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<document id="C0424AAB44939D31E1FE967439E2E6A6" ID-DOI="10.11646/phytotaxa.323.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13696684" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725566521507" checkinUser="felipe" docAuthor="Henderson, Andrew &amp; Dung, Nguyen Quoc" docDate="2017" docId="03C38781FF81FFFAFF776BBBFF15F788" docLanguage="en" docName="phytotaxa.323.2.4.pdf" docOrigin="Phytotaxa 323 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.323.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Licuala nuichuaensis Henderson &amp; Nguyen Quoc Dung 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="168" masterDocId="FFFAFFF9FF88FFF3FFFF6F1DFFD3FFA9" masterDocTitle="New species of Lanonia, Licuala, and Pinanga (Arecaceae) from Vietnam" masterLastPageNumber="172" masterPageNumber="159" pageNumber="168" updateTime="1725567521063" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="52CD0829B58A1C5348D347E1B683C46A">New species of Lanonia, Licuala, and Pinanga (Arecaceae) from Vietnam</mods:title>
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<mods:name id="61321D959AD81A6CC52AE3994117AFEC" type="personal">
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<mods:namePart id="EE3BDC5CF0D258F6E731D3CFF7306DC5">Henderson, Andrew</mods:namePart>
<mods:affiliation id="D4BB42586EE34DC9BCCAF92C73CA5FFD">Institute of Systematic Botany, The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
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<mods:namePart id="50D62F036AAE0D3FBC888E0C6390F45C">Dung, Nguyen Quoc</mods:namePart>
<mods:affiliation id="3F2C108FC5C824059BF27CA4720F381D">Forest Inventory and Planning Institute, Thanh Tri, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="0C6AE8D99FA96BDB53705BAEFA192579">Phytotaxa</mods:title>
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<mods:date id="1A7E2F6FDC503696F1840D4AE61FE036">2017</mods:date>
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<mods:number id="BA165F40B9A1B75332067C525FDE7CBB">2017-09-29</mods:number>
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<mods:identifier id="50AC7A930D76766082C1A57E533006D2" type="DOI">10.11646/phytotaxa.323.2.4</mods:identifier>
<mods:identifier id="5F36393624C7DC30CC67A30FEC8A5983" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="9238DEFB52E5CA739DEFD90D3455F4A8" type="Zenodo-Dep">13696684</mods:identifier>
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<treatment id="03C38781FF81FFFAFF776BBBFF15F788" ID-DOI="http://doi.org/10.5281/zenodo.13696728" ID-Zenodo-Dep="13696728" LSID="urn:lsid:plazi:treatment:03C38781FF81FFFAFF776BBBFF15F788" httpUri="http://treatment.plazi.org/id/03C38781FF81FFFAFF776BBBFF15F788" lastPageNumber="168" pageId="9" pageNumber="168">
<subSubSection id="C370651CFF81FFFAFF776BBBFC6DFB68" box="[136,958,1190,1217]" pageId="9" pageNumber="168" type="nomenclature">
<paragraph id="8BD53697FF81FFFAFF776BBBFC6DFB68" blockId="9.[136,958,1190,1217]" box="[136,958,1190,1217]" pageId="9" pageNumber="168">
<heading id="D09D81FBFF81FFFAFF776BBBFC6DFB68" box="[136,958,1190,1217]" fontSize="11" level="2" pageId="9" pageNumber="168" reason="3">
<taxonomicName id="4C6A4D14FF81FFFAFF776BBBFCD4FB68" ID-CoL="8WX52" authority="Henderson &amp; Nguyen Quoc Dung" authorityName="Henderson &amp; Nguyen Quoc Dung" authorityYear="2017" box="[136,775,1190,1217]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="nuichuaensis" status="sp. nov.">
<emphasis id="B91EEA85FF81FFFAFF776BBBFE51FB68" bold="true" box="[136,386,1190,1217]" italics="true" pageId="9" pageNumber="168">Licuala nuichuaensis</emphasis>
Henderson &amp; Nguyen Quoc Dung
</taxonomicName>
<emphasis id="B91EEA85FF81FFFAFCF16BB5FCB0FB68" box="[782,867,1192,1217]" italics="true" pageId="9" pageNumber="168">
<taxonomicNameLabel id="A22D57FEFF81FFFAFCF16BB5FCB0FB68" box="[782,867,1192,1217]" pageId="9" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13512A12FF81FFFAFC8D6BB5FC66FB68" box="[882,949,1192,1217]" captionStart="FIGURE 9" captionStartId="10.[136,229,1087,1108]" captionTargetBox="[154,1435,190,1061]" captionTargetId="figure-20@10.[151,1435,190,1061]" captionTargetPageId="10" captionText="FIGURE 9. Licuala nuichuaensis. Left, leaf. Right, leaf base with elongate ligules." figureDoi="http://doi.org/10.5281/zenodo.13696720" httpUri="https://zenodo.org/record/13696720/files/figure.png" pageId="9" pageNumber="168">Fig. 9</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C370651CFF81FFFAFF776BD2FEFDF8A8" pageId="9" pageNumber="168" type="materials_examined">
<paragraph id="8BD53697FF81FFFAFF776BD2FAE7FAA1" blockId="9.[136,1452,1231,1288]" pageId="9" pageNumber="168">
Type:—
<collectingCountry id="F37D7607FF81FFFAFF296BD2FE9BFB4D" box="[214,328,1231,1252]" name="Vietnam" pageId="9" pageNumber="168">VIETNAM</collectingCountry>
.
<collectingRegion id="49AEF875FF81FFFAFEAD6BD2FDF5FB4D" box="[338,550,1231,1252]" country="Vietnam" name="Ninh Thuan" pageId="9" pageNumber="168">Ninh Thuan Province</collectingRegion>
: Ninh Hai District, Nui Chua National Park, near summit of mountain,
<geoCoordinate id="EE5E5050FF81FFFAFB206BD2FAFBFB4D" box="[1247,1320,1231,1252]" degrees="11.80" direction="north" orientation="latitude" pageId="9" pageNumber="168" precision="555" value="11.8">11.80N</geoCoordinate>
,
<geoCoordinate id="EE5E5050FF81FFFAFACD6BD2FA56FB4D" box="[1330,1413,1231,1252]" degrees="109.16" direction="east" orientation="longitude" pageId="9" pageNumber="168" precision="555" value="109.16">109.16E</geoCoordinate>
, ca.
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,
<date id="FFD41057FF81FFFAFEF96BEEFEACFAA1" box="[262,383,1267,1288]" pageId="9" pageNumber="168" value="2009-10-26">26 Oct 2009</date>
,
<emphasis id="B91EEA85FF81FFFAFE766BEEFBC2FAA1" box="[393,1041,1267,1288]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED7BAE52FF81FFFAFE766BEEFE48FAA1" box="[393,411,1267,1288]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="9" pageNumber="168" type="Herbarium">A</collectionCode>
. Henderson, Bui Van Than, Nguyen Quoc Dat, &amp;
<collectionCode id="ED7BAE52FF81FFFAFC866BEEFC54FAA1" box="[889,903,1267,1288]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="9" pageNumber="168" type="Herbarium">A</collectionCode>
Ruc Cui 3642
</emphasis>
(
<typeStatus id="54D18835FF81FFFAFBE16BEEFBA7FAA1" box="[1054,1140,1267,1288]" pageId="9" pageNumber="168" type="holotype">holotype</typeStatus>
<collectionCode id="ED7BAE52FF81FFFAFB866BEEFB71FAA1" box="[1145,1186,1267,1288]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="9" pageNumber="168" type="Herbarium">HN</collectionCode>
!,
<typeStatus id="54D18835FF81FFFAFB4F6BEEFB26FAA1" box="[1200,1269,1267,1288]" pageId="9" pageNumber="168" type="isotype">isotype</typeStatus>
<collectionCode id="ED7BAE52FF81FFFAFB046BEEFAF6FAA1" box="[1275,1317,1267,1288]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="9" pageNumber="168" type="Herbarium">NY</collectionCode>
!).
</paragraph>
<paragraph id="8BD53697FF81FFFAFF776A25FADDFADC" blockId="9.[136,1452,1336,2081]" pageId="9" pageNumber="168">
It is similar to
<taxonomicName id="4C6A4D14FF81FFFAFED06A25FDCDFAF8" box="[303,542,1336,1361]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="bachmaensis">
<emphasis id="B91EEA85FF81FFFAFED06A25FDCDFAF8" box="[303,542,1336,1361]" italics="true" pageId="9" pageNumber="168">Licuala bachmaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A4D14FF81FFFAFDA96A25FCDBFAF8" box="[598,776,1336,1361]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="dakrongensis">
<emphasis id="B91EEA85FF81FFFAFDA96A25FCDBFAF8" box="[598,776,1336,1361]" italics="true" pageId="9" pageNumber="168">L. dakrongensis</emphasis>
</taxonomicName>
in its central segments wider at the apices and there briefly joined, but differs in its segments only briefly indented at the apices and closely spaced flower clusters.
</paragraph>
<paragraph id="8BD53697FF81FFFAFF426A9DFEFDF8A8" blockId="9.[136,1452,1336,2081]" pageId="9" pageNumber="168">
Stems solitary, 1.0 m tall, 5.0 cm diameter. Leaf number not recorded; sheath length not recorded; ligules 70.0 cm long, persistent and hanging down below the leaves; petioles length not recorded, with stout, black thorns to
<quantity id="4C929B72FF81FFFAFAA16AB9FA78FA14" box="[1374,1451,1444,1469]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="168" unit="cm" value="0.6">0.6 cm</quantity>
long; blades 140.0 cm wide, split into 22 segments, some briefly joined at their apices, with straight sides, wider at the apex; costa continuous into central segment, not forming a petiolule; middle segment
<quantity id="4C929B72FF81FFFAFBAC6AF1FB7DF9AC" box="[1107,1198,1516,1541]" metricMagnitude="-1" metricUnit="m" metricValue="6.55" pageId="9" pageNumber="168" unit="cm" value="65.5">65.5 cm</quantity>
long,
<quantity id="4C929B72FF81FFFAFB096AF1FA82F9AC" box="[1270,1361,1516,1541]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="9" pageNumber="168" unit="cm" value="10.3">10.3 cm</quantity>
wide at apex; lateral segment
<quantity id="4C929B72FF81FFFAFE7F690DFE08F980" box="[384,475,1552,1577]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" pageId="9" pageNumber="168" unit="cm" value="57.5">57.5 cm</quantity>
long,
<quantity id="4C929B72FF81FFFAFDDB690DFDA2F980" box="[548,625,1552,1577]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="168" unit="cm" value="3.3">3.3 cm</quantity>
wide at apex; segments briefly indented at the apex, with minute, brown scales abaxially. Inflorescences erect amongst the leaves,
<quantity id="4C929B72FF81FFFAFCE46929FC8EF9E4" box="[795,861,1588,1613]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="168" unit="m" value="1.5">1.5 m</quantity>
long, branched to 2 orders; prophylls not recorded; rachis bracts to
<quantity id="4C929B72FF81FFFAFEBD6945FE4CF9D8" box="[322,415,1624,1649]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" pageId="9" pageNumber="168" unit="cm" value="17.5">17.5 cm</quantity>
long, tubular, fraying at the apices; partial inflorescences 2; rachillae
<quantity id="4C929B72FF81FFFAFB316945FABAF9D8" box="[1230,1385,1624,1649]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="9" pageNumber="168" unit="cm" value="12.5" valueMax="13.5" valueMin="11.5">11.513.5 cm</quantity>
long, densely covered with felty, brown hairs, these tending to be arranged in short rows; flowers
<quantity id="4C929B72FF81FFFAFB7F6961FB06F93C" box="[1152,1237,1660,1685]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="168" unit="mm" value="3.5">3.5 mm</quantity>
long, 23 borne on a raised pedicel, these closely and irregularly arranged; calyx
<quantity id="4C929B72FF81FFFAFCC369BDFC41F910" box="[828,914,1696,1721]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="168" unit="mm" value="3.5">3.5 mm</quantity>
long, hairy as the rachillae, 3-lobed at the apex, not tubular basally; corolla
<quantity id="4C929B72FF81FFFAFE3C69D9FDC8F974" box="[451,539,1732,1757]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="168" unit="mm" value="2.5">2.5 mm</quantity>
long, split into 3, valvate petals, with dense, appressed hairs; stamens not seen; fruits not seen.
</paragraph>
</subSubSection>
<subSubSection id="C370651CFF81FFFAFF426811FF15F788" pageId="9" pageNumber="168" type="distribution">
<paragraph id="8BD53697FF81FFFAFF426811FE44F8E0" blockId="9.[136,1452,1336,2081]" pageId="9" pageNumber="168">
<emphasis id="B91EEA85FF81FFFAFF426811FE0CF88C" bold="true" box="[189,479,1804,1829]" pageId="9" pageNumber="168">Distribution and habitat</emphasis>
:—
<collectingRegion id="49AEF875FF81FFFAFDFF6811FD21F88C" box="[512,754,1804,1829]" country="Vietnam" name="Ninh Thuan" pageId="9" pageNumber="168">Ninh Thuan Province</collectingRegion>
in Nui Chua National Park in low forest on steep, rocky slopes at
<quantity id="4C929B72FF81FFFAFF5C682DFEF1F8E0" box="[163,290,1840,1865]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="9" pageNumber="168" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600800 m</quantity>
elevation.
</paragraph>
<paragraph id="8BD53697FF81FFFAFF426849FE5AF81C" blockId="9.[136,1452,1336,2081]" pageId="9" pageNumber="168">
<emphasis id="B91EEA85FF81FFFAFF426849FE7CF8C4" bold="true" box="[189,431,1876,1901]" pageId="9" pageNumber="168">Taxonomic notes:—</emphasis>
<taxonomicName id="4C6A4D14FF81FFFAFE506849FD71F8C4" authorityName="Henderson &amp; Nguyen Quoc Dung" authorityYear="2017" box="[431,674,1876,1901]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="nuichuaensis">
<emphasis id="B91EEA85FF81FFFAFE506849FD71F8C4" box="[431,674,1876,1901]" italics="true" pageId="9" pageNumber="168">Licuala nuichuaensis</emphasis>
</taxonomicName>
shares with
<taxonomicName id="4C6A4D14FF81FFFAFCC46849FC3EF8C4" box="[827,1005,1876,1901]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="bachmaensis">
<emphasis id="B91EEA85FF81FFFAFCC46849FC3EF8C4" box="[827,1005,1876,1901]" italics="true" pageId="9" pageNumber="168">L. bachmaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A4D14FF81FFFAFBD46849FB32F8C4" box="[1067,1249,1876,1901]" class="Liliopsida" family="Arecaceae" genus="Licuala" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="dakrongensis">
<emphasis id="B91EEA85FF81FFFAFBD46849FB32F8C4" box="[1067,1249,1876,1901]" italics="true" pageId="9" pageNumber="168">L. dakrongensis</emphasis>
</taxonomicName>
central segments briefly joined at the apices. It differs from them in its elongate ligules, shallowly lobed segment apices, and closely spaced flower clusters.
</paragraph>
<paragraph id="8BD53697FF81FFFAFF4268DDFF15F788" blockId="9.[136,1452,1336,2081]" pageId="9" pageNumber="168">
<emphasis id="B91EEA85FF81FFFAFF4268DDFDFEF870" bold="true" box="[189,557,1984,2009]" pageId="9" pageNumber="168">Additional specimen examined</emphasis>
.
<collectingCountry id="F37D7607FF81FFFAFDC368DDFD12F870" box="[572,705,1984,2009]" name="Vietnam" pageId="9" pageNumber="168">VIETNAM</collectingCountry>
.
<collectingRegion id="49AEF875FF81FFFAFD3468DDFC85F870" box="[715,854,1984,2009]" country="Vietnam" name="Ninh Thuan" pageId="9" pageNumber="168">Ninh Thuan</collectingRegion>
: Ninh Hai Distr., Vinh Hai Municipality, NEE slopes of Nui Chua mountains to SW of point 11˚4352”N 109˚0835”E, about
<quantity id="4C929B72FF81FFFAFC5268F9FBFEF854" box="[941,1069,2020,2045]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="168" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600700 m</quantity>
,
<emphasis id="B91EEA85FF81FFFAFBC768F9FABFF854" box="[1080,1388,2020,2045]" italics="true" pageId="9" pageNumber="168">J. Regalado et al. HLF4211</emphasis>
(HN, MO).
</paragraph>
</subSubSection>
</treatment>
</document>