261 lines
34 KiB
XML
261 lines
34 KiB
XML
<document id="420FA150330DB98D5006F2C6E052D486" ID-DOI="10.11646/palaeoentomology.6.2.12" ID-ISSN="2624-2834" ID-Zenodo-Dep="7929038" ID-ZooBank="C6C7A7D6-AD18-4F89-A0B4-3D538C718801" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683887917584" checkinUser="plazi" docAuthor="OYAMA, NOZOMU, SHINODA, KENJI, TAKAHASHI, HUMIO, DOI, EIJI & BÉTHOUX, OLIVIER" docDate="2023" docId="03C387B0FFC2F525DC030B4B24B6231C" docLanguage="en" docName="palaeoentomology.6.2.12.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.12" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Ideliopsina aristovi Oyama & Bethoux 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="208" masterDocId="FFFAFFC8FFC3F526DF290E5024542471" masterDocTitle="A new species of the Triassic genus Ideliopsina (Grylloblattida: Ideliidae) from the Ominé locality (Momonoki Formation, southwest Japan)" masterLastPageNumber="213" masterPageNumber="205" pageNumber="206" updateTime="1683889131799" updateUser="ExternalLinkService">
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<mods:title id="2DAF7320AC86FA4230DEE2A72A1DFE46">A new species of the Triassic genus Ideliopsina (Grylloblattida: Ideliidae) from the Ominé locality (Momonoki Formation, southwest Japan)</mods:title>
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<mods:namePart id="9B9CD88D646A087BEB952284B011B6BA">OYAMA, NOZOMU</mods:namePart>
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<mods:namePart id="0A47EEDF6D813CCCA536B0E4D14782E9">SHINODA, KENJI</mods:namePart>
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<mods:roleTerm id="14FF31216C78DC24D045FF1DF82CDF63">Author</mods:roleTerm>
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<mods:namePart id="3734A116E44B4FBE1CC5AF61C8961A57">TAKAHASHI, HUMIO</mods:namePart>
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<mods:namePart id="374E3D1A5257EEE9065F0670D820DCEF">DOI, EIJI</mods:namePart>
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<mods:namePart id="15A124E60C935E7C9B97B7B50232CAA4">BÉTHOUX, OLIVIER</mods:namePart>
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<mods:title id="571700BF7E9303BF60CAC1557F35F413">Palaeoentomology</mods:title>
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<mods:part id="9DE2FCC95680D6697CDBD354C27743FB">
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<mods:date id="C33E7BC1AB5A6B2A6D8AA6F521740BC3">2023</mods:date>
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<mods:number id="E331F261E2DB58919F8012467096E423">2023-04-28</mods:number>
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<mods:number id="F2270804B3FAD034CC9091498FACAA7B">6</mods:number>
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<mods:url id="90D9F4D50CB77AB91BBFDA6C39F2131F">http://dx.doi.org/10.11646/palaeoentomology.6.2.12</mods:url>
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<mods:classification id="1894F8DDF57F1AC2FA3C3AACE04E3BB0">journal article</mods:classification>
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<mods:identifier id="9FB1A1862EBEC99565D6E14EEFF82D40" type="DOI">10.11646/palaeoentomology.6.2.12</mods:identifier>
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<mods:identifier id="3C6408A930B7E1C676A809A34D8BD9E3" type="ISSN">2624-2834</mods:identifier>
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<mods:identifier id="57BB4E4F8423B2B23F91927295B17088" type="Zenodo-Dep">7929038</mods:identifier>
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<mods:identifier id="40852FEED5999197B4AC207EC25A9AC4" type="ZooBank">C6C7A7D6-AD18-4F89-A0B4-3D538C718801</mods:identifier>
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<subSubSection id="C370652DFFC2F527DC030B4B211B2147" box="[810,1359,1307,1334]" pageId="1" pageNumber="206" type="nomenclature">
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<paragraph id="8BD536A6FFC2F527DC030B4B211B2147" blockId="1.[810,1359,1307,1370]" box="[810,1359,1307,1334]" pageId="1" pageNumber="206">
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<heading id="D09D81CAFFC2F527DC030B4B211B2147" bold="true" box="[810,1359,1307,1334]" fontSize="10" level="2" pageId="1" pageNumber="206" reason="3">
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<emphasis id="B91EEAB4FFC2F527DC030B4B211B2147" bold="true" box="[810,1359,1307,1334]" pageId="1" pageNumber="206">
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<taxonomicName id="4C6A4D25FFC2F527DC030B4B20BB2147" authority="Oyama & Bethoux" authorityName="Oyama & Bethoux" authorityYear="2023" box="[810,1263,1307,1334]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
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<emphasis id="B91EEAB4FFC2F527DC030B4B205C2147" bold="true" box="[810,1032,1307,1334]" italics="true" pageId="1" pageNumber="206">Ideliopsina aristovi</emphasis>
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Oyama & Béthoux
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</taxonomicName>
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<taxonomicNameLabel id="A22D57CFFFC2F527DBDC0B4C211B2147" box="[1269,1359,1308,1334]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C370652DFFC2F527DC030B1027C9212B" box="[810,925,1344,1370]" pageId="1" pageNumber="206" type="description">
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<paragraph id="8BD536A6FFC2F527DC030B1027C9212B" blockId="1.[810,1359,1307,1370]" box="[810,925,1344,1370]" pageId="1" pageNumber="206">
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(
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<figureCitation id="13512A23FFC2F527DC1B0B1027C0212B" box="[818,916,1344,1370]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,235,1209,1233]" captionStartId-1="3.[136,235,1193,1217]" captionStartId-2="4.[136,235,1079,1103]" captionTargetBox-0="[331,1256,179,1185]" captionTargetBox-1="[330,1257,162,1159]" captionTargetBox-2="[145,1443,200,1054]" captionTargetId-0="figure-390@2.[331,1256,179,1185]" captionTargetId-1="figure-357@3.[330,1257,162,1169]" captionTargetId-2="figure-327@4.[145,1443,200,1055]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect). A, Photograph. B, Reflectance transforming imaging (RTI) extract in normals visualization (see Dryad Digital Repository)." captionText-1="FIGURE 2. Ideliopsina aristovi sp. nov., holotype specimen. A, Photograph (MMHF-3-00101, ventral aspect, under ethanol). B, Line drawing." captionText-2="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi-0="http://doi.org/10.5281/zenodo.7929040" figureDoi-1="http://doi.org/10.5281/zenodo.7929044" figureDoi-2="http://doi.org/10.5281/zenodo.7929046" httpUri-0="https://zenodo.org/record/7929040/files/figure.png" httpUri-1="https://zenodo.org/record/7929044/files/figure.png" httpUri-2="https://zenodo.org/record/7929046/files/figure.png" pageId="1" pageNumber="206">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C370652DFFC2F527DC030BD827B4219A" pageId="1" pageNumber="206" type="materials_examined">
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<paragraph id="8BD536A6FFC2F527DC030BD827B4219A" blockId="1.[810,1452,1416,2093]" pageId="1" pageNumber="206">
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<emphasis id="B91EEAB4FFC2F527DC030BD827CA21D3" bold="true" box="[810,926,1416,1442]" pageId="1" pageNumber="206">
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<typeStatus id="54D18804FFC2F527DC030BD827CD21D3" box="[810,921,1416,1442]" pageId="1" pageNumber="206">Holotype</typeStatus>
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.
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</emphasis>
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MMHF-3-00101, a right forewing with base and apical area incomplete (two sides, in dorsal and ventral aspects).
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</paragraph>
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</subSubSection>
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<subSubSection id="C370652DFFC2F527DC760BA527A0220A" pageId="1" pageNumber="206" type="etymology">
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<paragraph id="8BD536A6FFC2F527DC760BA527A0220A" blockId="1.[810,1452,1416,2093]" pageId="1" pageNumber="206">
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<emphasis id="B91EEAB4FFC2F527DC760BA527B2227E" bold="true" box="[863,998,1525,1551]" pageId="1" pageNumber="206">Etymology.</emphasis>
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The specific epithet honors the late Daniil S. Aristov, for his extensive contribution to our knowledge of the evolution of
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<taxonomicName id="4C6A4D25FFC2F527DB83086D211E2226" authorityName="Walker" authorityYear="1914" box="[1194,1354,1597,1623]" order="Grylloblattida" pageId="1" pageNumber="213" rank="order">Grylloblattida</taxonomicName>
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, and of insects in general.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C370652DFFC2F527DC7608D520CE23ED" pageId="1" pageNumber="206" type="diagnosis">
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<paragraph id="8BD536A6FFC2F527DC7608D5214D237A" blockId="1.[810,1452,1416,2093]" pageId="1" pageNumber="206">
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<emphasis id="B91EEAB4FFC2F527DC7608D5278D22EE" bold="true" box="[863,985,1669,1695]" pageId="1" pageNumber="206">Diagnosis.</emphasis>
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<taxonomicName id="4C6A4D25FFC2F527DCD108D520BF22EE" authority="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX, 2023" authorityName="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX" authorityYear="2023" box="[1016,1259,1669,1695]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="1" pageNumber="206" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
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<emphasis id="B91EEAB4FFC2F527DCD108D520BF22EE" box="[1016,1259,1669,1695]" italics="true" pageId="1" pageNumber="206">Ideliopsina aristovi</emphasis>
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</taxonomicName>
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<emphasis id="B91EEAB4FFC2F527DA2008D5212E22EE" bold="true" box="[1289,1402,1669,1695]" pageId="1" pageNumber="206">
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<taxonomicNameLabel id="A22D57CFFFC2F527DA2008D5212E22EE" box="[1289,1402,1669,1695]" pageId="1" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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is distinguished from the congeners by the following combination of characters: first fork of MA located basal to the first fork of RP; MP and CuP straight.
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</paragraph>
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<paragraph id="8BD536A6FFC2F527DC76094520CE23ED" blockId="1.[810,1452,1416,2093]" pageId="1" pageNumber="206">
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<emphasis id="B91EEAB4FFC2F527DC7609452034235E" bold="true" box="[863,1120,1813,1840]" pageId="1" pageNumber="206">Locality and horizon.</emphasis>
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Momonoki Formation (Mine Group); Ominé locality, Mine City area,
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<collectingRegion id="49AEF844FFC2F527DA04096A21FF2325" box="[1325,1451,1850,1876]" country="Japan" name="Yamaguchi" pageId="1" pageNumber="206">Yamaguchi</collectingRegion>
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(
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<collectingCountry id="F37D7636FFC2F527DC1B090E272C2309" box="[818,888,1886,1912]" name="Japan" pageId="1" pageNumber="206">Japan</collectingCountry>
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); Carnian (Upper Triassic;
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<bibRefCitation id="EFFB4B57FFC2F527DB9A090E21632308" author="Hase, A." box="[1203,1335,1886,1913]" pageId="1" pageNumber="206" pagination="72 - 89" refId="ref5282" refString="Hase, A. (1951) The Triassic in Yamaguchi Prefecture. In: Geological Survey of Japan (Ed.), Triassic stratigraphy of Japan, Geological Survey of Japan, pp. 72 - 89. https: // doi. org / 10.5575 / geosoc. 96.771" type="book chapter" year="1951">Hase, 1951</bibRefCitation>
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;
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<bibRefCitation id="EFFB4B57FFC2F527DA6F090E27FF23ED" author="Ishibashi, T. & Hashimoto, K. & Nohara, M. & Yoshida, K." pageId="1" pageNumber="206" pagination="771 - 774" refId="ref5406" refString="Ishibashi, T., Hashimoto, K., Nohara, M. & Yoshida, K. (1990) Triassic ammonites newly discovered in the western district of Yamaguchi Prefecture, Japan. Journal Geological Society of Japan, 96, 771 - 774." type="journal article" year="1990">
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Ishibashi
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<emphasis id="B91EEAB4FFC2F527DC0309D3270F23ED" box="[810,859,1922,1948]" italics="true" pageId="1" pageNumber="206">et al</emphasis>
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||
., 1990
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</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFFB4B57FFC2F527DC9F09D220DA23ED" author="Wakita, K. & Nakagawa, T. & Sakata, M. & Tanaka, N. & Oyama, N." box="[950,1166,1922,1948]" pageId="1" pageNumber="206" pagination="1 - 17" refId="ref6972" refString="Wakita, K., Nakagawa, T., Sakata, M., Tanaka, N. & Oyama, N. (2021) Phanerozoic accretionary history of Japan and the western pacific margin. Geological Magazine, 158, 1 - 17. https: // doi. org / 10.1017 / s 0016756818000742" type="journal article" year="2021">
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Wakita
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<emphasis id="B91EEAB4FFC2F527DB2509D3206923ED" box="[1036,1085,1922,1948]" italics="true" pageId="1" pageNumber="206">et al</emphasis>
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||
., 2021
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</bibRefCitation>
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||
).
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C370652DFFC2F524DC7609F6270D2190" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206" type="description">
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<paragraph id="8BD536A6FFC2F524DC7609F6270D2190" blockId="1.[810,1452,1416,2093]" lastBlockId="2.[810,1452,1332,2093]" lastPageId="2" lastPageNumber="207" pageId="1" pageNumber="206">
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<emphasis id="B91EEAB4FFC2F527DC7609F627A523B1" bold="true" box="[863,1009,1958,1984]" pageId="1" pageNumber="206">Description.</emphasis>
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Forewing
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<quantity id="4C929B43FFC2F527DB4209F6208423B0" box="[1131,1232,1958,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="1" pageNumber="206" unit="mm" value="10.3">10.3 mm</quantity>
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long, 4.0 mm wide (at its widest), without coloration pattern; anterior wing margin thin; in the median area of the wing, anterior and posterior wing margins not parallel, forming a low angle (10 degrees) instead; area between anterior wing margin and ScP broad (
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||
<quantity id="4C929B43FFC1F524DE6A0B0925CA2105" box="[323,414,1369,1396]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="207" unit="mm" value="3.6">3.6 mm</quantity>
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||
at its widest;
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||
<quantity id="4C929B43FFC1F524DD6E0B0926F52105" box="[583,673,1369,1396]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="207" unit="mm" value="3.3">3.3 mm</quantity>
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opposite the RA/RP fork); ScP mainly anteriorly pectinate, with 12 main veinlets, sometimes branched (up to three orders of branching, as preserved), with a total 18 veinlets reaching the wing margin (as preserved); RA/RP fork located distally to the MA/MP fork; RA simple, with one anterior veinlet preserved; RP branched
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<quantity id="4C929B43FFC1F524DEC1086526682221" box="[488,572,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="207" unit="mm" value="5.4">5.4 mm</quantity>
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distal to its origin, with a total of 4 branches (as preserved); MA simple for along distance (
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<quantity id="4C929B43FFC1F524DE5A082F25B722E8" box="[371,483,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="207" unit="mm" value="12.7">12.7 mm</quantity>
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), with at least one fork; MP desclerotized in the middle of its course (or, wing disrupted or creased in this area), simple, straight; MP– CuA area broader than other areas; CuA main posterior pectinate, with a strong main stem giving rise to 11 distal branches reaching the posterior wing margin, 1 distal veinlet vanishing in the cross-venation, and 4 proximal, well-developed veinlets vanishing in CuA–CuP area (or reaching CuP); CuP weak, straight; cross-venation forming (i) very small cells in the area between anterior wing margin and ScP (
|
||
<figureCitation id="13512A23FFC1F524DEAD09BE25892C79" box="[388,477,2030,2056]" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="2" pageNumber="207">Fig. 3A</figureCitation>
|
||
), (ii) strong, straight cross-veins connected by weaker ones in the ScP–RA area (
|
||
<figureCitation id="13512A23FFC1F524DC1B0B6427DD213F" box="[818,905,1332,1358]" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="2" pageNumber="207">Fig. 3B</figureCitation>
|
||
), (iii) S-shaped cross-veins near the base of CuA branches (
|
||
<figureCitation id="13512A23FFC1F524DC890B0927AE2102" box="[928,1018,1369,1395]" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="2" pageNumber="207">Fig. 3C</figureCitation>
|
||
), and (iv) a fine reticulation near the posterior wing margin (
|
||
<figureCitation id="13512A23FFC1F524DB100B2E20C221E9" box="[1081,1174,1406,1432]" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="2" pageNumber="207">Fig. 3D</figureCitation>
|
||
); posterior wing margin with crenulation (alternating bulges and depressions;
|
||
<figureCitation id="13512A23FFC1F524DA560BF227182190" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="2" pageNumber="207">Fig. 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF15662EFFC1F524DFA10AE9207B2084" ID-DOI="http://doi.org/10.5281/zenodo.7929040" ID-Zenodo-Dep="7929040" httpUri="https://zenodo.org/record/7929040/files/figure.png" pageId="2" pageNumber="207" startId="2.[136,235,1209,1233]" targetBox="[331,1256,179,1185]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BD536A6FFC1F524DFA10AE9207B2084" blockId="2.[136,1452,1209,1270]" pageId="2" pageNumber="207">
|
||
<emphasis id="B91EEAB4FFC1F524DFA10AE9255320A0" bold="true" box="[136,263,1209,1233]" pageId="2" pageNumber="207">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C6A4D25FFC1F524DE260AE9258320A0" authority="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX, 2023" authorityName="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX" authorityYear="2023" box="[271,471,1209,1233]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||
<emphasis id="B91EEAB4FFC1F524DE260AE9258320A0" box="[271,471,1209,1233]" italics="true" pageId="2" pageNumber="207">Ideliopsina aristovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEAB4FFC1F524DEC90AEA266120A0" bold="true" box="[480,565,1210,1234]" pageId="2" pageNumber="207">
|
||
<taxonomicNameLabel id="A22D57CFFFC1F524DEC90AEA266120A0" box="[480,565,1210,1234]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype specimen (MMHF-3-00101, dorsal aspect).
|
||
<emphasis id="B91EEAB4FFC1F524DB470AE920D420A0" bold="true" box="[1134,1152,1209,1233]" pageId="2" pageNumber="207">A</emphasis>
|
||
, Photograph.
|
||
<emphasis id="B91EEAB4FFC1F524DA3D0AEA217120A3" bold="true" box="[1300,1317,1210,1234]" pageId="2" pageNumber="207">B</emphasis>
|
||
, Reflectance transforming imaging (RTI) extract in normals visualization (see Dryad Digital Repository).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C370652DFFC1F525DC760BBC24B6231C" lastPageId="3" lastPageNumber="208" pageId="2" pageNumber="207" type="discussion">
|
||
<paragraph id="8BD536A6FFC1F525DC760BBC25B12217" blockId="2.[810,1452,1332,2093]" lastBlockId="3.[136,778,1312,1901]" lastPageId="3" lastPageNumber="208" pageId="2" pageNumber="207">
|
||
<emphasis id="B91EEAB4FFC1F524DC760BBC27872277" bold="true" box="[863,979,1516,1542]" pageId="2" pageNumber="207">Remarks.</emphasis>
|
||
Following
|
||
<bibRefCitation id="EFFB4B57FFC1F524DB720BBC21142277" author="Storozhenko, S. Y." box="[1115,1344,1516,1542]" pageId="2" pageNumber="207" refId="ref6733" refString="Storozhenko, S. Y. (1998) Sistematika, filogeniya i evolyutsiya grilloblattidovykh nasekomykh (Insecta: Grylloblattida). Vladivostok, Dal'nauka, 207 pp." type="book" year="1998">Storozhenko (1998)</bibRefCitation>
|
||
, the new species can be assigned to the family
|
||
<taxonomicName id="4C6A4D25FFC1F524DB970840214A225B" authorityName="Zalessky" authorityYear="1929" box="[1214,1310,1552,1578]" class="Insecta" family="Ideliidae" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="207" phylum="Arthropoda" rank="family">Ideliidae</taxonomicName>
|
||
owing to the following combination of character states: area between anterior wing margin and ScP broad; CuA with many branches; CuA not clearly differentiated into CuA1 and CuA2; CuA with posterior, proximal veinlets vanishing in the CuA–CuP area or reaching CuP; cross-venation very fine in the whole wing. Within this family, our specimen shares the diagnostic features of the genus
|
||
<taxonomicName id="4C6A4D25FFC1F524DA0F094121F0235A" authorityName="Storozhenko" authorityYear="1996" box="[1318,1444,1809,1835]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="2" pageNumber="207" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEAB4FFC1F524DA0F094121F0235A" box="[1318,1444,1809,1835]" italics="true" pageId="2" pageNumber="207">Ideliopsina</emphasis>
|
||
</taxonomicName>
|
||
, namely area between anterior wing margin and ScP very broad (about 2.7 times broader than maximum width of RA–RP area); RA simple (or, with mere distal anterior veinlets); R–M area (basal to the RA/RP fork) very narrow; MA straight; CuA proximal branches well-developed; all CuA branches directed towards the posterior wing margin (as opposed to ‘3–5 distal branches of CuA running alongside the posterior wing margin’). Compared with species belonging to
|
||
<taxonomicName id="4C6A4D25FFC0F525DEE90B15266A212E" authorityName="Storozhenko" authorityYear="1996" box="[448,574,1349,1375]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EEAB4FFC0F525DEE90B15266A212E" box="[448,574,1349,1375]" italics="true" pageId="3" pageNumber="208">Ideliopsina</emphasis>
|
||
</taxonomicName>
|
||
, the new species can be clearly distinguished from
|
||
<taxonomicName id="4C6A4D25FFC0F525DD0A0B3C26B121F4" authorityName="Aristov" authorityYear="2005" box="[547,741,1387,1413]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="kenderlykensis">
|
||
<emphasis id="B91EEAB4FFC0F525DD0A0B3C26B121F4" box="[547,741,1387,1413]" italics="true" pageId="3" pageNumber="208">I. kenderlykensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6A4D25FFC0F525DDD00B3C249421DB" authorityName="Storozhenko" authorityYear="1996" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="nana">
|
||
<emphasis id="B91EEAB4FFC0F525DDD00B3C249421DB" italics="true" pageId="3" pageNumber="208">I. nana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6A4D25FFC0F525DFE70BC1256421DB" authorityName="Storozhenko" authorityYear="1996" box="[206,304,1425,1450]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B91EEAB4FFC0F525DFE70BC1256421DB" box="[206,304,1425,1450]" italics="true" pageId="3" pageNumber="208">I. ornata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6A4D25FFC0F525DE450BC125BC21DB" authorityName="Storozhenko" authorityYear="1996" box="[364,488,1424,1451]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="stupenda">
|
||
<emphasis id="B91EEAB4FFC0F525DE450BC1252121DB" box="[364,373,1425,1450]" italics="true" pageId="3" pageNumber="208">I</emphasis>
|
||
.
|
||
<emphasis id="B91EEAB4FFC0F525DEAA0BC025BC21DB" box="[387,488,1424,1450]" italics="true" pageId="3" pageNumber="208">stupenda</emphasis>
|
||
</taxonomicName>
|
||
owing to first fork of MA located basal to the first fork of RP. Moreover, unlike
|
||
<taxonomicName id="4C6A4D25FFC0F525DDD00BE725792184" authorityName="Aristov" authorityYear="2005" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="kenderlykensis">
|
||
<emphasis id="B91EEAB4FFC0F525DDD00BE725792184" italics="true" pageId="3" pageNumber="208">I. kenderlykensis</emphasis>
|
||
</taxonomicName>
|
||
, it has a straight CuP. The new species resembles
|
||
<taxonomicName id="4C6A4D25FFC0F525DE2A085225D6226A" authorityName="Storozhenko" authorityYear="1996" box="[259,386,1537,1563]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="ruginosa">
|
||
<emphasis id="B91EEAB4FFC0F525DE2A08522558226A" box="[259,268,1538,1563]" italics="true" pageId="3" pageNumber="208">I</emphasis>
|
||
.
|
||
<emphasis id="B91EEAB4FFC0F525DE37085225D6226A" box="[286,386,1538,1563]" italics="true" pageId="3" pageNumber="208">ruginosa</emphasis>
|
||
</taxonomicName>
|
||
in the respective positions of the first forks of RP and MA but differs from it by a larger forewing, with a straighter MP.
|
||
</paragraph>
|
||
<caption id="DF15662EFFC0F525DFA10AF925432097" ID-DOI="http://doi.org/10.5281/zenodo.7929044" ID-Zenodo-Dep="7929044" httpUri="https://zenodo.org/record/7929044/files/figure.png" pageId="3" pageNumber="208" startId="3.[136,235,1193,1217]" targetBox="[330,1257,162,1159]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BD536A6FFC0F525DFA10AF925432097" blockId="3.[136,1451,1193,1254]" pageId="3" pageNumber="208">
|
||
<emphasis id="B91EEAB4FFC0F525DFA10AF9255020B3" bold="true" box="[136,260,1193,1218]" pageId="3" pageNumber="208">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6A4D25FFC0F525DE230AFA258420B0" authority="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX, 2023" authorityName="OYAMA & SHINODA & TAKAHASHI & DOI & BÉTHOUX" authorityYear="2023" box="[266,464,1194,1218]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
|
||
<emphasis id="B91EEAB4FFC0F525DE230AFA258420B0" box="[266,464,1194,1218]" italics="true" pageId="3" pageNumber="208">Ideliopsina aristovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91EEAB4FFC0F525DEFE0AFA267D20B0" bold="true" box="[471,553,1194,1218]" pageId="3" pageNumber="208">
|
||
<taxonomicNameLabel id="A22D57CFFFC0F525DEFE0AFA267D20B0" box="[471,553,1194,1218]" pageId="3" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype specimen.
|
||
<emphasis id="B91EEAB4FFC0F525DC2B0AF9274020B0" bold="true" box="[770,788,1193,1217]" pageId="3" pageNumber="208">A</emphasis>
|
||
, Photograph (MMHF-3-00101, ventral aspect, under ethanol).
|
||
<emphasis id="B91EEAB4FFC0F525DABD0AFA21F120B3" bold="true" box="[1428,1445,1194,1218]" pageId="3" pageNumber="208">B</emphasis>
|
||
, Line drawing.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD536A6FFC0F525DF94082224B6231C" blockId="3.[136,778,1312,1901]" pageId="3" pageNumber="208">
|
||
Structures reminiscent of trichosors, as occurring in Neuroptera, have been reported by
|
||
<bibRefCitation id="EFFB4B57FFC0F525DD0E08C7275E22C0" author="Storozhenko, S. Y." box="[551,778,1687,1713]" pageId="3" pageNumber="208" pagination="13 - 19" refId="ref6701" refString="Storozhenko, S. Y. (1996) New Triassic grylloblattids of the family Ideliidae (Plecopteroidea, Grylloblattida). Russian Entomological Journal, 5, 13 - 19." type="journal article" year="1996">Storozhenko (1996)</bibRefCitation>
|
||
in
|
||
<taxonomicName id="4C6A4D25FFC0F525DF8F08ED255D22A7" authorityName="Storozhenko" authorityYear="1996" box="[166,265,1725,1751]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B91EEAB4FFC0F525DF8F08ED24FB22A7" box="[166,175,1725,1750]" italics="true" pageId="3" pageNumber="208">I</emphasis>
|
||
.
|
||
<emphasis id="B91EEAB4FFC0F525DF9708ED255D22A7" box="[190,265,1725,1750]" italics="true" pageId="3" pageNumber="208">ornata</emphasis>
|
||
</taxonomicName>
|
||
. In the new material we observed bulges and depressions alternating along the posterior wing margin (
|
||
<figureCitation id="13512A23FFC0F525DFB9095725592353" box="[144,269,1799,1826]" captionStart="FIGURE 3" captionStartId="4.[136,235,1079,1103]" captionTargetBox="[145,1443,200,1054]" captionTargetId="figure-327@4.[145,1443,200,1055]" captionTargetPageId="4" captionText="FIGURE 3. Ideliopsina aristovi sp. nov., holotype specimen (MMHF-3-00101, dorsal aspect), morphological details (except for E, locations as indicated in Fig. 1A). A–C, Details of cross-venation. A, Detail of the area between the anterior wing margin and ScP. B, Detail of the area between ScP and RP. C, Detail of the area between CuA branches (substantiated by dotted lines; ‘CuA’ label indicating the main stem of this vein). D, E, Detail of the posterior wing margin showing crenulation. E, Detail as indicated in D." figureDoi="http://doi.org/10.5281/zenodo.7929046" httpUri="https://zenodo.org/record/7929046/files/figure.png" pageId="3" pageNumber="208">Fig. 3D, E</figureCitation>
|
||
), we propose to term ‘crenulation’. Whether this structure is similar to that known in
|
||
<taxonomicName id="4C6A4D25FFC0F525DD74097E26952336" authorityName="Storozhenko" authorityYear="1996" box="[605,705,1838,1863]" class="Insecta" family="Ideliidae" genus="Ideliopsina" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="208" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="B91EEAB4FFC0F525DD74097E26952336" box="[605,705,1838,1863]" italics="true" pageId="3" pageNumber="208">I. ornata</emphasis>
|
||
</taxonomicName>
|
||
is not evident.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |