treatments-xml/data/03/C3/87/03C38782FFD6FF97C9AE157E4D96FF7F.xml
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<document id="FBD44A6F0172BEFA90BE99C9288EA1C0" ID-CLB-Dataset="46543" ID-DOI="10.5281/zenodo.278052" ID-GBIF-Dataset="89803f24-d147-45a0-b77b-995f45bc0425" ID-ISSN="1175-5326" ID-Zenodo-Dep="278052" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460377557173" checkinUser="plazi" docAuthor="Larsen, Torben B. &amp; Congdon, Colin E." docDate="2011" docId="03C38782FFD6FF97C9AE157E4D96FF7F" docLanguage="en" docName="zt02931p058.pdf" docOrigin="Zootaxa 2931" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triskelionia tricerata Larsen &amp; Congdon, 2011, comb. nov." docType="treatment" docVersion="8" lastPageNumber="54" masterDocId="FFFAFFFAFFD7FF95C93916604C7CFFAB" masterDocTitle="Triskelionia, a new African genus of the Celaenorrhinini (Lepidoptera: Hesperiidae) and the promotion of T. compacta to species-status." masterLastPageNumber="58" masterPageNumber="53" pageNumber="54" updateTime="1698296419481" updateUser="ExternalLinkService">
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<mods:title id="B30B05A86D71C39837FD9B9400FC9346">Triskelionia, a new African genus of the Celaenorrhinini (Lepidoptera: Hesperiidae) and the promotion of T. compacta to species-status.</mods:title>
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<mods:namePart id="977C773D1B1531F2F0DC5EDAD0C6B7F7">Larsen, Torben B.</mods:namePart>
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<mods:namePart id="1238695111EFC9690B2BE08C3ACFA245">Congdon, Colin E.</mods:namePart>
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<treatment id="03C38782FFD6FF97C9AE157E4D96FF7F" ID-DOI="http://doi.org/10.5281/zenodo.6188372" ID-GBIF-Taxon="119413335" ID-Zenodo-Dep="6188372" LSID="urn:lsid:plazi:treatment:03C38782FFD6FF97C9AE157E4D96FF7F" httpUri="http://treatment.plazi.org/id/03C38782FFD6FF97C9AE157E4D96FF7F" lastPageId="2" lastPageNumber="54" pageId="1" pageNumber="54">
<subSubSection id="C370651FFFD6FF94C9AE157E48F1F8AA" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8BD53694FFD6FF94C9AE157E4E5FFC93" blockId="1.[151,547,798,824]" box="[151,547,798,824]" pageId="1" pageNumber="54">
<heading id="D09D81F8FFD6FF94C9AE157E4E5FFC93" bold="true" box="[151,547,798,824]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="B91EEA86FFD6FF94C9AE157E4E5FFC93" bold="true" box="[151,547,798,824]" pageId="1" pageNumber="54">
<taxonomicName id="4C6A4D17FFD6FF94C9AE157E4DEAFC93" box="[151,406,798,824]" class="Insecta" family="Hesperiidae" genus="Triskelionia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="tricerata" status="comb. nov.">
<emphasis id="B91EEA86FFD6FF94C9AE157E4DEAFC93" bold="true" box="[151,406,798,824]" italics="true" pageId="1" pageNumber="54">Triskelionia tricerata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57FDFFD6FF94C8A4157E4E5FFC93" box="[413,547,798,824]" pageId="1" pageNumber="54">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE15054826FC0A" blockId="1.[151,1435,869,930]" pageId="1" pageNumber="54">
The genus is defined by
<taxonomicName id="4C6A4D17FFD6FF94C88815064E4CFCD6" box="[433,560,870,893]" class="Insecta" family="Hesperiidae" genus="Triskelionia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="tricerata">
<emphasis id="B91EEA86FFD6FF94C88815064E4CFCD6" box="[433,560,870,893]" italics="true" pageId="1" pageNumber="54">T. tricerata</emphasis>
</taxonomicName>
, the
<typeStatus id="54D18836FFD6FF94CB5315064EE6FCD5" box="[618,666,870,894]" pageId="1" pageNumber="54">type</typeStatus>
species, and is characterized by a number of features that unite the two species, while differentiating them from other genera in the
<taxonomicName id="4C6A4D17FFD6FF94CA5E15E9486BFC0A" box="[871,1047,905,929]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini</taxonomicName>
tribe:
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE15B14E09FBA6" blockId="1.[151,1436,977,1613]" pageId="1" pageNumber="54">
a) as already mentioned by
<bibRefCitation id="EFFB4B65FFD6FF94C8CF15B24EEEFC42" author="Evans" box="[502,658,977,1001]" pageId="1" pageNumber="54" refString="Evans, W. H. (1937) A catalogue of the African Hesperiidae indicating the classification and nomenclature adopted in the British Museum. British Museum (Natural History), London, UK, 212 pp." type="book" year="1937">Evans (1937)</bibRefCitation>
the palps are longer and the antennal club more arcuate than in
<taxonomicName id="4C6A4D17FFD6FF94C9F015964D43FBA6" box="[201,319,1014,1037]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94C9F015964D43FBA6" box="[201,319,1014,1037]" italics="true" pageId="1" pageNumber="54">Sarangesa</emphasis>
</taxonomicName>
, “approaching
<taxonomicName id="4C6A4D17FFD6FF94C8D215954E1DFBA6" box="[491,609,1013,1037]" class="Insecta" family="Hesperiidae" genus="Calleagris" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94C8D215954E1DFBA6" box="[491,609,1013,1037]" italics="true" pageId="1" pageNumber="54">Calleagris</emphasis>
</taxonomicName>
”.
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE12794ED8FBFE" blockId="1.[151,1436,977,1613]" pageId="1" pageNumber="54">
b) the three-pronged spot in the forewing cell is different from all other
<taxonomicName id="4C6A4D17FFD6FF94CAD6127948E3FB9A" box="[1007,1183,1049,1073]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini</taxonomicName>
(slightly similar in
<taxonomicName id="4C6A4D17FFD6FF94CCBE127A4D69FBFE" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="seineri">
<emphasis id="B91EEA86FFD6FF94CCBE127A4D69FBFE" italics="true" pageId="1" pageNumber="54">S. seineri</emphasis>
</taxonomicName>
which has very different genitalia).
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE12014810FBD2" blockId="1.[151,1436,977,1613]" box="[151,1132,1121,1145]" pageId="1" pageNumber="54">
c) the wing shape, especially of
<taxonomicName id="4C6A4D17FFD6FF94CB2B12024E08FBD2" box="[530,628,1122,1145]" class="Insecta" family="Hesperiidae" genus="Triskelionia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="tricerata">
<emphasis id="B91EEA86FFD6FF94CB2B12024E08FBD2" box="[530,628,1122,1145]" italics="true" pageId="1" pageNumber="54">tricerata</emphasis>
</taxonomicName>
, is closer to that of
<taxonomicName id="4C6A4D17FFD6FF94CA6B12024FE8FBD2" box="[850,916,1122,1145]" class="Insecta" family="Hesperiidae" genus="Eretis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CA6B12024FE8FBD2" box="[850,916,1122,1145]" italics="true" pageId="1" pageNumber="54">Eretis</emphasis>
</taxonomicName>
than to
<emphasis id="B91EEA86FFD6FF94CAD712024810FBD2" box="[1006,1132,1122,1145]" italics="true" pageId="1" pageNumber="54">
<taxonomicName id="4C6A4D17FFD6FF94CAD712024814FBD2" box="[1006,1128,1122,1145]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">Sarangesa</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE12E54DC0FADE" blockId="1.[151,1436,977,1613]" pageId="1" pageNumber="54">
d) the male genitalia are very different from any
<taxonomicName id="4C6A4D17FFD6FF94CBF012E64F43FB36" box="[713,831,1158,1181]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CBF012E64F43FB36" box="[713,831,1158,1181]" italics="true" pageId="1" pageNumber="54">Sarangesa</emphasis>
</taxonomicName>
and show affinity with some sections of
<taxonomicName id="4C6A4D17FFD6FF94CC3312E54C8CFB6A" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CC3312E54C8CFB6A" italics="true" pageId="1" pageNumber="54">Celaenorrhinus</emphasis>
</taxonomicName>
. The uncus is small and pointed, without lateral side branches. The very large gnathos consists of two fully fused branches, forming an almost box-like structure, the external surfaces of which are finely ribbed and spined. It is proportionately larger than in any
<taxonomicName id="4C6A4D17FFD6FF94CBE312914FF5FAA2" box="[730,905,1265,1289]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CBE312914FF5FAA2" box="[730,905,1265,1289]" italics="true" pageId="1" pageNumber="54">Celaenorrhinus</emphasis>
</taxonomicName>
; no
<taxonomicName id="4C6A4D17FFD6FF94CA8712924848FAA2" box="[958,1076,1266,1289]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CA8712924848FAA2" box="[958,1076,1266,1289]" italics="true" pageId="1" pageNumber="54">Sarangesa</emphasis>
</taxonomicName>
or
<taxonomicName id="4C6A4D17FFD6FF94CD65129248E2FAA2" box="[1116,1182,1266,1289]" class="Insecta" family="Hesperiidae" genus="Eretis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CD65129248E2FAA2" box="[1116,1182,1266,1289]" italics="true" pageId="1" pageNumber="54">Eretis</emphasis>
</taxonomicName>
has this
<typeStatus id="54D18836FFD6FF94CC3D12924948FAA1" box="[1284,1332,1266,1290]" pageId="1" pageNumber="54">type</typeStatus>
of structure; at most the gnathos is composed of narrow branches that do not fuse. The vinculum continues narrowly to fuse basally rather than forming a regular, solid saccus. The shape and proportion of the penis is very different from
<taxonomicName id="4C6A4D17FFD6FF94C83F133D4DC9FADE" box="[262,437,1373,1397]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94C83F133D4DC9FADE" box="[262,437,1373,1397]" italics="true" pageId="1" pageNumber="54">Celaenorrhinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE13E14FC9FA16" blockId="1.[151,1436,977,1613]" pageId="1" pageNumber="54">
e) the pupa has no free proboscis-sheath as is the case in all known
<taxonomicName id="4C6A4D17FFD6FF94CAA613E24869FA32" box="[927,1045,1410,1433]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CAA613E24869FA32" box="[927,1045,1410,1433]" italics="true" pageId="1" pageNumber="54">Sarangesa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A4D17FFD6FF94CD7513E248F1FA32" box="[1100,1165,1410,1433]" class="Insecta" family="Hesperiidae" genus="Eretis" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94CD7513E248F1FA32" box="[1100,1165,1410,1433]" italics="true" pageId="1" pageNumber="54">Eretis</emphasis>
</taxonomicName>
, and which is very long in
<taxonomicName id="4C6A4D17FFD6FF94C9DD13C54DEFFA16" box="[228,403,1445,1469]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94C9DD13C54DEFFA16" box="[228,403,1445,1469]" italics="true" pageId="1" pageNumber="54">Celaenorrhinus</emphasis>
</taxonomicName>
, extending well beyond the end of the abdomen.
</paragraph>
<paragraph id="8BD53694FFD6FF94C9AE13A94C89F9E6" blockId="1.[151,1436,977,1613]" pageId="1" pageNumber="54">
f) the larval host-plant is a member of the
<taxonomicName id="4C6A4D17FFD6FF94CBB813A94E90FA4A" box="[641,748,1481,1505]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="54" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
, not known as host-plant for any other African member of the
<taxonomicName id="4C6A4D17FFD6FF94C9FE138D4D06F9AE" box="[199,378,1517,1541]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini</taxonomicName>
, although used by
<taxonomicName id="4C6A4D17FFD6FF94CB70138D4F50F9AE" box="[585,812,1517,1541]" class="Insecta" family="Hesperiidae" genus="Calleagris" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="jamesoni">
<emphasis id="B91EEA86FFD6FF94CB70138D4F50F9AE" box="[585,812,1517,1541]" italics="true" pageId="1" pageNumber="54">Calleagris jamesoni</emphasis>
</taxonomicName>
in the tribe
<taxonomicName id="4C6A4D17FFD6FF94CA89138D4866F9AE" box="[944,1050,1517,1541]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="tribe" tribe="Tagiadini">Tagiadini</taxonomicName>
and many species in the Odontoptilina (which should receive full status as a tribe and is very distant from
<taxonomicName id="4C6A4D17FFD6FF94CD38107148FBF982" box="[1025,1159,1553,1577]" class="Insecta" family="Hesperiidae" genus="Triskelionia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91EEA86FFD6FF94CD38107148FBF982" box="[1025,1159,1553,1577]" italics="true" pageId="1" pageNumber="54">Triskelionia</emphasis>
</taxonomicName>
and other
<taxonomicName id="4C6A4D17FFD6FF94CC3910714C95F9E6" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini</taxonomicName>
).
</paragraph>
<paragraph id="8BD53694FFD6FF94C9FE101E48F1F8AA" blockId="1.[151,1436,1661,2010]" pageId="1" pageNumber="54">
We attach special importance to the following characters: the longer palps; the large fully-fused gnathos structure; the lack of a free proboscis sheath in the pupa; and the use of
<taxonomicName id="4C6A4D17FFD6FF94CA4710C14F9AF912" box="[894,998,1697,1721]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="54" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
as host-plant. None of these characters is found in any of the
<taxonomicName id="4C6A4D17FFD6FF94C8A110A64E72F976" box="[408,526,1734,1757]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEA86FFD6FF94C8A110A64E72F976" box="[408,526,1734,1757]" italics="true" pageId="1" pageNumber="54">Sarangesa</emphasis>
</taxonomicName>
; each is certainly independent of any of the other characters. The simultaneous presence of four such characters is not compatible with retaining the species in
<emphasis id="B91EEA86FFD6FF94CD36108A48F1F8AA" box="[1039,1165,1770,1793]" italics="true" pageId="1" pageNumber="54">
<taxonomicName id="4C6A4D17FFD6FF94CD36108A48F5F8AA" box="[1039,1161,1770,1793]" class="Insecta" family="Hesperiidae" genus="Sarangesa" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">Sarangesa</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C370651FFFD6FF97C9FE116D4D96FF7F" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="etymology">
<paragraph id="8BD53694FFD6FF97C9FE116D4D96FF7F" blockId="1.[151,1436,1661,2010]" lastBlockId="2.[151,1435,151,212]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
<emphasis id="B91EEA86FFD6FF94C9FE116D4D32F88E" bold="true" box="[199,334,1805,1829]" pageId="1" pageNumber="54">Etymology.</emphasis>
The genus name
<taxonomicName id="4C6A4D17FFD6FF94CB19116D4EDAF88E" box="[544,678,1805,1829]" class="Insecta" family="Hesperiidae" genus="Triskelionia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91EEA86FFD6FF94CB19116D4EDAF88E" box="[544,678,1805,1829]" italics="true" pageId="1" pageNumber="54">Triskelionia</emphasis>
</taxonomicName>
refers to the unique, conjoined three-pronged shape (from Greek ρισκέλιον) of the fused spots in the forewing cell of the two species included in the genus (triskelion is colloquially called the “three-legged cross” in
<collectingCountry id="F37D7604FFD6FF94CB6311354ED1F8C6" box="[602,685,1877,1901]" name="United Kingdom" pageId="1" pageNumber="54">Britain</collectingCountry>
). This ancient symbol was common on Mycenaean pottery from where it entered modern heraldry. It is used in the flags or seals of several modern territories or institutions, including the
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and Ingushetia. It is also the symbol of the Global Fund to fight AIDS, Tuberculosis and Malaria, and is in use by the
<collectingCountry id="F37D7604FFD6FF94C8DD11A14E74F872" box="[484,520,1985,2009]" name="United States of America" pageId="1" pageNumber="54">US</collectingCountry>
Department of Transportation.
<emphasis id="B91EEA86FFD6FF94CA4911A14F9CF872" box="[880,992,1985,2009]" italics="true" pageId="1" pageNumber="54">Triskelion</emphasis>
was preoccupied by a genus of extinct Eocene flagellate Protozoa in the order Ebriida; however, the name seemed so descriptive that a modification of triskelion seemed appropriate.
</paragraph>
</subSubSection>
</treatment>
</document>