404 lines
53 KiB
XML
404 lines
53 KiB
XML
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<mods:title id="2616CB775FF26501B62868C6203E65EA">The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species</mods:title>
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<mods:namePart id="EC686EAC03BDEED3DB84A4EDFE8BDB2E">Valdés, Ángel</mods:namePart>
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<mods:affiliation id="0CB63ADE75EB5090B68C0192A0511D86">Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & aavaldes @ cpp. edu; https: // orcid. org / 0000 - 0002 - 2347 - 4896</mods:affiliation>
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<mods:namePart id="3495E97C82D0B857ACCB264B3B0ACC50">Feliciano, Kendall</mods:namePart>
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<mods:affiliation id="037E978389C1DBC221DAC65DF9DFBD2E">Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & https: // orcid. org / 0000 - 0003 - 0536 - 2628</mods:affiliation>
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<mods:namePart id="21405A609E26350D80E81689C1F65198">Malaquias, Manuel A. E.</mods:namePart>
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<mods:affiliation id="B8A8938860828B8D41BEF66DB8CD9D88">Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, N- 5020 Bergen, Norway. https: // orcid. org / 0000 - 0002 - 9668 - 945 X</mods:affiliation>
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<mods:title id="4405FB6E31DD1092536C3E2A7CC38FC6">Zootaxa</mods:title>
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<treatment id="03C38798FF9F467C8ED72D8F4091FBCB" ID-DOI="http://doi.org/10.5281/zenodo.7860384" ID-GBIF-Taxon="209569267" ID-Zenodo-Dep="7860384" LSID="urn:lsid:plazi:treatment:03C38798FF9F467C8ED72D8F4091FBCB" httpUri="http://treatment.plazi.org/id/03C38798FF9F467C8ED72D8F4091FBCB" lastPageId="19" lastPageNumber="490" pageId="18" pageNumber="489">
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<paragraph id="8BD5368EFF9F467D8ED72D8F40C6FE1A" blockId="18.[151,508,402,465]" box="[151,508,402,429]" pageId="18" pageNumber="489">
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<heading id="D09D81E2FF9F467D8ED72D8F40C6FE1A" bold="true" box="[151,508,402,429]" fontSize="11" level="1" pageId="18" pageNumber="489" reason="1">
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<taxonomicName id="4C6A4D0DFF9F467D8ED72D8F40C6FE1A" ID-CoL="4QJL9" authority="(Reeve, 1842)" baseAuthorityName="Reeve" baseAuthorityYear="1842" box="[151,508,402,429]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
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<emphasis id="B91EEA9CFF9F467D8ED72D8F40C6FE1A" bold="true" box="[151,508,402,429]" pageId="18" pageNumber="489">
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<emphasis id="B91EEA9CFF9F467D8ED72D8F4077FE1B" bold="true" box="[151,333,402,429]" italics="true" pageId="18" pageNumber="489">Pupa tessellata</emphasis>
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(Reeve, 1842)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3706505FF9F467D8ED72DAB41C0FE67" box="[151,250,439,465]" pageId="18" pageNumber="489" type="description">
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<paragraph id="8BD5368EFF9F467D8ED72DAB41C0FE67" blockId="18.[151,508,402,465]" box="[151,250,439,465]" pageId="18" pageNumber="489">
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(
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<figureCitation id="13512A0BFF9F467D8EDF2DAB41CBFE67" box="[159,241,439,465]" captionStart="FIGURE 12" captionStartId="18.[152,255,1693,1717]" captionTargetBox="[151,1436,1121,1669]" captionTargetId="figure-345@18.[151,1436,1121,1669]" captionTargetPageId="18" captionText="FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP." figureDoi="http://doi.org/10.5281/zenodo.7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="18" pageNumber="489">Fig. 12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3706505FF9F467D8ED72DE74471FDE6" pageId="18" pageNumber="489" type="reference_group">
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<paragraph id="8BD5368EFF9F467D8ED72DE74413FDA5" blockId="18.[151,1389,507,593]" box="[151,1321,507,532]" pageId="18" pageNumber="489">
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<treatmentCitationGroup id="AB7A11A0FF9F467D8ED72DE74413FDA5" box="[151,1321,507,532]" pageId="18" pageNumber="489">
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<treatmentCitation id="0ACB109FFF9F467D8ED72DE74311FDA5" author="Reeve, L. A." box="[151,555,507,531]" page="147" pageId="18" pageNumber="489" year="1842">
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<taxonomicName id="4C6A4D0DFF9F467D8ED72DE74311FDA5" ID-CoL="9c8d552a-ebd5-4bb3-a190-76f237cddc40" authority="Reeve, 1842 a: 147" authorityName="Reeve" authorityPageNumber="147" authorityYear="1842" box="[151,555,507,531]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
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<emphasis id="B91EEA9CFF9F467D8ED72DE7405CFDA5" box="[151,358,507,531]" italics="true" pageId="18" pageNumber="489">Tornatella tessellata</emphasis>
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<bibRefCitation id="EFFB4B7FFF9F467D8F2C2DE04311FDA5" author="Reeve, L. A." box="[364,555,507,531]" pageId="18" pageNumber="489" pagination="58 - 62" refId="ref25717" refString="Reeve, L. A. (1842 a) Monograph of the genus Tornatella, a small group of Pectinibranchiate Mollusks of the family Plicacea, including descriptions of seven new species, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 10, 58 - 62." type="journal article" year="1842">Reeve, 1842a: 147</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, pl. 206, fig. 3; 1842b: 60.
