 11 a – g" captionStartId="20.[151,264,1683,1705]" captionTargetBox="[257,1326,640,1661]" captionTargetId="figure@20.[252,1334,637,1662]" captionTargetPageId="20" captionText="FIGURES 11 a – g. Megaphyllum monticola. Males from Korakas Mts, Greece, ZMB 1300: (11 a) gonopods, posterior view and (11 b) right gonopods, mesal view and (11 c) left promere, meso-posterior view. Males from Ostrovo, Greece, Inv. No. 8019: (11 d) Right opisthomere, lateral view, penis, (11 e) posterior and (11 f) right lateral views, (11 f *) another, bent penis, left lateral view. (11 g) Left vulva, posterior view, female from Korakas Mts, Greece (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223783/files/figure.png" pageId="3" pageNumber="58">Figs 11f and 11f</figureCitation>
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*). Second pair of legs with two pads on postfemur and tibia (
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<emphasis id="B915EAD7FFFAFFB4FC089A94FC30AEDA" bold="true" box="[939,962,946,971]" italics="true" pageId="3" pageNumber="58">pf</emphasis>
|
||
and
|
||
<emphasis id="B915EAD7FFFAFFB4FC599A95FBF0AEDB" bold="true" box="[1018,1026,947,970]" italics="true" pageId="3" pageNumber="58">t</emphasis>
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, respectively), coxa simple, without hump (Fig. 6n). All subsequent legs with three pads on femur (
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<emphasis id="B915EAD7FFFAFFB4FCC99AF0FC81AEFE" bold="true" box="[874,883,982,1007]" italics="true" pageId="3" pageNumber="58">f</emphasis>
|
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), postfemur (
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<emphasis id="B915EAD7FFFAFFB4FBB39AF0FBD5AEFE" bold="true" box="[1040,1063,982,1007]" italics="true" pageId="3" pageNumber="58">pf</emphasis>
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) and tibia (
|
||
<emphasis id="B915EAD7FFFAFFB4FB0E9AF1FB47AEFF" bold="true" box="[1197,1205,983,1006]" italics="true" pageId="3" pageNumber="58">t</emphasis>
|
||
) (Fig.
|
||
<quantity id="4C999B20FFFAFFB4FAA59AF0FADFAEFE" box="[1286,1325,982,1007]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="58" unit="m" value="6.0">6m</quantity>
|
||
). Pads on postfemur and tibia sometimes rippled (as if additional pads would exist on the same article) (see
|
||
<emphasis id="B915EAD7FFFAFFB4FB759ADCFB1FA902" bold="true" box="[1238,1261,1018,1043]" italics="true" pageId="3" pageNumber="58">pf</emphasis>
|
||
on Fig.
|
||
<quantity id="4C999B20FFFAFFB4FAE99ADCFA83A902" box="[1354,1393,1018,1043]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="58" unit="m" value="6.0">6m</quantity>
|
||
), or with vestigial proximal parts (
|
||
<emphasis id="B915EAD7FFFAFFB4FE529D06FDE6A926" box="[497,532,1056,1079]" italics="true" pageId="3" pageNumber="58">i.e.</emphasis>
|
||
only the distal part visible). Tarsus of midbody legs shorter than tibia: (tarsus/ tibia)=0.65–0.9, although an unusual individual with ratio 1.15 was also found. 7th pleurotergum protruding like a simple shovel on the border of pro- and metazonum.
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFAFFB4FF649DABFCCAA84B" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
|
||
Male gonopods: Promere (
|
||
<emphasis id="B915EAD7FFFAFFB4FE5E9DA8FDFCA9B4" bold="true" box="[509,526,1166,1189]" italics="true" pageId="3" pageNumber="58">P</emphasis>
|
||
) very large, latero-anteriorly embracing the opisthomere. Connection between promeres due only to basal, transversal muscle fibres, and to a basal chitinous node between their mesal posterior margins,
|
||
<emphasis id="B915EAD7FFFAFFB4FEA29DF0FEEFA9FC" box="[257,285,1238,1261]" italics="true" pageId="3" pageNumber="58">i.e</emphasis>
|
||
. no additional connecting structure present. Promere bases relatively (compared to other congeners) well-developed, roundly protruding laterally. Posterior surface of the promere with an oblique ridge (
|
||
<emphasis id="B915EAD7FFFAFFB4FAB29DDCFAEEA800" bold="true" box="[1297,1308,1274,1297]" italics="true" pageId="3" pageNumber="58">r</emphasis>
|
||
) forming a groove (
|
||
<emphasis id="B915EAD7FFFAFFB4FF569C38FEF1A824" bold="true" box="[245,259,1310,1333]" italics="true" pageId="3" pageNumber="58">g</emphasis>
|
||
); and in some species an apical pit (“Fenestra”,
|
||
<emphasis id="B915EAD7FFFAFFB4FC859C38FCBBA824" box="[806,841,1310,1333]" italics="true" pageId="3" pageNumber="58">i.e.</emphasis>
|
||
window; see
|
||
<emphasis id="B915EAD7FFFAFFB4FC469C38FBFFA824" box="[997,1037,1310,1333]" italics="true" pageId="3" pageNumber="58">e.g.</emphasis>
|
||
Verhoeff 1897b)—as an imprint of the opisthomere’s anterior process. Flagellum long and thin.
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFAFFB4FF649C43FF1FAA3F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
|
||
All species groups with a basic arrangement of opisthomere parts (
|
||
<figureCitation id="135A2A40FFFAFFB4FC6B9C43FBBBA86F" box="[968,1097,1381,1406]" captionStart="FIGURES 1 a – e" captionStartId="4.[151,264,1450,1472]" captionTargetBox="[200,1386,1061,1424]" captionTargetId="figure@4.[197,1389,1058,1426]" captionTargetPageId="4" captionText="FIGURES 1 a – e. Schematic opisthomere drawings (right opisthomeres, lateral view). (1 a) Generalized scheme for all Megaphyllum sensu stricto. (1 b) Scheme for the M. projectum species group. (1 c) Scheme for the M. austriacum species group. (1 d) Scheme for the M. transsylvanicum species group. (1 e) Scheme for the M. unilineatum species group. Abbreviations: ah: anterior hump, ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere." httpUri="https://zenodo.org/record/223774/files/figure.png" pageId="3" pageNumber="58">Figs 1b–1e</figureCitation>
|
||
), while shape of opisthomere parts different within the actual species groups. Basically (
|
||
<figureCitation id="135A2A40FFFAFFB4FC969CAFFC7AA8B3" box="[821,904,1417,1442]" captionStart="FIGURES 1 a – e" captionStartId="4.[151,264,1450,1472]" captionTargetBox="[200,1386,1061,1424]" captionTargetId="figure@4.[197,1389,1058,1426]" captionTargetPageId="4" captionText="FIGURES 1 a – e. Schematic opisthomere drawings (right opisthomeres, lateral view). (1 a) Generalized scheme for all Megaphyllum sensu stricto. (1 b) Scheme for the M. projectum species group. (1 c) Scheme for the M. austriacum species group. (1 d) Scheme for the M. transsylvanicum species group. (1 e) Scheme for the M. unilineatum species group. Abbreviations: ah: anterior hump, ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere." httpUri="https://zenodo.org/record/223774/files/figure.png" pageId="3" pageNumber="58">Fig. 1a</figureCitation>
|
||
): opisthomere with a well-developed posterior process (
|
||
<emphasis id="B915EAD7FFFAFFB4FF599C88FEE4A8D4" bold="true" box="[250,278,1454,1477]" italics="true" pageId="3" pageNumber="58">pp</emphasis>
|
||
) in every species group—most likely this was called “
|
||
<emphasis id="B915EAD7FFFAFFB4FCD79C88FC10A8D4" box="[884,994,1454,1477]" italics="true" pageId="3" pageNumber="58">Paracoxit</emphasis>
|
||
” or “
|
||
<emphasis id="B915EAD7FFFAFFB4FB839C8BFB45A8D4" box="[1056,1207,1453,1477]" italics="true" pageId="3" pageNumber="58">Coxalfortsatz</emphasis>
|
||
” (
|
||
<emphasis id="B915EAD7FFFAFFB4FB709C88FB04A8D4" box="[1235,1270,1454,1477]" italics="true" pageId="3" pageNumber="58">i.e.</emphasis>
|
||
coxal process) by Attems (
|
||
<date id="FFDF1005FFFAFFB4FEBF9CF4FE3AA8FB" box="[284,456,1489,1514]" pageId="3" pageNumber="58" value="1927" valueMax="1940">1927 and 1940</date>
|
||
, respectively) and all the other parts of the opisthomere were probably gathered under the name “Telopodit” (Attems 1927: fig. 287.). Opisthomere usually also with an anterior process or anterior ridge in same position (
|
||
<emphasis id="B915EAD7FFFAFFB4FECC9F3CFE79AB20" bold="true" box="[367,395,1562,1585]" italics="true" pageId="3" pageNumber="58">ap</emphasis>
|
||
). Occasionally the apical part of the opisthomere is in the form of a hump around the solenomere, around the end of the flagellum-conducting lamella, posteriorly and/or anteriorly variably developed (
|
||
<emphasis id="B915EAD7FFFAFFB4FF039F47FF4FAB6B" bold="true" box="[160,189,1633,1658]" italics="true" pageId="3" pageNumber="58">ah</emphasis>
|
||
for anterior hump and
|
||
<emphasis id="B915EAD7FFFAFFB4FE1D9F47FE29AB6B" bold="true" box="[446,475,1633,1658]" italics="true" pageId="3" pageNumber="58">ph</emphasis>
|
||
for posterior hump)—probably this hump was called “
|
||
<emphasis id="B915EAD7FFFAFFB4FBE39F47FB63AB68" box="[1088,1169,1633,1657]" italics="true" pageId="3" pageNumber="58">Polster</emphasis>
|
||
” (
|
||
<emphasis id="B915EAD7FFFAFFB4FB0E9F44FB22AB68" box="[1197,1232,1634,1657]" italics="true" pageId="3" pageNumber="58">i.e.</emphasis>
|
||
pillow or pad, see
|
||
<emphasis id="B915EAD7FFFAFFB4FF349FA0FF4DAB8C" box="[151,191,1670,1693]" italics="true" pageId="3" pageNumber="58">e.g.</emphasis>
|
||
Attems 1927, 1940). Solenomere always with an anterior and a posterior process (
|
||
<emphasis id="B915EAD7FFFAFFB4FB2A9FA0FB42AB8C" bold="true" box="[1161,1200,1670,1693]" italics="true" pageId="3" pageNumber="58">asp</emphasis>
|
||
and
|
||
<emphasis id="B915EAD7FFFAFFB4FB4E9FA0FAE6AB8C" bold="true" box="[1261,1300,1670,1693]" italics="true" pageId="3" pageNumber="58">psp</emphasis>
|
||
) displaying several shapes and a great variation in length within species groups and species; sometimes even intraspecifically. Anterior solenomere process tubular (see
|
||
<figureCitation id="135A2A40FFFAFFB4FDC99FEBFD1BABF7" box="[618,745,1741,1766]" captionStart-0="FIGURES 9 a – g" captionStart-1="FIGURES 10 a – g" captionStart-2="FIGURES 11 a – g" captionStartId-0="18.[151,264,1626,1648]" captionStartId-1="19.[151,264,1833,1855]" captionStartId-2="20.[151,264,1683,1705]" captionTargetBox-0="[240,1350,593,1604]" captionTargetBox-1="[178,1409,801,1810]" captionTargetBox-2="[257,1326,640,1661]" captionTargetId-0="figure@18.[232,1355,591,1604]" captionTargetId-1="figure@19.[174,1413,797,1811]" captionTargetId-2="figure@20.[252,1334,637,1662]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURES 9 a – g. Megaphyllum dentatum males and a female from Ljuboten, Serbia, Inv. No. 8018: (9 a) gonopods, posterior view; (9 b) right gonopods, mesal view; (9 c) right promere, meso-posterior view; (9 d) right opisthomere, lateral view; penis, (9 e) posterior and (9 f) left lateral views; (9 g) right vulva, anterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, ph: posterior hump, pp: posterior process, pr: protrusion, psp: posterior process of the solenomere, sl: serrated lobe, P: promere. Scale bars: 0.2 mm." captionText-1="FIGURES 10 a – g. Megaphyllum glossulifer males and a female from Rhodopi Mts, Bulgaria: (10 a) gonopods, posterior view; (10 b) left gonopods, mesal view; (10 c) left promere, meso-posterior view; (10 d) left opisthomere, lateral view; penis, (10 e) posterior and (10 f) left lateral views; (10 g) left vulva, meso-posterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." captionText-2="FIGURES 11 a – g. Megaphyllum monticola. Males from Korakas Mts, Greece, ZMB 1300: (11 a) gonopods, posterior view and (11 b) right gonopods, mesal view and (11 c) left promere, meso-posterior view. Males from Ostrovo, Greece, Inv. No. 8019: (11 d) Right opisthomere, lateral view, penis, (11 e) posterior and (11 f) right lateral views, (11 f *) another, bent penis, left lateral view. (11 g) Left vulva, posterior view, female from Korakas Mts, Greece (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri-0="https://zenodo.org/record/223781/files/figure.png" httpUri-1="https://zenodo.org/record/223782/files/figure.png" httpUri-2="https://zenodo.org/record/223783/files/figure.png" pageId="3" pageNumber="58">Figs 9–11d</figureCitation>
|
||
,
|
||
<figureCitation id="135A2A40FFFAFFB4FD579FEBFCEEABF7" box="[756,796,1741,1766]" captionStart="FIGURES 15 a – g" captionStartId="28.[151,264,1408,1430]" captionTargetBox="[198,1393,271,1384]" captionTargetId="figure@28.[190,1397,265,1386]" captionTargetPageId="28" captionText="FIGURES 15 a – g. Megaphyllum hercules. Male from Kechrokambos, Lekanis Mts, Greece: (15 a) gonopods, posterior view; (15 b) right gonopods, mesal view; (15 c) left promere, mesal view; (15 d) left opisthomere, mesal view. Male from Demir Kapija, Republic of Macedonia: penis, (15 e) posterior and (15 f) right lateral views. (15 g) Female from the Rodopi Mts, Greece: left vulva, antero-mesal view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223787/files/figure.png" pageId="3" pageNumber="58">15d</figureCitation>
|
||
)—this may be the sperm channel,
|
||
<emphasis id="B915EAD7FFFAFFB4FB059FE8FB3BABF4" box="[1190,1225,1742,1765]" italics="true" pageId="3" pageNumber="58">i.e.</emphasis>
|
||
“
|
||
<emphasis id="B915EAD7FFFAFFB4FB7F9FE8FA9FABF4" box="[1244,1389,1742,1765]" italics="true" pageId="3" pageNumber="58">Spermarinne</emphasis>
|
||
” by Lohmander (1936, p. 107) while the opening for the flagellum is basal to the anterior solenomere process (see Figs 6p, 8d).
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFAFFB4FF649E1FFA68AAF3" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="58">
|
||
Gonopod complex compact,
|
||
<emphasis id="B915EAD7FFFAFFB4FDA99E1CFDBDAA40" box="[522,591,1850,1873]" italics="true" pageId="3" pageNumber="58">in situ</emphasis>
|
||
not informative. Thus, complete dissection of gonopods needed for species identification,
|
||
<emphasis id="B915EAD7FFFAFFB4FEE29E78FE9BAA64" box="[321,361,1886,1909]" italics="true" pageId="3" pageNumber="58">e.g.</emphasis>
|
||
at least separating right from left gonopods, sometimes also promere from opisthomere. Connection between promere–opisthomere stronger than between promere–promere or opisthomere–opisthomere. Opisthomere lying in the groove (
|
||
<emphasis id="B915EAD7FFFAFFB4FDB99E80FDDAAAAC" bold="true" box="[538,552,1958,1981]" italics="true" pageId="3" pageNumber="58">g</emphasis>
|
||
) of the promere. Solenomere sometimes exceeding the promere apically (
|
||
<emphasis id="B915EAD7FFFAFFB4FAD79E80FEB5AAF0" italics="true" pageId="3" pageNumber="58">
|
||
e.g.
|
||
<taxonomicName id="4C614D46FFFAFFB4FF349EECFEB5AAF0" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[151,327,1994,2017]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="rosenauense">M. rosenauense</taxonomicName>
|
||
</emphasis>
|
||
(Verhoeff, 1897)). The opisthomere’s posterior process (
|
||
<emphasis id="B915EAD7FFFAFFB4FC669EECFC13AAF0" bold="true" box="[965,993,1994,2017]" italics="true" pageId="3" pageNumber="58">pp</emphasis>
|
||
) never exceeding the promere apically.
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFDFFB3FF6499B1FA83AC99" blockId="4.[151,1436,151,645]" pageId="4" pageNumber="59">
|
||
Females: First two leg-pairs of adult females slightly swollen. Vulvae deeply hidden in vulval sacs, cylindrical and apically rounded, with distinct apical margin. Apical opening short; usually not, or only slightly exceeding the apical margin. Operculum (
|
||
<emphasis id="B915EAD7FFFDFFB3FE7699C6FE03ADE6" bold="true" box="[469,497,224,247]" italics="true" pageId="4" pageNumber="59">op</emphasis>
|
||
) positioned latero-anteriorly, shorter than bursa (see
|
||
<emphasis id="B915EAD7FFFDFFB3FBF999C7FB70ADE9" box="[1114,1154,225,248]" italics="true" pageId="4" pageNumber="59">e.g.</emphasis>
|
||
<figureCitation id="135A2A40FFFDFFB3FB2F99F9FB03ADE9" box="[1164,1265,223,248]" captionStart="FIGURES 20 a – g" captionStartId="35.[151,264,1484,1506]" captionTargetBox="[266,1320,497,1467]" captionTargetId="figure@35.[262,1324,493,1468]" captionTargetPageId="35" captionText="FIGURES 20 a – g. Megaphyllum unilineatum. Male from Sminthi, Xanthi county, Greece: (20 a) gonopods, posterior view; (20 b) right gonopods, mesal view; (20 c) left promere, meso-posterior view. Male from the shore of Lake Vistonida, Lagos, Greece: (20 d) left opisthomere, lateral view. Male from near Sminthi, Kamertsi Mts, Greece: penis, (20 e) posterior and (20 f) right lateral views. Female from Nestos Delta, Kavala county, Greece: (20 g) left vulva, latero-posterior view (setae omitted). Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/223792/files/figure.png" pageId="4" pageNumber="59">
|
||
Fig.
|
||
<quantity id="4C999B20FFFDFFB3FB6099C6FB03ADE9" box="[1219,1265,224,248]" metricMagnitude="-2" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="59" unit="g" value="20.0">20g</quantity>
|
||
</figureCitation>
|
||
). Bursa with a double receptaculum seminis consisting of two ampullae: central ampulla (
|
||
<emphasis id="B915EAD7FFFDFFB3FC599822FBE7AC0A" bold="true" box="[1018,1045,260,283]" italics="true" pageId="4" pageNumber="59">ca</emphasis>
|
||
) subequally wide along its whole length, opening to the surface in the apical opening; posterior ampulla (
|
||
<emphasis id="B915EAD7FFFDFFB3FC65980EFC10AC2E" bold="true" box="[966,994,296,319]" italics="true" pageId="4" pageNumber="59">pa</emphasis>
|
||
) drop-shaped, connected to the central one through a significantly more narrow spiral connecting tube (
|
||
<emphasis id="B915EAD7FFFDFFB3FCC7986AFC8AAC72" bold="true" box="[868,888,332,355]" italics="true" pageId="4" pageNumber="59">ct</emphasis>
|
||
), (except for
|
||
<taxonomicName id="4C614D46FFFDFFB3FBB3986BFB4DAC75" box="[1040,1215,333,356]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="montivagum">
|
||
<emphasis id="B915EAD7FFFDFFB3FBB3986BFB4DAC75" box="[1040,1215,333,356]" italics="true" pageId="4" pageNumber="59">M. montivagum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C614D46FFFDFFB3FB57986BFEA1AC99" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="carniolense">
|
||
<emphasis id="B915EAD7FFFDFFB3FB57986BFA6EAC75" box="[1268,1436,332,356]" italics="true" pageId="4" pageNumber="59">M. carniolense</emphasis>
|
||
(Verhoeff, 1897)
|
||
</taxonomicName>
|
||
, with gradually widening connecting tube into the posterior ampulla, without defined border).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B654EFFFDFFB3FF6498B2FC5DAEFA" pageId="4" pageNumber="59" type="discussion">
|
||
<paragraph id="8BDE36C5FFFDFFB3FF6498B2FB7DAF95" blockId="4.[151,1436,151,645]" pageId="4" pageNumber="59">
|
||
<emphasis id="B915EAD7FFFDFFB3FF6498B2FEC9ACBC" bold="true" box="[199,315,404,429]" pageId="4" pageNumber="59">Remarks.</emphasis>
|
||
True sexual dimorphism in body coloration is detected in three species:
|
||
<taxonomicName id="4C614D46FFFDFFB3FB3398B3FF20ACC1" authority="Verhoeff, 1894" authorityName="Verhoeff" authorityYear="1894" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="projectum">
|
||
<emphasis id="B915EAD7FFFDFFB3FB3398B3FADBACBD" box="[1168,1321,405,428]" italics="true" pageId="4" pageNumber="59">M. projectum</emphasis>
|
||
Verhoeff, 1894
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C614D46FFFDFFB3FF7D989FFDAAACC1" authority="(Verhoeff 1897a)" baseAuthorityName="Verhoeff" baseAuthorityYear="1897" box="[222,600,439,464]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="rosenauense">
|
||
<emphasis id="B915EAD7FFFDFFB3FF7D989FFE7DACC1" box="[222,399,441,464]" italics="true" pageId="4" pageNumber="59">M. rosenauense</emphasis>
|
||
(Verhoeff 1897a)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C614D46FFFDFFB3FD32989FFCD9ACC1" box="[657,811,440,464]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="silvaticum">
|
||
<emphasis id="B915EAD7FFFDFFB3FD32989FFCD9ACC1" box="[657,811,440,464]" italics="true" pageId="4" pageNumber="59">M. silvaticum</emphasis>
|
||
</taxonomicName>
|
||
. Females of these species have two bright longitudinal bands dorsally on a brownish ground, while males are dark grey with a black longitudinal line medially. Other species show three colour
|
||
<typeStatus id="54DA8867FFFDFFB3FE739B26FDFDAF09" box="[464,527,512,536]" pageId="4" pageNumber="59">types</typeStatus>
|
||
: (1) two longitudinal bright bands on brownish or dark grey ground (
|
||
<emphasis id="B915EAD7FFFDFFB3FAE99B27FF0DAF2D" italics="true" pageId="4" pageNumber="59">
|
||
e.g.
|
||
<taxonomicName id="4C614D46FFFDFFB3FADE9B27FF0DAF2D" authority="Verhoeff, 1898" authorityName="Verhoeff" authorityYear="1898" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="dentatum">M. dentatum</taxonomicName>
|
||
</emphasis>
|
||
(Verhoeff, 1898))—in these cases females are usually slightly more brightly colored; (2) one medial longitudinal yellowish or reddish band/dot-line (
|
||
<emphasis id="B915EAD7FFFDFFB3FD199B6FFC64AF71" box="[698,918,584,608]" italics="true" pageId="4" pageNumber="59">
|
||
e.g.
|
||
<taxonomicName id="4C614D46FFFDFFB3FD499B6FFC64AF71" authority="C. L. Koch, 1838" authorityName="C. L. Koch" authorityYear="1838" box="[746,918,584,608]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="unilineatum">M. unilineatum</taxonomicName>
|
||
</emphasis>
|
||
(C. L. Koch, 1838)) on dark grey ground; (3) brownish body without any bands (
|
||
<emphasis id="B915EAD7FFFDFFB3FD809B4BFDB9AF95" box="[547,587,621,644]" italics="true" pageId="4" pageNumber="59">e.g.</emphasis>
|
||
<taxonomicName id="4C614D46FFFDFFB3FDF19B4BFC31AF95" authority="(Verhoeff, 1928)" authorityName="Verhoeff" authorityYear="1928" baseAuthorityName="Verhoeff" baseAuthorityYear="1928" box="[594,963,620,645]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="rhodopinum">
|
||
<emphasis id="B915EAD7FFFDFFB3FDF19B4BFD0DAF95" box="[594,767,620,644]" italics="true" pageId="4" pageNumber="59">M. rhodopinum</emphasis>
|
||
(Verhoeff, 1928)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C614D46FFFDFFB3FC6C9B4BFB8CAF95" box="[975,1150,621,644]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="montivagum">
|
||
<emphasis id="B915EAD7FFFDFFB3FC6C9B4BFB8CAF95" box="[975,1150,621,644]" italics="true" pageId="4" pageNumber="59">M. montivagum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFDFFB3FF349BFEFF15AE02" blockId="4.[151,730,727,787]" pageId="4" pageNumber="59">
|
||
<emphasis id="B915EAD7FFFDFFB3FF349BFEFDE7AFE0" bold="true" box="[151,533,727,754]" pageId="4" pageNumber="59">
|
||
1. The
|
||
<taxonomicName id="4C614D46FFFDFFB3FF4F9BF1FDE7AFE0" box="[236,533,727,753]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="projectum">
|
||
<emphasis id="B915EAD7FFFDFFB3FF4F9BF1FDE7AFE0" bold="true" box="[236,533,727,753]" italics="true" pageId="4" pageNumber="59">Megaphyllum projectum</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<emphasis id="B915EAD7FFFDFFB3FD929BFEFD28AFE3" bold="true" box="[561,730,728,754]" pageId="4" pageNumber="59">species group</emphasis>
|
||
<figureCitation id="135A2A40FFFDFFB3FF349BDCFF15AE02" box="[151,231,762,787]" captionStart="FIGURES 1 a – e" captionStartId="4.[151,264,1450,1472]" captionTargetBox="[200,1386,1061,1424]" captionTargetId="figure@4.[197,1389,1058,1426]" captionTargetPageId="4" captionText="FIGURES 1 a – e. Schematic opisthomere drawings (right opisthomeres, lateral view). (1 a) Generalized scheme for all Megaphyllum sensu stricto. (1 b) Scheme for the M. projectum species group. (1 c) Scheme for the M. austriacum species group. (1 d) Scheme for the M. transsylvanicum species group. (1 e) Scheme for the M. unilineatum species group. Abbreviations: ah: anterior hump, ap: anterior process, asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere." httpUri="https://zenodo.org/record/223774/files/figure.png" pageId="4" pageNumber="59">Fig. 1b</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="8BDE36C5FFFDFFB3FF349A64FC5DAEFA" blockId="4.[151,1436,834,1003]" pageId="4" pageNumber="59">
|
||
Posterior opisthomere process (
|
||
<emphasis id="B915EAD7FFFDFFB3FE5B9A65FDE6AE4B" bold="true" box="[504,532,835,858]" italics="true" pageId="4" pageNumber="59">pp</emphasis>
|
||
) occasionally rudimentary (in
|
||
<taxonomicName id="4C614D46FFFDFFB3FCCF9A65FBF7AE4A" box="[876,1029,835,859]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="silvaticum">
|
||
<emphasis id="B915EAD7FFFDFFB3FCCF9A65FBF7AE4A" box="[876,1029,835,859]" italics="true" pageId="4" pageNumber="59">M. silvaticum</emphasis>
|
||
</taxonomicName>
|
||
); most species with anterior process (
|
||
<emphasis id="B915EAD7FFFDFFB3FF039A41FF4EAE6F" bold="true" box="[160,188,871,894]" italics="true" pageId="4" pageNumber="59">ap</emphasis>
|
||
) (except for
|
||
<taxonomicName id="4C614D46FFFDFFB3FEFA9A41FD21AE6E" authority="Latzel, 1884" authorityName="Latzel" authorityYear="1884" box="[345,723,870,895]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="erythronotum">
|
||
<emphasis id="B915EAD7FFFDFFB3FEFA9A41FDEBAE6F" box="[345,537,870,894]" italics="true" pageId="4" pageNumber="59">M. erythronotum</emphasis>
|
||
(Latzel, 1884))
|
||
</taxonomicName>
|
||
. No apical hump, neither anteriorly nor posteriorly. Posterior solenomere process (
|
||
<emphasis id="B915EAD7FFFDFFB3FE269AADFE5EAEB3" bold="true" box="[389,428,907,930]" italics="true" pageId="4" pageNumber="59">psp</emphasis>
|
||
) cockscomb-like, with a hook facing the anterior solenomere process (
|
||
<emphasis id="B915EAD7FFFDFFB3FB779AADFB09AEB3" bold="true" box="[1236,1275,907,930]" italics="true" pageId="4" pageNumber="59">asp</emphasis>
|
||
) (seen mostly from mesal view)—called “
|
||
<emphasis id="B915EAD7FFFDFFB3FE419A88FD40AED7" box="[482,690,942,966]" italics="true" pageId="4" pageNumber="59">Flagellumhäkchen</emphasis>
|
||
”,
|
||
<emphasis id="B915EAD7FFFDFFB3FD709A89FD07AED7" box="[723,757,943,966]" italics="true" pageId="4" pageNumber="59">i.e.</emphasis>
|
||
flagellal hook, by Verhoeff (1907). Anterior solenomere process (
|
||
<emphasis id="B915EAD7FFFDFFB3FF589AF5FED3AEFB" bold="true" box="[251,289,979,1002]" italics="true" pageId="4" pageNumber="59">asp</emphasis>
|
||
) usually shorter, mostly quite wide and easily discernible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |