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<document id="24799929B3CDF14E622383ECA9AE8D5C" ID-DOI="10.11646/zootaxa.5506.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13747211" ID-ZooBank="F7F39F20-07D0-434B-9DB7-160C19D519E5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726054659219" checkinUser="plazi" docAuthor="Volynkin, Anton V." docDate="2024" docId="03C887CEFFFAFFEDFF7CBAFAFD87FA6B" docLanguage="en" docName="zootaxa.5506.3.7.pdf" docOrigin="Zootaxa 5506 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thysanoplisna grandimacula Volynkin, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="433" masterDocId="FFF1FFB6FFFFFFE8FFEBBB04FFDAFF8E" masterDocTitle="Taxonomic review of the Thysanoptyx Hampson generic complex (Lepidoptera: Erebidae: Arctiinae: Lithosiini). II. The genus Thysanoplisna Volynkin with descriptions of seven new species from Vietnam and mainland China" masterLastPageNumber="450" masterPageNumber="428" pageNumber="433" updateTime="1726055383621" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6AE7192D8901D015513F886D71AF06F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="12FEE7FC78C25BB267ED3F162EBDE936">
<mods:title id="24387C7E8D78709CC1D922787DBBC5C7">Taxonomic review of the Thysanoptyx Hampson generic complex (Lepidoptera: Erebidae: Arctiinae: Lithosiini). II. The genus Thysanoplisna Volynkin with descriptions of seven new species from Vietnam and mainland China</mods:title>
</mods:titleInfo>
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<mods:role id="4A430DA33C83EDF39159A4DBA347DB1A">
<mods:roleTerm id="45001C197023CD53D1C893B2A31D6A20">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CBF7656E9EBCD048AD5296CAFFDFA547">Volynkin, Anton V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C15C38DC99233F3D73DBE1F55D1A431D">text</mods:typeOfResource>
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<mods:title id="404FBE6BAAEB714D128DB5BB92A9C731">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="77EB1DA824FD327FDE0B9D748152CBAC">
<mods:date id="9FDF835DE92C587CDF79C394DFE5F374">2024</mods:date>
<mods:detail id="F404D552929A064186C576087EDE26BE" type="pubDate">
<mods:number id="A493473D21F343C913D6544909D44927">2024-09-09</mods:number>
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<mods:detail id="C3A7FF2EB199070025D96055EC3A7D95" type="volume">
<mods:number id="5D5161524F2F67E4E2A77062A2A7EB14">5506</mods:number>
</mods:detail>
<mods:detail id="44DF960D2AE30C4F090389CAFE320DEB" type="issue">
<mods:number id="143E82A129341F4E16C3672C3FBB1B4E">3</mods:number>
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<mods:start id="C9335F95B4E6231D2B2AD72D930C580B">428</mods:start>
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<mods:location id="660F57BDF86EC1F5577601F20F6EA5F7">
<mods:url id="9277B8BB8D776DF1671EB9A8EF669471">http://dx.doi.org/10.11646/zootaxa.5506.3.7</mods:url>
</mods:location>
<mods:classification id="EDE8C89F7F6D0A34E175E829508F4CF2">journal article</mods:classification>
<mods:identifier id="5EB418D8A763E9C45973C56D704D25EA" type="DOI">10.11646/zootaxa.5506.3.7</mods:identifier>
<mods:identifier id="282EF89E97B61C5D44E2EB18BFBB8C4C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3BBEF1140C8BCC0319E7F061BD4F7799" type="Zenodo-Dep">13747211</mods:identifier>
<mods:identifier id="88A7C6F16F8B1AA50249F0632E5B0B6D" type="ZooBank">F7F39F20-07D0-434B-9DB7-160C19D519E5</mods:identifier>
</mods:mods>
<treatment id="03C887CEFFFAFFEDFF7CBAFAFD87FA6B" ID-DOI="http://doi.org/10.5281/zenodo.13747114" ID-Zenodo-Dep="13747114" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B" httpUri="http://treatment.plazi.org/id/03C887CEFFFAFFEDFF7CBAFAFD87FA6B" lastPageNumber="433" pageId="5" pageNumber="433">
<subSubSection id="C37B6553FFFAFFEDFF7CBAFAFD85FD97" box="[151,607,510,537]" pageId="5" pageNumber="433" type="nomenclature">
<paragraph id="8BDE36D8FFFAFFEDFF7CBAFAFD85FD97" blockId="5.[151,607,510,573]" box="[151,607,510,537]" pageId="5" pageNumber="433">
<heading id="D09681B4FFFAFFEDFF7CBAFAFD85FD97" bold="true" box="[151,607,510,537]" fontSize="11" level="1" pageId="5" pageNumber="433" reason="1">
<emphasis id="B915EACAFFFAFFEDFF7CBAFAFD85FD97" bold="true" box="[151,607,510,537]" pageId="5" pageNumber="433">
<taxonomicName id="4C614D5BFFFAFFEDFF7CBAFAFE20FD97" authorityName="Volynkin" authorityYear="2024" box="[151,506,510,537]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFAFFEDFF7CBAFAFE20FD97" bold="true" box="[151,506,510,537]" italics="true" pageId="5" pageNumber="433">Thysanoplisna grandimacula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657B1FFFAFFEDFDEABAFBFD85FD97" box="[513,607,511,537]" pageId="5" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFAFFEDFF7CB927FE15FDB3" box="[151,463,546,573]" pageId="5" pageNumber="433" type="description">
<paragraph id="8BDE36D8FFFAFFEDFF7CB927FE15FDB3" blockId="5.[151,607,510,573]" box="[151,463,546,573]" pageId="5" pageNumber="433">
(
<figureCitation id="135A2A5DFFFAFFEDFF74B927FEFAFDB3" box="[159,288,547,573]" captionStart="FIGURES 9–16" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[199,1388,182,1909]" captionTargetId="figure-15@6.[186,1401,181,1919]" captionTargetPageId="6" captionText="FIGURES 9–16. Adults of Thysanoplisna spp. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747215" httpUri="https://zenodo.org/record/13747215/files/figure.png" pageId="5" pageNumber="433">Figs 13–16</figureCitation>
,
<figureCitation id="135A2A5DFFFAFFEDFEC7B927FE91FDB3" box="[300,331,547,573]" captionStart="FIGURES 41–44" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[226,1371,185,1912]" captionTargetId="figure-26@15.[213,1375,181,1919]" captionTargetPageId="15" captionText="FIGURES 41–44. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747225" httpUri="https://zenodo.org/record/13747225/files/figure.png" pageId="5" pageNumber="433">44</figureCitation>
,
<figureCitation id="135A2A5DFFFAFFEDFEBDB926FEAEFDB3" box="[342,372,546,573]" captionStart="FIGURES 45–48" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[280,1308,181,1916]" captionTargetId="figure-15@16.[268,1320,181,1919]" captionTargetPageId="16" captionText="FIGURES 45–48. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747229" httpUri="https://zenodo.org/record/13747229/files/figure.png" pageId="5" pageNumber="433">45</figureCitation>
,
<figureCitation id="135A2A5DFFFAFFEDFE6BB926FE1DFDB3" box="[384,455,546,573]" captionStart="FIGURES 56–61" captionStartId="19.[151,264,1932,1956]" captionTargetBox="[195,1392,184,1895]" captionTargetId="figure-26@19.[182,1405,181,1908]" captionTargetPageId="19" captionText="FIGURES 56–61. Female genitalia of Thysanoplisna spp. Depositories of the specimens dissected: 56 and 57 in NHMUK (©The Trustees of NHMUK); 58–61 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747235" httpUri="https://zenodo.org/record/13747235/files/figure.png" pageId="5" pageNumber="433">59, 60</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFAFFEDFF7CB96FFDF2FC47" pageId="5" pageNumber="433" type="materials_examined">
<paragraph id="8BDE36D8FFFAFFEDFF7CB96FFD08FD7F" blockId="5.[151,1437,618,1509]" pageId="5" pageNumber="433">
<emphasis id="B915EACAFFFAFFEDFF7CB96FFEE7FD0A" bold="true" box="[151,317,618,645]" pageId="5" pageNumber="433">Type material</emphasis>
.
<materialsCitation id="3B093C85FFFAFFEDFEA1B96FFD08FD7F" collectingDate="1995-04-20" collectingDateMax="1995-04-30" collectingDateMin="1995-04-20" collectionCode="V, WITT, ZSM" county="Nord" elevation="1600" latitude="22.283333" location="Primarurwald" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" pageId="5" pageNumber="433" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="B915EACAFFFAFFEDFEA1B96FFE6DFD0B" bold="true" box="[330,439,619,645]" pageId="5" pageNumber="433">
<typeStatus id="54DA887AFFFAFFEDFEA1B96FFE6DFD0B" box="[330,439,619,645]" pageId="5" pageNumber="433" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="135A2A5DFFFAFFEDFE2FB96FFDC0FD0B" box="[452,538,619,645]" captionStart="FIGURES 9–16" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[199,1388,182,1909]" captionTargetId="figure-15@6.[186,1401,181,1919]" captionTargetPageId="6" captionText="FIGURES 9–16. Adults of Thysanoplisna spp. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747215" httpUri="https://zenodo.org/record/13747215/files/figure.png" pageId="5" pageNumber="433">Figs 13</figureCitation>
,
<figureCitation id="135A2A5DFFFAFFEDFDCEB96FFD9CFD0B" box="[549,582,619,645]" captionStart="FIGURES 41–44" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[226,1371,185,1912]" captionTargetId="figure-26@15.[213,1375,181,1919]" captionTargetPageId="15" captionText="FIGURES 41–44. Male genitalia of Thysanoplisna spp. The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13747225" httpUri="https://zenodo.org/record/13747225/files/figure.png" pageId="5" pageNumber="433">44</figureCitation>
):
<specimenCount id="9D67FD51FFFAFFEDFDB3B96FFDB5FD0B" box="[600,623,619,645]" count="1" pageId="5" pageNumber="433" type="male">♂</specimenCount>
, “N-Vietnam
<quantity id="4C999B3DFFFAFFEDFCE3B96FFC8FFD0B" box="[776,853,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="433" unit="m" value="1600.0">
<elevation id="004CD1EBFFFAFFEDFCE3B96FFC8FFD0B" box="[776,853,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="433" unit="m" value="1600.0">1600m</elevation>
</quantity>
| [
<collectingRegion id="49A5F83AFFFAFFEDFC85B96FFBECFD0B" box="[878,1078,619,645]" country="Vietnam" name="Lao Cai" pageId="5" pageNumber="433">Lào Cai Province</collectingRegion>
] Mt. Fan-si-pan(
<collectingCounty id="62BF4E54FFFAFFEDFB1BB96FFAF1FD0B" box="[1264,1323,619,645]" pageId="5" pageNumber="433">Nord</collectingCounty>
) |
<collectingMunicipality id="6BBAACA2FFFAFFEDFAAFB96FFA42FD0B" box="[1348,1432,619,645]" pageId="5" pageNumber="433">Cha-pa</collectingMunicipality>
,
<location id="8EBE6003FFFAFFEDFF7CB98BFEEBFD27" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFF7CB98BFEEBFD27" box="[151,305,655,681]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Primarurwald" pageId="5" pageNumber="433" stateProvince="Lao Cai">Primärurwald</location>
[primary jungle] | 22.17''N 103.44''E [
<geoCoordinate id="EE55501FFFFAFFEDFD36B98BFCE6FD27" box="[733,828,655,681]" degrees="22" direction="north" minutes="17" orientation="latitude" pageId="5" pageNumber="433" precision="925" value="22.283333">22°17'N</geoCoordinate>
<geoCoordinate id="EE55501FFFFAFFEDFCA8B98BFC76FD27" box="[835,940,655,681]" degrees="103" direction="east" minutes="44" orientation="longitude" pageId="5" pageNumber="433" precision="925" value="103.73333">103°44'E</geoCoordinate>
] |
<date id="FFDF1018FFFAFFEDFC2CB98AFBAEFD27" box="[967,1140,654,681]" pageId="5" pageNumber="433" value="1995-04-20" valueMax="1995-04-30" valueMin="1995-04-20">
<collectingDate id="EF9BE9F0FFFAFFEDFC2CB98AFBAEFD27" box="[967,1140,654,681]" pageId="5" pageNumber="433" value="1995-04-20" valueMax="1995-04-30" valueMin="1995-04-20">20.–30.IV.1995</collectingDate>
</date>
| leg.
<collectionCode id="ED70AE1DFFFAFFEDFB5DB98BFB10FD27" box="[1206,1226,655,681]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="433" type="Museum">V</collectionCode>
.
<location id="8EBE6003FFFAFFEDFB3EB98BFAF3FD27" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFB3EB98BFAF3FD27" box="[1237,1321,655,681]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Sinjaev" pageId="5" pageNumber="433" stateProvince="Lao Cai">Sinjaev</location>
[
<location id="8EBE6003FFFAFFEDFADDB98BFA4CFD27" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFADDB98BFA4CFD27" box="[1334,1430,655,681]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Sinyaev" pageId="5" pageNumber="433" stateProvince="Lao Cai">Sinyaev</location>
] &amp; | Einheim. Sammler [native collector]” | vertical inscription “
<location id="8EBE6003FFFAFFEDFCB5B9B7FC26FD43" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFCB5B9B7FC26FD43" box="[862,1020,691,717]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Museum Witt" pageId="5" pageNumber="433" stateProvince="Lao Cai">Museum Witt</location>
” / “
<location id="8EBE6003FFFAFFEDFBCDB9B7FB0DFD43" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFBCDB9B7FB0DFD43" box="[1062,1239,691,717]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Genitalpraparat" pageId="5" pageNumber="433" stateProvince="Lao Cai">Genitalpräparat</location>
|
<location id="8EBE6003FFFAFFEDFB00B9B7FABCFD43" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFB00B9B7FABCFD43" box="[1259,1382,691,717]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Heterocera" pageId="5" pageNumber="433" stateProvince="Lao Cai">Heterocera</location>
| Nr. 36.023 |
<location id="8EBE6003FFFAFFEDFF13B9D3FE80FD7F" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFF13B9D3FE80FD7F" box="[248,346,727,753]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Museum" pageId="5" pageNumber="433" stateProvince="Lao Cai">Museum</location>
<collectionCode id="ED70AE1DFFFAFFEDFE8BB9D3FE7FFD7F" box="[352,421,727,753]" pageId="5" pageNumber="433">WITT</collectionCode>
<location id="8EBE6003FFFAFFEDFE46B9D3FDC3FD7F" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFE46B9D3FDC3FD7F" box="[429,537,727,753]" county="Nord" latitude="22.283333" longLatPrecision="1260" longitude="103.73333" municipality="Cha-pa" name="Munchen" pageId="5" pageNumber="433" stateProvince="Lao Cai">München</location>
” (MWM/
<collectionCode id="ED70AE1DFFFAFFEDFD61B9D3FD1CFD7F" box="[650,710,727,753]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="433" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDE36D8FFFAFFEDFF2CB9FFFDF2FC47" blockId="5.[151,1437,618,1509]" pageId="5" pageNumber="433">
<materialsCitation id="3B093C85FFFAFFEDFF2CB9FFFD78FC9B" box="[199,674,762,789]" country="Vietnam" location="VIETNAM" pageId="5" pageNumber="433" specimenCount="159" specimenCount-female="70" specimenCount-male="89" typeStatus="paratype">
<emphasis id="B915EACAFFFAFFEDFF2CB9FFFEE5FC9B" bold="true" box="[199,319,763,789]" pageId="5" pageNumber="433">
<typeStatus id="54DA887AFFFAFFEDFF2CB9FFFEE5FC9B" box="[199,319,763,789]" pageId="5" pageNumber="433" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="B915EACAFFFAFFEDFEBAB9FEFE02FC9A" bold="true" box="[337,472,762,788]" pageId="5" pageNumber="433">
<collectingCountry id="F3767648FFFAFFEDFEBAB9FEFE02FC9A" box="[337,472,762,788]" name="Vietnam" pageId="5" pageNumber="433">VIETNAM</collectingCountry>
</emphasis>
:
<specimenCount id="9D67FD51FFFAFFEDFE07B9FFFD9AFC9B" box="[492,576,763,789]" count="89" pageId="5" pageNumber="433" type="male">89 ♂♂</specimenCount>
,
<specimenCount id="9D67FD51FFFAFFEDFDBBB9FFFD78FC9B" box="[592,674,763,789]" count="70" pageId="5" pageNumber="433" type="female">70 ♀♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B093C85FFFAFFEDFD5FB9FFFD5AFCB7" collectingDate="1995-04-20" collectingDateMax="1995-04-30" collectingDateMin="1995-04-20" collectionCode="V, WITT, ZSM" country="Vietnam" county="Nord" elevation="1600" latitude="35.961" location="Nos." longLatPrecision="71" longitude="36.025" municipality="Cha-pa" pageId="5" pageNumber="433" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="paratype">
same data as holotype, gen. prep.
<location id="8EBE6003FFFAFFEDFBA1B9FFFB58FC9B" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFBA1B9FFFB58FC9B" box="[1098,1154,763,789]" country="Vietnam" latitude="35.961" longLatPrecision="71" longitude="36.025" name="Nos." pageId="5" pageNumber="433">Nos.</location>
: MWM 35.960, 35.962, 35.963, 36.024 (
<specimenCount id="9D67FD51FFFAFFEDFEBBB81BFEA7FCB7" box="[336,381,799,825]" count="1" pageId="5" pageNumber="433" type="male">♂♂</specimenCount>
),
<geoCoordinate id="EE55501FFFFAFFEDFE79B81AFE38FCB7" box="[402,482,798,825]" degrees="35.961" direction="north" orientation="latitude" pageId="5" pageNumber="433" precision="55" value="35.961">35.961</geoCoordinate>
,
<geoCoordinate id="EE55501FFFFAFFEDFE05B81AFDE1FCB7" box="[494,571,798,825]" degrees="36.025" direction="east" orientation="longitude" pageId="5" pageNumber="433" precision="55" value="36.025">36.025</geoCoordinate>
(
<specimenCount id="9D67FD51FFFAFFEDFDA6B81BFDAFFCB7" box="[589,629,799,825]" count="1" pageId="5" pageNumber="433" type="female">♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B093C85FFFAFFEDFD65B81BFD63FCB7" box="[654,697,799,825]" collectingDate="1995-04-20" collectingDateMax="1995-04-30" collectingDateMin="1995-04-20" collectionCode="V, WITT, ZSM" country="Vietnam" county="Nord" elevation="1600" isTailEnumeration="true" latitude="35.961" location="Nos." longLatPrecision="71" longitude="36.025" municipality="Cha-pa" pageId="5" pageNumber="433" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="9D67FD51FFFAFFEDFD65B81BFD63FCB7" box="[654,697,799,825]" count="1" pageId="5" pageNumber="433" type="female">1 ♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B093C85FFFAFFEDFD2CB81BFC54FCD3" collectingDate="1994-09" collectorName="Mong" country="Vietnam" county="Mt." elevation="1700" latitude="22.333334" location="Sapa" longLatPrecision="1260" longitude="103.666664" municipality="Cha-pa" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Lao Cai" typeStatus="paratype">
[
<collectingRegion id="49A5F83AFFFAFFEDFD25B81BFC43FCB7" box="[718,921,799,825]" country="Vietnam" name="Lao Cai" pageId="5" pageNumber="433">Lào Cai Province</collectingRegion>
]
<quantity id="4C999B3DFFFAFFEDFC4DB81BFBE7FCB7" box="[934,1085,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="5" pageNumber="433" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="004CD1EBFFFAFFEDFC4DB81BFBE7FCB7" box="[934,1085,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="5" pageNumber="433" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800m</elevation>
</quantity>
,
<collectingCounty id="62BF4E54FFFAFFEDFBA3B81BFBABFCB7" box="[1096,1137,799,825]" pageId="5" pageNumber="433">Mt.</collectingCounty>
Fan-si-pan (west),
<collectingMunicipality id="6BBAACA2FFFAFFEDFAA0B81BFA46FCB7" box="[1355,1436,799,825]" pageId="5" pageNumber="433">Cha-pa</collectingMunicipality>
(=
<location id="8EBE6003FFFAFFEDFF44B847FF30FCD3" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFF44B847FF30FCD3" box="[175,234,835,861]" country="Vietnam" county="Mt." latitude="22.333334" longLatPrecision="1260" longitude="103.666664" municipality="Cha-pa" name="Sapa" pageId="5" pageNumber="433" stateProvince="Lao Cai">Sapa</location>
),
<geoCoordinate id="EE55501FFFFAFFEDFEEBB847FE86FCD3" box="[256,348,835,861]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="5" pageNumber="433" precision="925" value="22.333334">22°20'N</geoCoordinate>
<geoCoordinate id="EE55501FFFFAFFEDFE8EB847FE15FCD3" box="[357,463,835,861]" degrees="103" direction="east" minutes="40" orientation="longitude" pageId="5" pageNumber="433" precision="925" value="103.666664">103°40'E</geoCoordinate>
,
<date id="FFDF1018FFFAFFEDFE37B847FDEAFCD3" box="[476,560,835,861]" pageId="5" pageNumber="433" value="1994-09">
<collectingDate id="EF9BE9F0FFFAFFEDFE37B847FDEAFCD3" box="[476,560,835,861]" pageId="5" pageNumber="433" value="1994-09">ix.1994</collectingDate>
</date>
(secondary forest),
<collectorName id="2694530EFFFAFFEDFCFDB847FC83FCD3" box="[790,857,835,861]" pageId="5" pageNumber="433">Mong</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="9D67FD51FFFAFFEDFC70B847FC36FCD3" box="[923,1004,835,861]" count="17" pageId="5" pageNumber="433" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D67FD51FFFAFFEDFC11B847FBE1FCD3" box="[1018,1083,835,861]" count="8" pageId="5" pageNumber="433" type="female">8 ♀♀</specimenCount>
,
<materialsCitation id="3B093C85FFFAFFEDFBA1B847FEB2FC2B" collectingDate="1995-04" collectionCode="V" collectorName="Sinyaev &amp; No." country="Vietnam" elevation="950" latitude="21.566668" location="Tam Dao" longLatPrecision="1263" longitude="105.333336" pageId="5" pageNumber="433" specimenCount="1" specimenCount-female="1" stateProvince="Tuyen Quang" typeStatus="paratype">
[
<collectingRegion id="49A5F83AFFFAFFEDFBBBB847FA87FCD3" box="[1104,1373,835,861]" country="Vietnam" name="Tuyen Quang" pageId="5" pageNumber="433">Tuyên Quang Province</collectingRegion>
]
<location id="8EBE6003FFFAFFEDFA80B847FF1CFC0F" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFA80B847FF1CFC0F" country="Vietnam" latitude="21.566668" longLatPrecision="1263" longitude="105.333336" name="Tam Dao" pageId="5" pageNumber="433" stateProvince="Tuyen Quang">Tam Dao</location>
(secondary forest),
<locationDeviation id="9A6C2FFAFFFAFFEDFE40B863FD40FC0F" box="[427,666,871,897]" location="Hanoi" pageId="5" pageNumber="433">
<quantity id="4C999B3DFFFAFFEDFE40B863FE2EFC0F" box="[427,500,871,897]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="433" unit="km" value="60.0">60 km</quantity>
NW of
<collectingRegion id="49A5F83AFFFAFFEDFDB8B863FD40FC0F" box="[595,666,871,897]" country="Vietnam" name="Ha Noi" pageId="5" pageNumber="433">Hanoi</collectingRegion>
</locationDeviation>
,
<quantity id="4C999B3DFFFAFFEDFD4CB862FD30FC0F" box="[679,746,870,897]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="433" unit="m" value="950.0">
<elevation id="004CD1EBFFFAFFEDFD4CB862FD30FC0F" box="[679,746,870,897]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="433" unit="m" value="950.0">950m</elevation>
</quantity>
,
<geoCoordinate id="EE55501FFFFAFFEDFD1CB863FC89FC0F" box="[759,851,871,897]" degrees="21" direction="north" minutes="34" orientation="latitude" pageId="5" pageNumber="433" precision="925" value="21.566668">21°34'N</geoCoordinate>
<geoCoordinate id="EE55501FFFFAFFEDFCB7B862FC1CFC0F" box="[860,966,870,897]" degrees="105" direction="east" minutes="20" orientation="longitude" pageId="5" pageNumber="433" precision="925" value="105.333336">105°20'E</geoCoordinate>
,
<date id="FFDF1018FFFAFFEDFC38B862FBF3FC0F" box="[979,1065,870,897]" pageId="5" pageNumber="433" value="1995-04">
<collectingDate id="EF9BE9F0FFFAFFEDFC38B862FBF3FC0F" box="[979,1065,870,897]" pageId="5" pageNumber="433" value="1995-04">iv.1995</collectingDate>
</date>
,
<collectionCode id="ED70AE1DFFFAFFEDFBDEB863FB93FC0F" box="[1077,1097,871,897]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="433" type="Museum">V</collectionCode>
.
<collectorName id="2694530EFFFAFFEDFBBDB863FB6AFC0F" box="[1110,1200,871,897]" pageId="5" pageNumber="433">Sinyaev</collectorName>
leg., gen. prep.
<collectorName id="2694530EFFFAFFEDFA80B863FA42FC0F" box="[1387,1432,871,897]" pageId="5" pageNumber="433">No.</collectorName>
: MWM 36.028 (
<specimenCount id="9D67FD51FFFAFFEDFEA1B88FFE87FC2B" box="[330,349,907,933]" count="1" pageId="5" pageNumber="433" type="female">♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B093C85FFFAFFEDFE9EB88FFE45FC2B" box="[373,415,907,933]" collectingDate="1995-04" collectionCode="V" collectorName="Sinyaev &amp; No." country="Vietnam" elevation="950" isTailEnumeration="true" latitude="21.566668" location="Tam Dao" longLatPrecision="1263" longitude="105.333336" pageId="5" pageNumber="433" specimenCount="1" specimenCount-female="1" stateProvince="Tuyen Quang" typeStatus="paratype">
<specimenCount id="9D67FD51FFFAFFEDFE9EB88FFE45FC2B" box="[373,415,907,933]" count="1" pageId="5" pageNumber="433" type="female">1 ♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B093C85FFFAFFEDFE46B88FFDFEFC47" collectingDate="1997-02-22" collectingDateMax="1997-03-05" collectingDateMin="1997-02-22" collectionCode="ZSM" collectorName="G. Csorba" country="Vietnam" elevation="300" latitude="22.05" location="Na Hang Nature Reserve" longLatPrecision="1261" longitude="105.083336" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Tuyen Quang" typeStatus="paratype">
<collectingRegion id="49A5F83AFFFAFFEDFE46B88FFD50FC2B" box="[429,650,907,933]" country="Vietnam" name="Tuyen Quang" pageId="5" pageNumber="433">Tuyên Quang Prov.</collectingRegion>
,
<location id="8EBE6003FFFAFFEDFD7FB88FFC6AFC2B" LSID="urn:lsid:plazi:treatment:03C887CEFFFAFFEDFF7CBAFAFD87FA6B:8EBE6003FFFAFFEDFD7FB88FFC6AFC2B" box="[660,944,907,933]" country="Vietnam" latitude="22.05" longLatPrecision="1261" longitude="105.083336" name="Na Hang Nature Reserve" pageId="5" pageNumber="433" stateProvince="Tuyen Quang">Na Hang Nature Reserve</location>
,
<quantity id="4C999B3DFFFAFFEDFC50B88FFC24FC2B" box="[955,1022,907,933]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="433" unit="m" value="300.0">
<elevation id="004CD1EBFFFAFFEDFC50B88FFC24FC2B" box="[955,1022,907,933]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="433" unit="m" value="300.0">300m</elevation>
</quantity>
,
<geoCoordinate id="EE55501FFFFAFFEDFBE2B88FFBBCFC2B" box="[1033,1126,907,933]" degrees="22" direction="north" minutes="03" orientation="latitude" pageId="5" pageNumber="433" precision="925" value="22.05">22°03'N</geoCoordinate>
<geoCoordinate id="EE55501FFFFAFFEDFB86B88EFB0CFC2B" box="[1133,1238,906,933]" degrees="105" direction="east" minutes="05" orientation="longitude" pageId="5" pageNumber="433" precision="925" value="105.083336">105°05'E</geoCoordinate>
,
<date id="FFDF1018FFFAFFEDFB09B88EFA4DFC2B" box="[1250,1431,906,933]" pageId="5" pageNumber="433" value="1997-02-22" valueMax="1997-03-05" valueMin="1997-02-22">
<collectingDate id="EF9BE9F0FFFAFFEDFB09B88EFA4DFC2B" box="[1250,1431,906,933]" pageId="5" pageNumber="433" value="1997-02-22" valueMax="1997-03-05" valueMin="1997-02-22">22.ii.–5.iii.1997</collectingDate>
</date>
,
<collectorName id="2694530EFFFAFFEDFF7CB8ABFED2FC47" box="[151,264,943,969]" pageId="5" pageNumber="433">G. Csorba</collectorName>
leg. (all in MWM/
<collectionCode id="ED70AE1DFFFAFFEDFE0AB8ABFDC6FC47" box="[481,540,943,969]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="433" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFAFFEDFF2CB8D6FAE4FADB" pageId="5" pageNumber="433" type="diagnosis">
<paragraph id="8BDE36D8FFFAFFEDFF2CB8D6FAE4FADB" blockId="5.[151,1437,618,1509]" pageId="5" pageNumber="433">
<emphasis id="B915EACAFFFAFFEDFF2CB8D6FEE0FC62" bold="true" box="[199,314,978,1004]" pageId="5" pageNumber="433">Diagnosis</emphasis>
. The forewing length is 15.5–18.0 mm in males and 18.5–19.0 mm in females.
<taxonomicName id="4C614D5BFFFAFFEDFB12B8D7FEEEFB9F" authorityName="Volynkin" authorityYear="2024" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFAFFEDFB12B8D7FEEEFB9F" italics="true" pageId="5" pageNumber="433">Thysanoplisna grandimacula</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFAFFEDFEABB8F3FE47FB9F" bold="true" box="[320,413,1015,1041]" pageId="5" pageNumber="433">
<taxonomicNameLabel id="A22657B1FFFAFFEDFEABB8F3FE47FB9F" box="[320,413,1015,1041]" pageId="5" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
superficially differs from the similar
<taxonomicName id="4C614D5BFFFAFFEDFCB7B8F3FBCDFB9F" authorityName="Alpheraky" authorityYear="1897" box="[860,1047,1015,1041]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFAFFEDFCB7B8F3FBCDFB9F" box="[860,1047,1015,1041]" italics="true" pageId="5" pageNumber="433">Th. brevimacula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D5BFFFAFFEDFBBDB8F3FB02FB9E" authorityName="Walker" authorityYear="1854" box="[1110,1240,1015,1041]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFAFFEDFBBDB8F3FB02FB9E" box="[1110,1240,1015,1041]" italics="true" pageId="5" pageNumber="433">Th. signata</emphasis>
</taxonomicName>
in the markedly longer medial spot on the forewing. The male genital capsule of
<taxonomicName id="4C614D5BFFFAFFEDFC96BF1FFB9DFBBB" authorityName="Volynkin" authorityYear="2024" box="[893,1095,1051,1077]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFAFFEDFC96BF1FFB9DFBBB" box="[893,1095,1051,1077]" italics="true" pageId="5" pageNumber="433">Th. grandimacula</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFAFFEDFBBBBF1FFB71FBBB" bold="true" box="[1104,1195,1051,1077]" pageId="5" pageNumber="433">
<taxonomicNameLabel id="A22657B1FFFAFFEDFBBBBF1FFB71FBBB" box="[1104,1195,1051,1077]" pageId="5" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C614D5BFFFAFFEDFF3DBF3BFE55FBD7" authorityName="Alpheraky" authorityYear="1897" box="[214,399,1087,1113]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFAFFEDFF3DBF3BFE55FBD7" box="[214,399,1087,1113]" italics="true" pageId="5" pageNumber="433">Th. brevimacula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D5BFFFAFFEDFE23BF3BFD92FBD6" authorityName="Walker" authorityYear="1854" box="[456,584,1087,1113]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFAFFEDFE23BF3BFD92FBD6" box="[456,584,1087,1113]" italics="true" pageId="5" pageNumber="433">Th. signata</emphasis>
</taxonomicName>
by the somewhat more rounded vinculum and the slightly straighter distal saccular process. The phallus of the new species is longer and broader than in the similar species. Compared to
<taxonomicName id="4C614D5BFFFAFFEDFA93BF67FEF8FB2F" authorityName="Alpheraky" authorityYear="1897" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFAFFEDFA93BF67FEF8FB2F" italics="true" pageId="5" pageNumber="433">Th. brevimacula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D5BFFFAFFEDFEBCBF83FE09FB2E" authorityName="Walker" authorityYear="1854" box="[343,467,1159,1185]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFAFFEDFEBCBF83FE09FB2E" box="[343,467,1159,1185]" italics="true" pageId="5" pageNumber="433">Th. signata</emphasis>
</taxonomicName>
, the vesica of
<taxonomicName id="4C614D5BFFFAFFEDFD98BF83FCE0FB2F" authorityName="Volynkin" authorityYear="2024" box="[627,826,1159,1185]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFAFFEDFD98BF83FCE0FB2F" box="[627,826,1159,1185]" italics="true" pageId="5" pageNumber="433">Th. grandimacula</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFAFFEDFCAABF83FC43FB2F" bold="true" box="[833,921,1159,1185]" pageId="5" pageNumber="433">
<taxonomicNameLabel id="A22657B1FFFAFFEDFCAABF83FC43FB2F" box="[833,921,1159,1185]" pageId="5" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is considerably broader and somewhat longer, has a shorter basal diverticulum, broader subbasal diverticula, a longer and bilobate dorsal medial diverticulum, a bilobate (foot-shaped) ventral medial diverticulum, and bears a longer and thicker cornutus. The female genitalia of
<taxonomicName id="4C614D5BFFFAFFEDFF5EBFF7FEA4FA83" authorityName="Volynkin" authorityYear="2024" box="[181,382,1267,1293]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="grandimacula" status="sp. nov.">
<emphasis id="B915EACAFFFAFFEDFF5EBFF7FEA4FA83" box="[181,382,1267,1293]" italics="true" pageId="5" pageNumber="433">Th. grandimacula</emphasis>
</taxonomicName>
<emphasis id="B915EACAFFFAFFEDFE6EBFF7FE04FA83" bold="true" box="[389,478,1267,1293]" pageId="5" pageNumber="433">
<taxonomicNameLabel id="A22657B1FFFAFFEDFE6EBFF7FE04FA83" box="[389,478,1267,1293]" pageId="5" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are distinguished from
<taxonomicName id="4C614D5BFFFAFFEDFD0CBFF7FC47FA83" authorityName="Alpheraky" authorityYear="1897" box="[743,925,1267,1293]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="brevimacula">
<emphasis id="B915EACAFFFAFFEDFD0CBFF7FC47FA83" box="[743,925,1267,1293]" italics="true" pageId="5" pageNumber="433">Th. brevimacula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D5BFFFAFFEDFC38BFF7FB8BFA82" authorityName="Walker" authorityYear="1854" box="[979,1105,1267,1293]" class="Insecta" family="Erebidae" genus="Lithosia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B915EACAFFFAFFEDFC38BFF7FB8BFA82" box="[979,1105,1267,1293]" italics="true" pageId="5" pageNumber="433">Th. signata</emphasis>
</taxonomicName>
by the anteriorly broader and more posteriorly tapered ductus bursae, the markedly broader posterior gelatinous enlargement of the corpus bursae bearing a large and heavily sclerotised lateral plate, and the shorter and distally narrower appendix bursae.
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFAFFEDFF2CBE5AFC2AFAF7" box="[199,1008,1374,1401]" pageId="5" pageNumber="433" type="distribution">
<paragraph id="8BDE36D8FFFAFFEDFF2CBE5AFC2AFAF7" blockId="5.[151,1437,618,1509]" box="[199,1008,1374,1401]" pageId="5" pageNumber="433">
<emphasis id="B915EACAFFFAFFEDFF2CBE5AFE83FAF6" bold="true" box="[199,345,1374,1400]" pageId="5" pageNumber="433">Distribution</emphasis>
. Northern
<collectingCountry id="F3767648FFFAFFEDFE38BE5BFDE8FAF7" box="[467,562,1375,1401]" name="Vietnam" pageId="5" pageNumber="433">Vietnam</collectingCountry>
(
<collectingRegion id="49A5F83AFFFAFFEDFDABBE5BFD46FAF7" box="[576,668,1375,1401]" country="Vietnam" name="Lao Cai" pageId="5" pageNumber="433">Lào Cai</collectingRegion>
and
<collectingRegion id="49A5F83AFFFAFFEDFD3ABE5BFCB0FAF7" box="[721,874,1375,1401]" country="Vietnam" name="Tuyen Quang" pageId="5" pageNumber="433">Tuyên Quang</collectingRegion>
Provinces).
</paragraph>
</subSubSection>
<subSubSection id="C37B6553FFFAFFEDFF2CBE87FD87FA6B" pageId="5" pageNumber="433" type="etymology">
<paragraph id="8BDE36D8FFFAFFEDFF2CBE87FD87FA6B" blockId="5.[151,1437,618,1509]" pageId="5" pageNumber="433">
<emphasis id="B915EACAFFFAFFEDFF2CBE87FE92FA13" bold="true" box="[199,328,1411,1437]" pageId="5" pageNumber="433">Etymology</emphasis>
. The specific epithet is derived from the Latin ‘grandis’ and ‘macula’ meaning ‘large’ and ‘spot’, respectively, and refers to the medial spot of the forewing, which is larger than in similar species. The name is a noun in the nominative singular in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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