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<document id="7CF50CBE1AFAFC52228BE37F667F3C34" ID-CLB-Dataset="32718" ID-DOI="10.11646/zootaxa.4290.2.1" ID-GBIF-Dataset="43ebda6c-201a-46af-8b54-e7980b8d79e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="828965" ID-ZooBank="52D6E3C3-BA0A-49C7-865C-00F3F5E5C5D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110315309" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2017" docId="03C887A8FFC5FFC9FF737FA25AF4B099" docLanguage="en" docName="zootaxa.4290.2.1.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Matthewsius" docType="key" docVersion="7" lastPageNumber="215" masterDocId="FFF1FFD0FFCBFFC7FFE47F3A5F6FB348" masterDocTitle="Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species" masterLastPageNumber="243" masterPageNumber="201" pageNumber="215" updateTime="1698465402479" updateUser="ExternalLinkService">
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<mods:title id="F706C66A5141AD6F7D85DC1D16AE661A">Two new genera of Australian dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) with the description of six new species and transfer of six described species</mods:title>
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<mods:namePart id="FD7BCA6A5042FADA22AC40EF25289226">Gunter, Nicole L.</mods:namePart>
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<treatment id="03C887A8FFC5FFC9FF737FA25AF4B099" ID-DOI="http://doi.org/10.5281/zenodo.6032990" ID-GBIF-Taxon="132549212" ID-Zenodo-Dep="6032990" LSID="urn:lsid:plazi:treatment:03C887A8FFC5FFC9FF737FA25AF4B099" httpUri="http://treatment.plazi.org/id/03C887A8FFC5FFC9FF737FA25AF4B099" lastPageNumber="215" pageId="14" pageNumber="215">
<subSubSection id="C37B6535FFC5FFC9FF737FA25E88B3FA" box="[151,487,151,178]" pageId="14" pageNumber="215" type="nomenclature">
<paragraph id="8BDE36BEFFC5FFC9FF737FA25E88B3FA" blockId="14.[151,487,151,178]" box="[151,487,151,178]" pageId="14" pageNumber="215">
<heading id="D09681D2FFC5FFC9FF737FA25E88B3FA" bold="true" box="[151,487,151,178]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
<emphasis id="B915EAACFFC5FFC9FF737FA25E88B3FA" bold="true" box="[151,487,151,178]" pageId="14" pageNumber="215">
Key to
<taxonomicName id="4C614D3DFFC5FFC9FF147FAD5EE6B3F9" ID-CoL="5M3C" box="[240,393,151,177]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="216" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B915EAACFFC5FFC9FF147FAD5EE6B3F9" bold="true" box="[240,393,151,177]" italics="true" pageId="14" pageNumber="215">Matthewsius</emphasis>
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species
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</paragraph>
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<subSubSection id="C37B6535FFC5FFC9FF737FE45AF4B099" pageId="14" pageNumber="215" type="key">
<keyStep id="30952E1BFFC5FFC9FF737FE45AF4B203" pageId="14" pageNumber="215">
<paragraph id="8BDE36BEFFC5FFC9FF737FE45AF3B258" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737FE45AF3B258" pageId="14" pageNumber="215">
1. Pygidium with transverse groove at base narrowed medially (
<figureCitation id="135A2A3BFFC5FFC9FCDB7FE45CEBB3BB" box="[831,900,222,243]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Figs. 3</figureCitation>
B, D); elytral striae narrow, shallowly impressed, with simple punctures; elytra glabrous or with extremely fine setae..................................................2
</keyLead>
</paragraph>
<paragraph id="8BDE36BEFFC5FFC9FF737E235AF4B203" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737E235AF4B203" pageId="14" pageNumber="215">
- Pygidium with transverse groove at base not narrowed medially (
<figureCitation id="135A2A3BFFC5FFC9FCB37E235CF6B266" box="[855,921,281,302]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Figs. 3</figureCitation>
A, C, E–F); elytral striae wider, geminate, containing simple punctures, crenulate apically or not; elytra glabrous or with thick setae.....................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E1BFFC5FFC9FF737E695AF9B2B3" pageId="14" pageNumber="215">
<paragraph id="8BDE36BEFFC5FFC9FF737E695AF4B2EB" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737E695AF4B2EB" pageId="14" pageNumber="215">
2. Elytra microreticulate, more evident apically, masking punctures on interstices; striae 7 short, as strongly impressed as 1–6, only present in basal 1/3; eyes separated by about 14 eye-diameters; protibiae without serrations between apical and second tooth; aedeagus with elongate parameres as in
<figureCitation id="135A2A3BFFC5FFC9FD7B7EB45DB8B2EB" box="[671,727,398,419]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
D; northern Queensland.....
<taxonomicName id="4C614D3DFFC5FFC9FC1B7EB45AF4B2EB" ID-CoL="3YF86" authority="Storey &amp; Monteith" authorityName="Storey &amp; Monteith" box="[1023,1435,398,419]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="speewah">
<emphasis id="B915EAACFFC5FFC9FC1B7EB45BBEB2EB" box="[1023,1233,398,419]" italics="true" pageId="14" pageNumber="215">Matthewsius speewah</emphasis>
(Storey &amp; Monteith)
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</keyLead>
</paragraph>
<paragraph id="8BDE36BEFFC5FFC9FF737E915AF9B2B3" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737E915AF9B2B3" pageId="14" pageNumber="215">
-
<materialsCitation id="3B093CE3FFC5FFC9FF057E915AF9B2B3" ID-GBIF-Occurrence="1572325585" location="South Australia" pageId="14" pageNumber="215" specimenCount="1" stateProvince="South Australia">
Elytra shiny, smooth; small punctures on interstices clearly visible; striae 7 much less impressed than 1–6, longer but only present in basal 2/3; eyes separated by more than 20 eye-diameters; protibiae with several serrations between apical and second tooth; aedeagus as in
<figureCitation id="135A2A3BFFC5FFC9FE3D7EDC5D7EB2B3" box="[473,529,486,507]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
B;
<collectingRegion id="49A5F85CFFC5FFC9FDCA7EDC5DACB2B3" box="[558,707,486,507]" country="Australia" name="South Australia" pageId="14" pageNumber="215">South Australia</collectingRegion>
........................
<taxonomicName id="4C614D3DFFC5FFC9FC167EDC5AF9B2B3" ID-CoL="3YF84" authority="Matthews &amp; Weir" authorityName="Matthews &amp; Weir" box="[1010,1430,486,507]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="penelopae">
<emphasis id="B915EAACFFC5FFC9FC167EDC5BBAB2B3" box="[1010,1237,486,507]" italics="true" pageId="14" pageNumber="215">Matthewsius penelopae</emphasis>
(Matthews &amp; Weir)
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</materialsCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E1BFFC5FFC9FF737D395AF3B1C6" pageId="14" pageNumber="215">
<paragraph id="8BDE36BEFFC5FFC9FF737D395AF4B11B" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737D395AF4B11B" pageId="14" pageNumber="215">
3. Clypeal edge with a blunt projection lateral to clypeal teeth; elytral striae punctate but not crenulate; protibiae without serrations between apical and second tooth; aedeagus as in
<figureCitation id="135A2A3BFFC5FFC9FD0E7D1B5C4DB17E" box="[746,802,545,566]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
F;
<materialsCitation id="3B093CE3FFC5FFC9FCD87D1B5AF4B11B" ID-GBIF-Occurrence="1572325631" location="Western Australia" pageId="14" pageNumber="215" specimenCount="1" stateProvince="Western Australia">
<collectingRegion id="49A5F85CFFC5FFC9FCD87D1B5C87B17E" box="[828,1000,545,566]" country="Australia" name="Western Australia" pageId="14" pageNumber="215">
<collectingRegion id="49A5F85CFFC5FFC9FCD87D1B5CE5B17E" box="[828,906,545,566]" country="American Samoa" name="Western" pageId="14" pageNumber="215">Western</collectingRegion>
Australia
</collectingRegion>
.............................................................................................
<taxonomicName id="4C614D3DFFC5FFC9FC747D045A70B11B" ID-CoL="3YF88" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" box="[912,1311,574,595]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="vanleeuweni" status="sp. nov.">
<emphasis id="B915EAACFFC5FFC9FC747D045BE7B11B" box="[912,1160,574,595]" italics="true" pageId="14" pageNumber="215">Matthewsius vanleeuweni</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A22657D7FFC5FFC9FACD7D045AF4B11B" box="[1321,1435,574,595]" pageId="14" pageNumber="215" rank="species">new species</taxonomicNameLabel>
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</keyLead>
</paragraph>
<paragraph id="8BDE36BEFFC5FFC9FF737D665AF3B1C6" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737D665AF3B1C6" pageId="14" pageNumber="215">- Clypeal edge without a projection lateral to clypeal teeth; elytral striae more-or-less crenulate, at least apically; protibiae with several serrations between apical and second tooth...........................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E1BFFC5FFC9FF737DAC5AF4B069" pageId="14" pageNumber="215">
<paragraph id="8BDE36BEFFC5FFC9FF737DAC5AF3B1AE" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737DAC5AF3B1AE" pageId="14" pageNumber="215">
4. Elytral surface smooth, intervals finely punctate, glabrous; male protibiae abruptly widened apically from second tooth; female pygidium not much more convex than that of male; male metatibiae slightly bent on inner side and slightly twisted; aedeagus as in
<figureCitation id="135A2A3BFFC5FFC9FEF07DEB5E25B1AE" box="[276,330,721,742]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
A;
<materialsCitation id="3B093CE3FFC5FFC9FE8D7DEB5AF3B1AE" ID-GBIF-Occurrence="1572325673" box="[361,1436,721,742]" location="New South Wales" pageId="14" pageNumber="215" specimenCount="1" stateProvince="New South Wales">
<collectingRegion id="49A5F85CFFC5FFC9FE8D7DEB5D7AB1AE" box="[361,533,721,742]" country="Australia" name="New South Wales" pageId="14" pageNumber="215">
New
<collectingRegion id="49A5F85CFFC5FFC9FE787DEB5D7AB1AE" box="[412,533,721,742]" inferredBySuffix="true" pageId="14" pageNumber="215">South Wales</collectingRegion>
</collectingRegion>
............................................
<taxonomicName id="4C614D3DFFC5FFC9FBCD7DEB5AF3B1AE" ID-CoL="3YF83" authority="Matthews" authorityName="Matthews" box="[1065,1436,721,742]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="illawarrensis">
<emphasis id="B915EAACFFC5FFC9FBCD7DEB5A48B1AE" box="[1065,1319,721,742]" italics="true" pageId="14" pageNumber="215">Matthewsius illawarrensis</emphasis>
(Matthews)
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="8BDE36BEFFC5FFC9FF737DD45AF4B069" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737DD45AF4B069" pageId="14" pageNumber="215">- Elytral surface slightly uneven, intervals with long or short setae; male protibiae gradually widened from base to apex; female pygidium markedly more convex than that of male; male metatibiae slightly bent on inner side but not twisted............5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30952E1BFFC5FFC9FF737C135AF4B099" pageId="14" pageNumber="215">
<paragraph id="8BDE36BEFFC5FFC9FF737C135AF3B031" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
<keyLead id="3090958BFFC5FFC9FF737C135AF3B031" pageId="14" pageNumber="215">
5. Elytral intervals with punctures bearing long, backwardly curving setae; setae as long as distance between punctures; punctures tending to become elongate laterally; striae deeply impressed; aedeagus as in
<figureCitation id="135A2A3BFFC5FFC9FC187C7C5B5AB013" box="[1020,1077,838,859]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
C;
<materialsCitation id="3B093CE3FFC5FFC9FBB57C7C5AF3B031" ID-GBIF-Occurrence="1572325610" location="New South Wales" pageId="14" pageNumber="215" specimenCount="1" stateProvince="New South Wales">
<collectingRegion id="49A5F85CFFC5FFC9FBB57C7C5B93B013" box="[1105,1276,838,859]" country="Australia" name="New South Wales" pageId="14" pageNumber="215">
New
<collectingRegion id="49A5F85CFFC5FFC9FB607C7C5B93B013" box="[1156,1276,838,859]" inferredBySuffix="true" pageId="14" pageNumber="215">South Wales</collectingRegion>
</collectingRegion>
..........................................................................................
<taxonomicName id="4C614D3DFFC5FFC9FB917C5E5AF3B031" ID-CoL="3YF85" authority="Matthews" authorityName="Matthews" box="[1141,1436,868,889]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="rossi">
<emphasis id="B915EAACFFC5FFC9FB917C5E5A49B031" box="[1141,1318,868,889]" italics="true" pageId="14" pageNumber="215">Matthewsius rossi</emphasis>
(Matthews)
</taxonomicName>
</materialsCitation>
</keyLead>
</paragraph>
<paragraph id="8BDE36BEFFC5FFC9FF737CBB5AF4B099" blockId="14.[151,1436,222,977]" pageId="14" pageNumber="215">
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- Elytral intervals with punctures bearing extremely short setae; setae barely longer than the width of a puncture; punctures mainly round laterally; striae less deeply impressed; aedeagus as in
<figureCitation id="135A2A3BFFC5FFC9FC8E7CA45CCDB0FB" box="[874,930,926,947]" captionStart="FIGURE 3" captionStartId="11.[151,250,1894,1916]" captionTargetBox="[164,1410,211,1857]" captionTargetId="figure@11.[163,1424,193,1872]" captionTargetPageId="11" captionText="FIGURE 3. Pygidium and aedeagus (left view, dorsal view, right view) of species of Matthewsius new genus, all scale bars 0.5 mm. A, Matthewsius illawarrensis new combination; B, M. penelopae new combination; C, M. rossi new combination; D, M. speewah new combination; E, M. stricklandensis new species; F, M. vanleeuweni new species." httpUri="https://zenodo.org/record/828971/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
E;
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New
<collectingRegion id="49A5F85CFFC5FFC9FC157CA45B03B0FB" box="[1009,1132,926,947]" inferredBySuffix="true" pageId="14" pageNumber="215">South Wales</collectingRegion>
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................................................................................
<taxonomicName id="4C614D3DFFC5FFC9FC9C7C865A70B099" ID-CoL="3YF87" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" box="[888,1311,956,977]" class="Insecta" family="Scarabaeidae" genus="Matthewsius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="stricklandensis" status="sp. nov.">
<emphasis id="B915EAACFFC5FFC9FC9C7C865BE7B099" box="[888,1160,956,977]" italics="true" pageId="14" pageNumber="215">Matthewsius stricklandensis</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
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</treatment>
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