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<typeStatus id="54D1882CFF9F467D8D7E2DE0424AFDA2" box="[830,880,508,532]" pageId="18" pageNumber="489">Type</typeStatus>
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locality: Sinum Persicum [= Persian Gulf].
|
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD5368EFF9F467D8ED72E064457FD84" blockId="18.[151,1389,507,593]" box="[151,1389,538,564]" pageId="18" pageNumber="489">
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<treatmentCitationGroup id="AB7A11A0FF9F467D8ED72E064457FD84" box="[151,1389,538,564]" pageId="18" pageNumber="489">
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?
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<taxonomicName id="4C6A4D0DFF9F467D8EE22E0640F3FD84" ID-CoL="7CHF4" authority="Souverbie" authorityName="Souverbie" authorityYear="1863" box="[162,457,538,562]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="alveola">
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||
<emphasis id="B91EEA9CFF9F467D8EE22E064066FD84" box="[162,348,538,562]" italics="true" pageId="18" pageNumber="489">Tornatella alveola</emphasis>
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||
Souverbie
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</taxonomicName>
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||
<emphasis id="B91EEA9CFF9F467D8F8F2E0640D9FD87" box="[463,483,538,561]" italics="true" pageId="18" pageNumber="489">in</emphasis>
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||
<treatmentCitation id="0ACB109FFF9F467D8FA92E064259FD84" author="Souverbie, S. - M. & Montrouzier, X." box="[489,867,538,562]" page="167" pageId="18" pageNumber="489" year="1863">
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||
<bibRefCitation id="EFFB4B7FFF9F467D8FA92E064259FD84" author="Souverbie, S. - M. & Montrouzier, X." box="[489,867,538,562]" pageId="18" pageNumber="489" pagination="161 - 176" refId="ref26745" refString="Souverbie, S. - M. & Montrouzier, X. (1863) Descriptions d'especes nouvelles de l'archipel caledonien, et des iles Salomon et Woodlark. Journal de Conchyliologie, Series 3, 3, 161 - 176, pl. 5." type="journal article" year="1863">Souverbie & Montrouzier, 1863: 167</bibRefCitation>
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</treatmentCitation>
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, pl. 5, fig. 9.
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<typeStatus id="54D1882CFF9F467D8DA92E064521FD84" box="[1001,1051,538,562]" pageId="18" pageNumber="489">Type</typeStatus>
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locality: Île Art,
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||
<collectingCountry id="F37D761EFF9F467D8A892E064453FD84" box="[1225,1385,538,562]" name="New Caledonia" pageId="18" pageNumber="489">New Caledonia</collectingCountry>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD5368EFF9F467D8ED72E254471FDE6" blockId="18.[151,1389,507,593]" box="[151,1355,568,593]" pageId="18" pageNumber="489">
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<treatmentCitationGroup id="AB7A11A0FF9F467D8ED72E254471FDE6" box="[151,1355,568,593]" pageId="18" pageNumber="489">
|
||
?
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<taxonomicName id="4C6A4D0DFF9F467D8EE22E24436BFDE6" ID-CoL="4Y7M6" authority="Pilsbry, 1917: 214-215" authorityName="Pilsbry" authorityPageNumber="214 - 215" authorityYear="1917" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[162,593,568,593]" class="Gastropoda" family="Acteonidae" genus="Solidula" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis id="B91EEA9CFF9F467D8EE22E2441CCFDE6" box="[162,246,568,592]" italics="true" pageId="18" pageNumber="489">Solidula</emphasis>
|
||
<treatmentCitation id="0ACB109FFF9F467D8EBD2E24436BFDE6" author="Pilsbry, H. A." box="[253,593,568,593]" page="215" pageId="18" pageNumber="489" year="1917">
|
||
<emphasis id="B91EEA9CFF9F467D8EBD2E244067FDE6" box="[253,349,568,592]" italics="true" pageId="18" pageNumber="489">thaanumi</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8F232E24436BFDE6" author="Pilsbry, H. A." box="[355,593,568,593]" pageId="18" pageNumber="489" pagination="207 - 230" refId="ref25559" refString="Pilsbry, H. A. (1917) Marine Mollusks of Hawaii, I - III. Proceedings of the Academy of Natural Sciences of Philadelphia, 69, 207 - 230, pls. 14 - 15." type="journal article" year="1917">Pilsbry, 1917: 214–215</bibRefCitation>
|
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</treatmentCitation>
|
||
</taxonomicName>
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||
, text fig. 1.
|
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<typeStatus id="54D1882CFF9F467D8C892E2543C1FDE7" box="[713,763,569,593]" pageId="18" pageNumber="489">Type</typeStatus>
|
||
locality: Off Honolulu, Hawaiian Islands,
|
||
<quantity id="4C929B6BFF9F467D8AED2E24443FFDE6" box="[1197,1285,568,592]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="18" pageNumber="489" unit="m" value="13.0" valueMax="15.0" valueMin="11.0">11–15 m</quantity>
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depth.
|
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</treatmentCitationGroup>
|
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</paragraph>
|
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</subSubSection>
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<subSubSection id="C3706505FF9F467D8ED72E63458FF83B" pageId="18" pageNumber="489" type="materials_examined">
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<paragraph id="8BD5368EFF9F467D8ED72E634087FD57" blockId="18.[151,1437,638,1097]" pageId="18" pageNumber="489">
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<emphasis id="B91EEA9CFF9F467D8ED72E63407EFD2E" bold="true" box="[151,324,638,665]" pageId="18" pageNumber="489">Type material</emphasis>
|
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.
|
||
<taxonomicName id="4C6A4D0DFF9F467D8F192E63437EFD2F" authorityName="Reeve" authorityYear="1842" box="[345,580,639,665]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
|
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<emphasis id="B91EEA9CFF9F467D8F192E63437EFD2F" box="[345,580,639,665]" italics="true" pageId="18" pageNumber="489">Tornatella tessellata</emphasis>
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</taxonomicName>
|
||
—
|
||
<specimenCount id="9D6CFD07FF9F467D8C202E6343DAFD2F" box="[608,736,639,665]" pageId="18" pageNumber="489" type="generic" typeStatus="syntypes">4 syntypes</specimenCount>
|
||
, dry shells,
|
||
<quantity id="4C929B6BFF9F467D8D382E6342D3FD2F" box="[888,1001,639,665]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="18" pageNumber="489" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
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long (
|
||
<collectionCode id="ED7BAE4BFF9F467D8A7D2E634593FD2F" box="[1085,1193,639,665]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="18" pageNumber="489" type="Museum">NHMUK</collectionCode>
|
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196954);
|
||
<taxonomicName id="4C6A4D0DFF9F467D8B692E6341D3FD0B" authorityName="Souverbie" authorityYear="1863" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="alveola">
|
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<emphasis id="B91EEA9CFF9F467D8B692E6341D3FD0B" italics="true" pageId="18" pageNumber="489">Tornatella alveola</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<typeStatus id="54D1882CFF9F467D8F472EBF404BFD0B" box="[263,369,675,701]" pageId="18" pageNumber="489" type="holotype">Holotype</typeStatus>
|
||
, dry shell,
|
||
<quantity id="4C929B6BFF9F467D8FAC2EBF4300FD0B" box="[492,570,675,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="489" unit="mm" value="11.0">11 mm</quantity>
|
||
long (MHNBx 2004.TY.29);
|
||
<emphasis id="B91EEA9CFF9F467D8DC82EBF456DFD0B" box="[904,1111,675,701]" italics="true" pageId="18" pageNumber="489">
|
||
<taxonomicName id="4C6A4D0DFF9F467D8DC82EBF42DFFD0B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[904,997,675,701]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="solidula">Solidula</taxonomicName>
|
||
<taxonomicName id="4C6A4D0DFF9F467D8DAD2EBF456DFD0B" authorityName="Pilsbry" authorityYear="1917" box="[1005,1111,675,701]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="thaanumi">thaanumi</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<specimenCount id="9D6CFD07FF9F467D8A332EBF45D7FD0B" box="[1139,1261,675,701]" pageId="18" pageNumber="489" type="generic" typeStatus="syntypes">2 syntypes</specimenCount>
|
||
, dry shells,
|
||
<quantity id="4C929B6BFF9F467D8B322EBF41F9FD57" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="18" pageNumber="489" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6–9 mm</quantity>
|
||
long (
|
||
<collectionCode id="ED7BAE4BFF9F467D8F4A2EDB4068FD57" box="[266,338,711,737]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="18" pageNumber="489">ANSP</collectionCode>
|
||
117069).
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9F467D8E872EF74026FBFF" blockId="18.[151,1437,638,1097]" pageId="18" pageNumber="489">
|
||
<emphasis id="B91EEA9CFF9F467D8E872EF740F3FCB3" bold="true" box="[199,457,747,773]" pageId="18" pageNumber="489">External morphology</emphasis>
|
||
. Live animal unknow. Shell solid, narrow, elongate, widest at midlenght, with convex sides, rounded to elongate anterior end (
|
||
<figureCitation id="13512A0BFF9F467D8C252F134380FC9F" box="[613,698,783,809]" captionStart="FIGURE 12" captionStartId="18.[152,255,1693,1717]" captionTargetBox="[151,1436,1121,1669]" captionTargetId="figure-345@18.[151,1436,1121,1669]" captionTargetPageId="18" captionText="FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP." figureDoi="http://doi.org/10.5281/zenodo.7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="18" pageNumber="489">Fig. 12</figureCitation>
|
||
). Body whorl large, about 3/4–4/5 of total length. Spire conical, with 4 whorls. Suture channeled. Aperture elongate, wider anteriorly, narrowing gradually towards posterior end, ending at 1/5 of the posterior end of first whorl. Columellar margin thickened, slightly oblique, with large, channeled anterior fold starting near anterior end of aperture; smaller, simple posterior fold located close to aperture mid-length, separated from anterior fold by short gap. Apex of all specimens examined damaged, protoconch not observed. Sculpture composed of numerous punctuated spiral grooves. Punctuations conspicuous, irregular, oval, situated next to each other, often fused together, within each groove. Grooves separated by gaps wider than grooves. Shell color cream with irregular, often zigzagging pale brown to pink patches, separated by gaps similar in size to the patches.
|
||
</paragraph>
|
||
<caption id="DF156606FF9F467D8ED82A81425AF895" ID-DOI="http://doi.org/10.5281/zenodo.7860394" ID-Zenodo-Dep="7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="18" pageNumber="489" startId="18.[152,255,1693,1717]" targetBox="[151,1436,1121,1669]" targetPageId="18" targetType="figure">
|
||
<paragraph id="8BD5368EFF9F467D8ED82A81425AF895" blockId="18.[151,1436,1693,1827]" pageId="18" pageNumber="489">
|
||
<emphasis id="B91EEA9CFF9F467D8ED82A81401AF903" bold="true" box="[152,288,1693,1717]" pageId="18" pageNumber="489">FIGURE 12</emphasis>
|
||
. Photographs of shell specimens of
|
||
<taxonomicName id="4C6A4D0DFF9F467D8CDB2A8242E8F903" authority="(Reeve, 1842)" baseAuthorityName="Reeve" baseAuthorityYear="1842" box="[667,978,1693,1717]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9F467D8CDB2A824203F903" box="[667,825,1693,1717]" italics="true" pageId="18" pageNumber="489">Pupa tessellata</emphasis>
|
||
(Reeve, 1842)
|
||
</taxonomicName>
|
||
and possible synonyms, ventral view.
|
||
<emphasis id="B91EEA9CFF9F467D8B252A8144ACF903" bold="true" box="[1381,1430,1693,1717]" pageId="18" pageNumber="489">A–C</emphasis>
|
||
. Three syntypes of
|
||
<taxonomicName id="4C6A4D0DFF9F467D8F1E2ADD4386F96F" authority="Reeve, 1842" authorityName="Reeve" authorityYear="1842" box="[350,700,1729,1753]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9F467D8F1E2ADD430AF96F" box="[350,560,1729,1753]" italics="true" pageId="18" pageNumber="489">Tornatella tessellata</emphasis>
|
||
Reeve, 1842
|
||
</taxonomicName>
|
||
(NHMUK 196954), ©NHMUK, photos by K. Webb.
|
||
<emphasis id="B91EEA9CFF9F467D8ABD2ADD4435F96F" bold="true" box="[1277,1295,1729,1753]" pageId="18" pageNumber="489">D</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C6A4D0DFF9F467D8ED72AF940F8F94B" authority="Souverbie" authorityName="Souverbie" authorityYear="1863" box="[151,450,1765,1789]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9F467D8ED72AF94069F94B" box="[151,339,1765,1789]" italics="true" pageId="18" pageNumber="489">Tornatella alveola</emphasis>
|
||
Souverbie
|
||
</taxonomicName>
|
||
<emphasis id="B91EEA9CFF9F467D8F892AFA40E7F94B" box="[457,477,1766,1789]" italics="true" pageId="18" pageNumber="489">in</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8FA52AF94215F94B" author="Souverbie, S. - M. & Montrouzier, X." box="[485,815,1765,1789]" pageId="18" pageNumber="489" pagination="161 - 176" refId="ref26745" refString="Souverbie, S. - M. & Montrouzier, X. (1863) Descriptions d'especes nouvelles de l'archipel caledonien, et des iles Salomon et Woodlark. Journal de Conchyliologie, Series 3, 3, 161 - 176, pl. 5." type="journal article" year="1863">Souverbie & Montrouzier, 1863</bibRefCitation>
|
||
(MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles.
|
||
<emphasis id="B91EEA9CFF9F467D8BC52AF944ACF94B" bold="true" box="[1413,1430,1765,1789]" pageId="18" pageNumber="489">E</emphasis>
|
||
. Syntype of
|
||
<taxonomicName id="4C6A4D0DFF9F467D8F4C2B15405AF897" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[268,352,1801,1825]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="solidula">
|
||
<emphasis id="B91EEA9CFF9F467D8F4C2B15405AF897" box="[268,352,1801,1825]" italics="true" pageId="18" pageNumber="489">Solidula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C6A4D0DFF9F467D8F272B154369F897" authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[359,595,1801,1825]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis id="B91EEA9CFF9F467D8F272B1540FDF897" box="[359,455,1801,1825]" italics="true" pageId="18" pageNumber="489">thaanumi</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8F8D2B154369F897" author="Pilsbry, H. A." box="[461,595,1801,1825]" pageId="18" pageNumber="489" pagination="207 - 230" refId="ref25559" refString="Pilsbry, H. A. (1917) Marine Mollusks of Hawaii, I - III. Proceedings of the Academy of Natural Sciences of Philadelphia, 69, 207 - 230, pls. 14 - 15." type="journal article" year="1917">Pilsbry, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
(ANSP 117069), ©ANSP.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5368EFF9F467D8E872B53458FF83B" blockId="18.[151,1437,1871,2042]" pageId="18" pageNumber="489">
|
||
<emphasis id="B91EEA9CFF9F467D8E872B53409BF8DF" bold="true" box="[199,417,1871,1897]" pageId="18" pageNumber="489">Geographic range</emphasis>
|
||
. Widespread in the Indo-Pacific region from East Africa to the West Pacific Ocean and possibly the Hawaiian Islands (
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8FB52B6F4356F83B" author="Kay, A. E." box="[501,620,1907,1933]" pageId="18" pageNumber="489" pagination="1 - 653" refId="ref24228" refString="Kay, A. E. (1979) Hawaiian marine shells. Reef and shore fauna of Hawaii: Section 4: Mollusca. Bishop Museum Special Publication, 64, 1 - 653." type="journal article" year="1979">Kay, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8C372B684234F83B" author="Yonow, N." box="[631,782,1907,1934]" pageId="18" pageNumber="489" refId="ref27463" refString="Yonow, N. (2008) Sea Slugs of the Red Sea. Pensoft, Sofia, 303 pp." type="book" year="2008">Yonow, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8D5A2B6F4538F83B" author="Gosliner, T. & Valdes, A. & Behrens, D. W." box="[794,1026,1907,1934]" pageId="18" pageNumber="489" refId="ref23324" refString="Gosliner, T., Valdes, A. & Behrens, D. W. (2018) Nudibranch and Sea Slug Identification: Indo-Pacific. New World Publications Inc., Jacksonville, Florida, 451 pp." type="book" year="2018">
|
||
Gosliner
|
||
<emphasis id="B91EEA9CFF9F467D8DC12B684280F83B" box="[897,954,1907,1933]" italics="true" pageId="18" pageNumber="489">et al.</emphasis>
|
||
, 2018
|
||
</bibRefCitation>
|
||
; present paper).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3706505FF9F467C8E872B8B4091FBCB" lastPageId="19" lastPageNumber="490" pageId="18" pageNumber="489" type="discussion">
|
||
<paragraph id="8BD5368EFF9F467C8E872B8B426BFE3F" blockId="18.[151,1437,1871,2042]" lastBlockId="19.[151,1437,151,1149]" lastPageId="19" lastPageNumber="490" pageId="18" pageNumber="489">
|
||
<emphasis id="B91EEA9CFF9F467D8E872B8B400EF807" bold="true" box="[199,308,1943,1969]" pageId="18" pageNumber="489">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8F062B8440DDF807" author="Reeve, L. A." box="[326,487,1943,1970]" pageId="18" pageNumber="489" pagination="58 - 62" refId="ref25717" refString="Reeve, L. A. (1842 a) Monograph of the genus Tornatella, a small group of Pectinibranchiate Mollusks of the family Plicacea, including descriptions of seven new species, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 10, 58 - 62." type="journal article" year="1842">Reeve (1842a</bibRefCitation>
|
||
, b) introduced the new species
|
||
<taxonomicName id="4C6A4D0DFF9F467D8D1D2B8B45DAF807" authority="Reeve, 1842" authorityName="Reeve" authorityYear="1842" box="[861,1248,1943,1970]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="18" pageNumber="489" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9F467D8D1D2B8B457CF807" box="[861,1094,1943,1969]" italics="true" pageId="18" pageNumber="489">Tornatella tessellata</emphasis>
|
||
Reeve, 1842
|
||
</taxonomicName>
|
||
based on shells collected by Dr. Ŗppell from the Red Sea, although the habitat is indicated as Sinum Persicum [= Persian
|
||
<collectingRegion id="49AEF86CFF9F467D8B192BA744A8F863" box="[1369,1426,1979,2005]" country="Papua New Guinea" name="Gulf" pageId="18" pageNumber="489">Gulf</collectingRegion>
|
||
].
|
||
<bibRefCitation id="EFFB4B7FFF9F467D8ED72BFC4005F84F" author="Reeve, L. A." box="[151,319,2015,2042]" pageId="18" pageNumber="489" pagination="58 - 62" refId="ref25717" refString="Reeve, L. A. (1842 a) Monograph of the genus Tornatella, a small group of Pectinibranchiate Mollusks of the family Plicacea, including descriptions of seven new species, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 10, 58 - 62." type="journal article" year="1842">Reeve (1842a)</bibRefCitation>
|
||
described these shells as “finely striated in a transverse direction, and the interstices are neatly tessellated with numerous pale flesh-coloured square spots.” The examination of photographs of
|
||
<specimenCount id="9D6CFD07FF9E467C8B412C8B44A6FF07" box="[1281,1436,151,177]" pageId="19" pageNumber="490" type="generic" typeStatus="syntypes">four syntypes</specimenCount>
|
||
(NHMUK 196954), three of them illustrated herein (
|
||
<figureCitation id="13512A0BFF9E467C8C962CA7425FFF63" box="[726,869,187,213]" captionStart="FIGURE 12" captionStartId="18.[152,255,1693,1717]" captionTargetBox="[151,1436,1121,1669]" captionTargetId="figure-345@18.[151,1436,1121,1669]" captionTargetPageId="18" captionText="FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP." figureDoi="http://doi.org/10.5281/zenodo.7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="19" pageNumber="490">Figs. 12A–C</figureCitation>
|
||
) revealed the shells of this species are characterized by having a cream to light brown background color with a checkered pattern of darker brown rectangular or irregular markings all over. The shells are solid, oval, widest mid-length, with convex sides, with a rounded anterior end and an elongate, conical spire with 3–4 whorls. None of the specimens here examined and sequenced match this description, therefore we assume that
|
||
<taxonomicName id="4C6A4D0DFF9E467C8C7A2D574385FED3" authorityName="Reeve" authorityYear="1842" box="[570,703,331,357]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8C7A2D574385FED3" box="[570,703,331,357]" italics="true" pageId="19" pageNumber="490">T. tessellata</emphasis>
|
||
</taxonomicName>
|
||
constitute a distinct species, currently regarded as a valid member of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8EF62D7341CBFE3E" authorityName="Roding" authorityYear="1798" box="[182,241,367,392]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91EEA9CFF9E467C8EF62D7341CBFE3E" box="[182,241,367,392]" italics="true" pageId="19" pageNumber="490">Pupa</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8F772D734097FE3F" author="Kay, A. E." box="[311,429,367,393]" pageId="19" pageNumber="490" pagination="1 - 653" refId="ref24228" refString="Kay, A. E. (1979) Hawaiian marine shells. Reef and shore fauna of Hawaii: Section 4: Mollusca. Bishop Museum Special Publication, 64, 1 - 653." type="journal article" year="1979">Kay, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8FF82D734375FE3F" author="Yonow, N." box="[440,591,367,393]" pageId="19" pageNumber="490" refId="ref27463" refString="Yonow, N. (2008) Sea Slugs of the Red Sea. Pensoft, Sofia, 303 pp." type="book" year="2008">Yonow, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8C1B2D73427FFE3F" author="Gosliner, T. & Valdes, A. & Behrens, D. W." box="[603,837,367,393]" pageId="19" pageNumber="490" refId="ref23324" refString="Gosliner, T., Valdes, A. & Behrens, D. W. (2018) Nudibranch and Sea Slug Identification: Indo-Pacific. New World Publications Inc., Jacksonville, Florida, 451 pp." type="book" year="2018">
|
||
Gosliner
|
||
<emphasis id="B91EEA9CFF9E467C8C822D7343C0FE3F" box="[706,762,367,393]" italics="true" pageId="19" pageNumber="490">et al.</emphasis>
|
||
, 2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9E467C8E872D8F404DFCA3" blockId="19.[151,1437,151,1149]" pageId="19" pageNumber="490">
|
||
Souverbie
|
||
<emphasis id="B91EEA9CFF9E467C8F002D8F406CFE1A" box="[320,342,403,428]" italics="true" pageId="19" pageNumber="490">in</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8F1E2D8F43E8FE1B" author="Souverbie, S. - M. & Montrouzier, X." box="[350,722,403,429]" pageId="19" pageNumber="490" pagination="161 - 176" refId="ref26745" refString="Souverbie, S. - M. & Montrouzier, X. (1863) Descriptions d'especes nouvelles de l'archipel caledonien, et des iles Salomon et Woodlark. Journal de Conchyliologie, Series 3, 3, 161 - 176, pl. 5." type="journal article" year="1863">Souverbie & Montrouzier (1863)</bibRefCitation>
|
||
introduced the name
|
||
<taxonomicName id="4C6A4D0DFF9E467C8D872D8F446FFE1B" authority="Souverbie, 1863" authorityName="Souverbie" authorityYear="1863" box="[967,1365,403,429]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8D872D8F45AFFE1B" box="[967,1173,403,429]" italics="true" pageId="19" pageNumber="490">Tornatella alveola</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8ADD2D8F446FFE1B" author="Souverbie, S. - M. & Montrouzier, X." box="[1181,1365,403,429]" pageId="19" pageNumber="490" pagination="161 - 176" refId="ref26745" refString="Souverbie, S. - M. & Montrouzier, X. (1863) Descriptions d'especes nouvelles de l'archipel caledonien, et des iles Salomon et Woodlark. Journal de Conchyliologie, Series 3, 3, 161 - 176, pl. 5." type="journal article" year="1863">Souverbie, 1863</bibRefCitation>
|
||
</taxonomicName>
|
||
based on a single shell collected from Île Art,
|
||
<collectingCountry id="F37D761EFF9E467C8C132DAB423EFE67" box="[595,772,439,465]" name="New Caledonia" pageId="19" pageNumber="490">New Caledonia</collectingCountry>
|
||
. The
|
||
<typeStatus id="54D1882CFF9E467C8D002DAB4298FE67" box="[832,930,439,465]" pageId="19" pageNumber="490" type="holotype">holotype</typeStatus>
|
||
was described as having a white background with flesh pink, quadrangular spots arranged in transverse series; these spots alternate with smaller, white areas of background color.
|
||
<taxonomicName id="4C6A4D0DFF9E467C8FCF2DE34364FDAF" authorityName="Souverbie" authorityYear="1863" box="[399,606,511,537]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8FCF2DE34364FDAF" box="[399,606,511,537]" italics="true" pageId="19" pageNumber="490">Tornatella alveola</emphasis>
|
||
</taxonomicName>
|
||
is considered a member of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8DE32DE342E4FDAE" authorityName="Roding" authorityYear="1798" box="[931,990,511,536]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91EEA9CFF9E467C8DE32DE342E4FDAE" box="[931,990,511,536]" italics="true" pageId="19" pageNumber="490">Pupa</emphasis>
|
||
</taxonomicName>
|
||
and by some authors a synonym of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8BC82DE341E7FD8B" baseAuthorityName="A. Adams" baseAuthorityYear="1855" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis id="B91EEA9CFF9E467C8BC82DE341E7FD8B" italics="true" pageId="19" pageNumber="490">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8F5C2E3F4084FD8B" author="Chaban, E. M." box="[284,446,547,573]" pageId="19" pageNumber="490" pagination="415 - 448" refId="ref22248" refString="Chaban, E. M. (2016) Heterobranch mollusks of the orders Acteonoidea and Cephalaspidea (Gastropoda: Heterobranchia) of Vietnam: Annotated check-list with illustrations of some species. In: Adrianov, A. V. & Lutaenko, K. A. (Eds.), Biodiversity of the Western Part of the South China Sea. Dalnauka, Vladivostok, pp. 415 - 448." type="book chapter" year="2016">Chaban, 2016</bibRefCitation>
|
||
). Again, no specimens matching the description of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8A482E3F454DFD8B" authorityName="Souverbie" authorityYear="1863" box="[1032,1143,547,573]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8A482E3F454DFD8B" box="[1032,1143,547,573]" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
were obtained from
|
||
<collectingCountry id="F37D761EFF9E467C8B272E3F4031FDD7" name="New Caledonia" pageId="19" pageNumber="490">New Caledonia</collectingCountry>
|
||
, but the examination of a photograph of the
|
||
<typeStatus id="54D1882CFF9E467C8D4F2E5B424BFDD7" box="[783,881,583,609]" pageId="19" pageNumber="490" type="holotype">holotype</typeStatus>
|
||
(MHNBx 2004.TY.29,
|
||
<figureCitation id="13512A0BFF9E467C8AC22E5B45D0FDD7" box="[1154,1258,583,609]" captionStart="FIGURE 12" captionStartId="18.[152,255,1693,1717]" captionTargetBox="[151,1436,1121,1669]" captionTargetId="figure-345@18.[151,1436,1121,1669]" captionTargetPageId="18" captionText="FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP." figureDoi="http://doi.org/10.5281/zenodo.7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="19" pageNumber="490">Fig. 12D</figureCitation>
|
||
), which is very similar to the
|
||
<typeStatus id="54D1882CFF9E467C8F762E7740A2FD33" box="[310,408,619,645]" pageId="19" pageNumber="490" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8F802E77437DFD33" authorityName="Reeve" authorityYear="1842" box="[448,583,619,645]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8F802E77437DFD33" box="[448,583,619,645]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
, except the patches are larger and more pinkish in
|
||
<taxonomicName id="4C6A4D0DFF9E467C8ACD2E7745C6FD33" authorityName="Souverbie" authorityYear="1863" box="[1165,1276,619,645]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8ACD2E7745C6FD33" box="[1165,1276,619,645]" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
, and the shell appears to be slightly narrower. The
|
||
<typeStatus id="54D1882CFF9E467C8C702E9343A8FD1F" box="[560,658,655,681]" pageId="19" pageNumber="490" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8CF72E93421FFD1F" authorityName="Souverbie" authorityYear="1863" box="[695,805,655,681]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8CF72E93421FFD1F" box="[695,805,655,681]" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
appears to fall within the morphological variation range of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8EF62EAF4006FD7B" authorityName="Reeve" authorityYear="1842" box="[182,316,691,717]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8EF62EAF4006FD7B" box="[182,316,691,717]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
. However, because complete specimens/DNA of these two species were not available for study and due to the distance between their type localities, we prefer to leave the possible synonymy of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8A872ECB4474FD47" authorityName="Reeve" authorityYear="1842" box="[1223,1358,727,753]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8A872ECB4474FD47" box="[1223,1358,727,753]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A4D0DFF9E467C8BC82ECB41D3FCA3" authorityName="Souverbie" authorityYear="1863" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8BC82ECB41D3FCA3" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
as tentative.
|
||
</paragraph>
|
||
<paragraph id="8BD5368EFF9E467C8E872F034091FBCB" blockId="19.[151,1437,151,1149]" pageId="19" pageNumber="490">
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8E872F034051FC8F" author="Pilsbry, H. A." box="[199,363,799,825]" pageId="19" pageNumber="490" pagination="207 - 230" refId="ref25559" refString="Pilsbry, H. A. (1917) Marine Mollusks of Hawaii, I - III. Proceedings of the Academy of Natural Sciences of Philadelphia, 69, 207 - 230, pls. 14 - 15." type="journal article" year="1917">Pilsbry (1917)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C6A4D0DFF9E467C8FAA2F03437DFC8F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[490,583,799,825]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="solidula">
|
||
<emphasis id="B91EEA9CFF9E467C8FAA2F03437DFC8F" box="[490,583,799,825]" italics="true" pageId="19" pageNumber="490">Solidula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C6A4D0DFF9E467C8C112F034261FC8F" authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[593,859,799,825]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis id="B91EEA9CFF9E467C8C112F034381FC8F" box="[593,699,799,825]" italics="true" pageId="19" pageNumber="490">thaanumi</emphasis>
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8C852F034261FC8F" author="Pilsbry, H. A." box="[709,859,799,825]" pageId="19" pageNumber="490" pagination="207 - 230" refId="ref25559" refString="Pilsbry, H. A. (1917) Marine Mollusks of Hawaii, I - III. Proceedings of the Academy of Natural Sciences of Philadelphia, 69, 207 - 230, pls. 14 - 15." type="journal article" year="1917">Pilsbry, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
based on shells collected off Honolulu, Hawaiian Islands between
|
||
<quantity id="4C929B6BFF9E467C8F162F5E4080FCEB" box="[342,442,834,861]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="19" pageNumber="490" unit="m" value="13.0" valueMax="15.0" valueMin="11.0">11–15 m</quantity>
|
||
depth. The shells were described as “oblong with acutely conic spire, solid, somewhat shining, whitish, unevenly tessellated with flesh pink, the spotting interrupted by pale bands on the sixth and twelfth spaces between spiral grooves.” We also examined a photograph of one of the
|
||
<typeStatus id="54D1882CFF9E467C8A4F2F97454BFC13" box="[1039,1137,907,933]" pageId="19" pageNumber="490" type="syntype">syntypes</typeStatus>
|
||
(ANSP 117069,
|
||
<figureCitation id="13512A0BFF9E467C8B722F9744AEFC13" box="[1330,1428,907,933]" captionStart="FIGURE 12" captionStartId="18.[152,255,1693,1717]" captionTargetBox="[151,1436,1121,1669]" captionTargetId="figure-345@18.[151,1436,1121,1669]" captionTargetPageId="18" captionText="FIGURE 12. Photographs of shell specimens of Pupa tessellata (Reeve, 1842) and possible synonyms, ventral view. A–C. Three syntypes of Tornatella tessellata Reeve, 1842 (NHMUK 196954), ©NHMUK, photos by K. Webb. D. Holotype of Tornatella alveola Souverbie in Souverbie & Montrouzier, 1863 (MHNBx 2004.TY.29), ©MHNBx, photo by L. Charles. E. Syntype of Solidula thaanumi Pilsbry, 1917 (ANSP 117069), ©ANSP." figureDoi="http://doi.org/10.5281/zenodo.7860394" httpUri="https://zenodo.org/record/7860394/files/figure.png" pageId="19" pageNumber="490">Fig. 12E</figureCitation>
|
||
) which is very similar to the type material of both
|
||
<taxonomicName id="4C6A4D0DFF9E467C8CFD2FB34278FC7F" authorityName="Reeve" authorityYear="1842" box="[701,834,943,969]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8CFD2FB34278FC7F" box="[701,834,943,969]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A4D0DFF9E467C8D382FB342DFFC7F" authorityName="Souverbie" authorityYear="1863" box="[888,997,943,969]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8D382FB342DFFC7F" box="[888,997,943,969]" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
, although narrower and more elongate.
|
||
<emphasis id="B91EEA9CFF9E467C8ED72FCF405DFC5B" box="[151,359,979,1005]" italics="true" pageId="19" pageNumber="490">
|
||
<taxonomicName id="4C6A4D0DFF9E467C8ED72FCF41CEFC5B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,244,979,1005]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="solidula">Solidula</taxonomicName>
|
||
<taxonomicName id="4C6A4D0DFF9E467C8EBD2FCF405DFC5B" authorityName="Pilsbry" authorityYear="1917" box="[253,359,979,1005]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="thaanumi">thaanumi</taxonomicName>
|
||
</emphasis>
|
||
is currently considered a member of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8D552FCF426AFC5A" authorityName="Roding" authorityYear="1798" box="[789,848,979,1004]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91EEA9CFF9E467C8D552FCF426AFC5A" box="[789,848,979,1004]" italics="true" pageId="19" pageNumber="490">Pupa</emphasis>
|
||
</taxonomicName>
|
||
, but its identity remains unclear. For example,
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8B2E2FCF41D8FBA7" author="Kay, A. E." pageId="19" pageNumber="490" pagination="1 - 653" refId="ref24228" refString="Kay, A. E. (1979) Hawaiian marine shells. Reef and shore fauna of Hawaii: Section 4: Mollusca. Bishop Museum Special Publication, 64, 1 - 653." type="journal article" year="1979">Kay (1979)</bibRefCitation>
|
||
regarded
|
||
<taxonomicName id="4C6A4D0DFF9E467C8F182FEB40DBFBA7" authorityName="Pilsbry" authorityYear="1917" box="[344,481,1015,1041]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis id="B91EEA9CFF9E467C8F182FEB40DBFBA7" box="[344,481,1015,1041]" italics="true" pageId="19" pageNumber="490">P. thaanumi</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8CF52FEB4204FBA7" authorityName="Reeve" authorityYear="1842" box="[693,830,1015,1041]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8CF52FEB4204FBA7" box="[693,830,1015,1041]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<bibRefCitation id="EFFB4B7FFF9E467C8D3D2FEB42C5FBA7" author="Beu, A. G." box="[893,1023,1015,1041]" pageId="19" pageNumber="490" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004)</bibRefCitation>
|
||
cited
|
||
<taxonomicName id="4C6A4D0DFF9E467C8A092FEB45E8FBA7" authorityName="Pilsbry" authorityYear="1917" box="[1097,1234,1015,1041]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis id="B91EEA9CFF9E467C8A092FEB45E8FBA7" box="[1097,1234,1015,1041]" italics="true" pageId="19" pageNumber="490">P. thaanumi</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8ED7280741C1FB83" baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[151,251,1051,1077]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis id="B91EEA9CFF9E467C8ED7280741C1FB83" box="[151,251,1051,1077]" italics="true" pageId="19" pageNumber="490">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
. Because we had no access to complete specimens from the Hawaiian Islands, we are unable to verify its synonymy with
|
||
<taxonomicName id="4C6A4D0DFF9E467C8F34282340C1FBEF" authorityName="Reeve" authorityYear="1842" box="[372,507,1087,1113]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="tessellata">
|
||
<emphasis id="B91EEA9CFF9E467C8F34282340C1FBEF" box="[372,507,1087,1113]" italics="true" pageId="19" pageNumber="490">P. tessellata</emphasis>
|
||
</taxonomicName>
|
||
and/or
|
||
<taxonomicName id="4C6A4D0DFF9E467C8C15282343FFFBEF" authorityName="Souverbie" authorityYear="1863" box="[597,709,1087,1113]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="alveola">
|
||
<emphasis id="B91EEA9CFF9E467C8C15282343FFFBEF" box="[597,709,1087,1113]" italics="true" pageId="19" pageNumber="490">P. alveola</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="4C6A4D0DFF9E467C8D422823425CFBEF" baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[770,870,1087,1113]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis id="B91EEA9CFF9E467C8D422823425CFBEF" box="[770,870,1087,1113]" italics="true" pageId="19" pageNumber="490">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
appears to be a completely different species (see description of
|
||
<taxonomicName id="4C6A4D0DFF9E467C8F79287F40A0FBCB" baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[313,410,1123,1149]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="19" pageNumber="490" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis id="B91EEA9CFF9E467C8F79287F40A0FBCB" box="[313,410,1123,1149]" italics="true" pageId="19" pageNumber="490">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |