223 lines
24 KiB
XML
223 lines
24 KiB
XML
<document id="6BEEDDD4016C5D68829D3D013CD5B9AF" ID-DOI="10.11646/zootaxa.5040.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531508" ID-ZooBank="60622671-2759-46AD-8AB0-04E9A71C2338" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749247351" checkinUser="plazi" docAuthor="Whorrall, Kyle & Philips, Keith" docDate="2021" docId="03C38780FFE5FFA9FF02FC141E7FF867" docLanguage="en" docName="zootaxa.5040.3.6.pdf" docOrigin="Zootaxa 5040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cordielytrum inca Whorrall & Philips 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="418" masterDocId="FFFAFFF8FFE1FFADFF95FF9E1B1CFFBA" masterDocTitle="Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips" masterLastPageNumber="427" masterPageNumber="414" pageNumber="418" updateTime="1699415554805" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="96B44111FE183202596DEA47CC8988CC">Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips</mods:title>
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<mods:name id="FE9B833754F187680B3D580B4796C55D" type="personal">
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<mods:roleTerm id="83D55EC41B11F2D3F6D159DAF923F200">Author</mods:roleTerm>
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<mods:namePart id="4F4B591F45899520C0DBDEF497275B0E">Whorrall, Kyle</mods:namePart>
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<mods:nameIdentifier id="DFA14AEF7160906A75B82FBBDA75F020" type="ORCID">0000-0002-1555-3076</mods:nameIdentifier>
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<mods:affiliation id="48E368B232304EFAD1DC08E8540F88C4">Manaaki Whenua-Landcare Research, 231 Morrin Road, St Johns, Auckland 1072, New Zealand. whorrallk @ landcareresearch. co. nz; https: // orcid. org / 0000 - 0002 - 1555 - 3076</mods:affiliation>
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<mods:nameIdentifier id="E4E537D972ABAABE4E8C92D800CA4EF6" type="email">whorrallk@landcareresearch.co.nz</mods:nameIdentifier>
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<mods:roleTerm id="B91C8FF91787F4F051BE984CFD5EF9E7">Author</mods:roleTerm>
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<mods:namePart id="7BFE6C88C8EB0ABB468F9F1E876B9762">Philips, Keith</mods:namePart>
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<mods:affiliation id="CD6408221CBD4AE1BCFB188603788BD9">Systematics and Evolution Laboratory, Department of Biology, Western Kentucky University, 1906 College Heights Blvd., Bowling Green, KY 42101 - 3576 USA.</mods:affiliation>
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<mods:title id="84E23C1B31D1A6C1FDB1A97E0F1412DE">Zootaxa</mods:title>
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<mods:date id="208D0B74B01324A13E9A09E79B40A356">2021</mods:date>
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<mods:number id="C7ED7CDCA2EE132E77722BD7A8D314AE">2021-09-22</mods:number>
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<taxonomicName id="4C6A4D15FFE5FFA9FF02FC141A68FC1F" authority="Whorrall & Philips, 2021" authorityName="Whorrall & Philips" authorityYear="2021" box="[151,372,906,933]" class="Insecta" family="Ptinidae" genus="Cordielytrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="inca" status="sp. nov.">
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<emphasis id="B91EEA84FFE5FFA9FF02FC141A68FC1F" bold="true" box="[151,372,906,933]" italics="true" pageId="4" pageNumber="418">Cordielytrum inca</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22D57FFFFE5FFA9FE16FC151AD9FC1F" box="[387,453,907,933]" pageId="4" pageNumber="418" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C370651DFFE5FFA9FF02FC311BF7FC73" box="[151,235,943,969]" pageId="4" pageNumber="418" type="description">
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<paragraph id="8BD53696FFE5FFA9FF02FC311BF7FC73" blockId="4.[151,453,906,969]" box="[151,235,943,969]" pageId="4" pageNumber="418">
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(
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C370651DFFE5FFA9FF02FC691AD5FB7F" pageId="4" pageNumber="418" type="materials_examined">
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<paragraph id="8BD53696FFE5FFA9FF02FC691BDCFBE3" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
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<materialsCitation id="3B023CCBFFE5FFA9FF02FC691AFAFB8F" collectingDate="2019-06-21" collectionCode="T" collectorName="la Caleta & K. A. Whorrall & K. Philips" country="Peru" latitude="-16.488" location="Caleta del Inca" longLatPrecision="7" longitude="-72.9465" pageId="4" pageNumber="418" specimenCount="1" stateProvince="Arequipa" typeStatus="holotype">
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<emphasis id="B91EEA84FFE5FFA9FF02FC691A18FBAB" bold="true" box="[151,260,1015,1041]" pageId="4" pageNumber="418">
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<typeStatus id="54D18834FFE5FFA9FF02FC691A18FBAB" box="[151,260,1015,1041]" pageId="4" pageNumber="418" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="F37D7606FFE5FFA9FE87FC691A56FBAB" box="[274,330,1015,1041]" name="Peru" pageId="4" pageNumber="418">Peru</collectingCountry>
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:
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<collectingRegion id="49AEF874FFE5FFA9FEC4FC691908FBAB" box="[337,532,1015,1041]" country="Peru" name="Arequipa" pageId="4" pageNumber="418">Arequipa Region</collectingRegion>
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, nr.
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<collectorName id="269F5340FFE5FFA9FDD5FC6919BEFBAB" box="[576,674,1015,1041]" pageId="4" pageNumber="418">
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la
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</collectorName>
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del Inca, dune veg. litter
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<date id="FFD41056FFE5FFA9FC2FFC691F20FBAB" box="[954,1084,1015,1041]" pageId="4" pageNumber="418" value="2019-06-21">
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<collectingDate id="EF90E9BEFFE5FFA9FC2FFC691F20FBAB" box="[954,1084,1015,1041]" pageId="4" pageNumber="418" value="2019-06-21">21.VI.2019</collectingDate>
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, 2570 m
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<collectorName id="269F5340FFE5FFA9FF02FB851A2EFB8F" box="[151,306,1051,1077]" pageId="4" pageNumber="418">K.A.Whorrall</collectorName>
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&
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.
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<collectorName id="269F5340FFE5FFA9FEE4FB851AFAFB8F" box="[369,486,1051,1077]" pageId="4" pageNumber="418">K. Philips</collectorName>
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.
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<materialsCitation id="3B023CCBFFE5FFA9FE66FB8518A2FB8F" box="[499,958,1051,1077]" collectionCode="MUSM" country="Peru" pageId="4" pageNumber="418" specimenCount="1" stateProvince="Arequipa" typeStatus="holotype">
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<typeStatus id="54D18834FFE5FFA9FE66FB851947FB8F" box="[499,603,1051,1077]" pageId="4" pageNumber="418" type="holotype">Holotype</typeStatus>
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has been deposited in
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<collectionCode id="ED7BAE53FFE5FFA9FCF0FB8518A2FB8F" box="[869,958,1051,1077]" pageId="4" pageNumber="418">MUSM</collectionCode>
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.
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<emphasis id="B91EEA84FFE5FFA9FC5EFB851F95FB8E" bold="true" box="[971,1161,1050,1077]" pageId="4" pageNumber="418">
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<typeStatus id="54D18834FFE5FFA9FC5EFB851F5FFB8F" box="[971,1091,1051,1077]" pageId="4" pageNumber="418" type="paratype">Paratypes</typeStatus>
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(131)
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</emphasis>
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: Same data as holotype
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<emphasis id="B91EEA84FFE5FFA9FF02FBA01BA4FBE2" bold="true" box="[151,184,1086,1112]" pageId="4" pageNumber="418">(7)</emphasis>
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;
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</paragraph>
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<paragraph id="8BD53696FFE5FFA9FF52FBFD1AD5FB7F" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
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<collectingCountry id="F37D7606FFE5FFA9FF52FBFD1BE3FBC7" box="[199,255,1123,1149]" name="Peru" pageId="4" pageNumber="418">Peru</collectingCountry>
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:
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<collectingRegion id="49AEF874FFE5FFA9FE9CFBFD1AD2FBC7" box="[265,462,1123,1149]" country="Peru" name="Arequipa" pageId="4" pageNumber="418">Arequipa Region</collectingRegion>
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,
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<location id="8EB5604DFFE5FFA9FE4FFBFD1967FBC7" LSID="urn:lsid:plazi:treatment:03C38780FFE5FFA9FF02FC141E7FF867:8EB5604DFFE5FFA9FE4FFBFD1967FBC7" box="[474,635,1123,1149]" country="Peru" name="Ocona Lomas" pageId="4" pageNumber="418" stateProvince="Arequipa">Ocoña Lomas</location>
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,
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<date id="FFD41056FFE5FFA9FD12FBFD19EDFBC7" box="[647,753,1123,1149]" pageId="4" pageNumber="418" value="2019-06-18" valueMax="2019-06-22" valueMin="2019-06-18">
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18–22.
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<collectionCode id="ED7BAE53FFE5FFA9FD47FBFD19EDFBC7" box="[722,753,1123,1149]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="418">VI</collectionCode>
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</date>
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<materialsCitation id="3B023CCBFFE5FFA9FD64FBFD1AB6FB1B" collectionCode="T" country="Peru" elevation="725" latitude="-16.4214" location="Whorrall" longLatPrecision="7" longitude="-73.049" pageId="4" pageNumber="418" specimenCount="1" stateProvince="Arequipa Region">
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.2019
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<geoCoordinate id="EE5E5051FFE5FFA9FCACFBFD18B3FBC7" box="[825,943,1123,1149]" degrees="16.4214" direction="south" orientation="latitude" pageId="4" pageNumber="418" precision="5" value="-16.4214">S16.4214°</geoCoordinate>
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</quantity>
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scattered veg, dung
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<collectingMethod id="522B4E81FFE5FFA9FAE0FBFD1BDBFB1B" pageId="4" pageNumber="418">pitfalls</collectingMethod>
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K.A.
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<location id="8EB5604DFFE5FFA9FE96FB191A72FB1B" LSID="urn:lsid:plazi:treatment:03C38780FFE5FFA9FF02FC141E7FF867:8EB5604DFFE5FFA9FE96FB191A72FB1B" box="[259,366,1159,1185]" country="Peru" latitude="-16.4214" longLatPrecision="7" longitude="-73.049" name="Whorrall" pageId="4" pageNumber="418" stateProvince="Arequipa Region">Whorrall</location>
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&
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<collectionCode id="ED7BAE53FFE5FFA9FE0CFB191AB6FB1B" box="[409,426,1159,1185]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="418">T</collectionCode>
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</materialsCitation>
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.
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<materialsCitation id="3B023CCBFFE5FFA9FE3BFB191A5EFB7F" collectionCode="MUSM, USNM, NHMUK, TAMU, KAWC, TKPC" country="Peru" location="Philips" pageId="4" pageNumber="418" specimenCount="1" stateProvince="Arequipa Region" typeStatus="paratype">
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K.
|
||
<location id="8EB5604DFFE5FFA9FE53FB191904FB1B" LSID="urn:lsid:plazi:treatment:03C38780FFE5FFA9FF02FC141E7FF867:8EB5604DFFE5FFA9FE53FB191904FB1B" box="[454,536,1159,1185]" country="Peru" name="Philips" pageId="4" pageNumber="418" stateProvince="Arequipa Region">Philips</location>
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<emphasis id="B91EEA84FFE5FFA9FDB6FB181943FB1A" bold="true" box="[547,607,1158,1184]" pageId="4" pageNumber="418">(122)</emphasis>
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.
|
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<typeStatus id="54D18834FFE5FFA9FDE4FB1919C2FB1B" box="[625,734,1159,1185]" pageId="4" pageNumber="418" type="paratype">Paratypes</typeStatus>
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have been deposited in
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<collectionCode id="ED7BAE53FFE5FFA9FB95FB191F44FB1B" box="[1024,1112,1159,1185]" pageId="4" pageNumber="418">MUSM</collectionCode>
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,
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<collectionCode id="ED7BAE53FFE5FFA9FBF2FB191FA7FB1B" box="[1127,1211,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="418" type="Museum">USNM</collectionCode>
|
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,
|
||
<collectionCode id="ED7BAE53FFE5FFA9FB5FFB191E2BFB1B" box="[1226,1335,1159,1185]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="418" type="Museum">NHMUK</collectionCode>
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,
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<collectionCode id="ED7BAE53FFE5FFA9FAD0FB191E84FB1B" box="[1349,1432,1159,1185]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;M University" pageId="4" pageNumber="418" type="Herbarium">TAMU</collectionCode>
|
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,
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||
<collectionCode id="ED7BAE53FFE5FFA9FF02FB351BF1FB7F" box="[151,237,1195,1221]" pageId="4" pageNumber="418">KAWC</collectionCode>
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,
|
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<collectionCode id="ED7BAE53FFE5FFA9FF6DFB351A5EFB7F" box="[248,322,1195,1221]" pageId="4" pageNumber="418">TKPC</collectionCode>
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</materialsCitation>
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, and
|
||
<collectionCode id="ED7BAE53FFE5FFA9FEE8FB351AD9FB7F" box="[381,453,1195,1221]" pageId="4" pageNumber="418">JBHC</collectionCode>
|
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.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C370651DFFE5FFA9FF52FB501F29FAEF" pageId="4" pageNumber="418" type="diagnosis">
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<paragraph id="8BD53696FFE5FFA9FF52FB501F29FAEF" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52FB501A58FB52" bold="true" box="[199,324,1230,1256]" pageId="4" pageNumber="418">Diagnosis:</emphasis>
|
||
This species is distinguished from its congeners by the pronounced carinae on its elytra with 4–5 rows of deep, open depressions between each. It can be distinguished from
|
||
<taxonomicName id="4C6A4D15FFE5FFA9FC75FB6D1F61FAB7" authorityName="Whorrall & Philips" authorityYear="2021" box="[992,1149,1267,1293]" class="Insecta" family="Ptinidae" genus="Cordielytrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="fimbriatum">
|
||
<emphasis id="B91EEA84FFE5FFA9FC75FB6D1F61FAB7" box="[992,1149,1267,1293]" italics="true" pageId="4" pageNumber="418">C. fimbriatum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13512A13FFE5FFA9FB19FB6D1FD2FAB7" box="[1164,1230,1266,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1978,2002]" captionTargetBox="[196,1391,596,1953]" captionTargetId="figure-175@6.[196,1391,596,1953]" captionTargetPageId="6" captionText="FIGURE 5. Cordielytrum fimbriatum. A Dorsal habitus, B ventral sclerites, C lateral view, D frontal close-up. Scale bars: 1.0 mm (A, C), 0.5 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.5531520" httpUri="https://zenodo.org/record/5531520/files/figure.png" pageId="4" pageNumber="418">Fig. 5</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C6A4D15FFE5FFA9FA98FB6D1E80FAB7" authorityName="Whorrall & Philips" authorityYear="2021" box="[1293,1436,1267,1293]" class="Insecta" family="Ptinidae" genus="Cordielytrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="obscurum">
|
||
<emphasis id="B91EEA84FFE5FFA9FA98FB6D1E80FAB7" box="[1293,1436,1267,1293]" italics="true" pageId="4" pageNumber="418">C. obscurum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13512A13FFE5FFA9FF35FA891BFFFA88" box="[160,227,1303,1330]" captionStart="FIGURE 6" captionStartId="7.[151,250,1966,1990]" captionTargetBox="[175,1412,597,1942]" captionTargetId="figure-160@7.[175,1412,597,1942]" captionTargetPageId="7" captionText="FIGURE 6. Cordielytrum obscurum. A Dorsal habitus, B ventral sclerites, C lateral view, D frontal close-up. Scale bars: 1.0 mm (A, C), 0.5 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.5531524" httpUri="https://zenodo.org/record/5531524/files/figure.png" pageId="4" pageNumber="418">Fig. 6</figureCitation>
|
||
) by its simple erect elytral setae and from
|
||
<taxonomicName id="4C6A4D15FFE5FFA9FD50FA891859FA8B" authorityName="Whorrall & Philips" authorityYear="2021" box="[709,837,1303,1329]" class="Insecta" family="Ptinidae" genus="Cordielytrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="418" phylum="Arthropoda" rank="species" species="adustum">
|
||
<emphasis id="B91EEA84FFE5FFA9FD50FA891859FA8B" box="[709,837,1303,1329]" italics="true" pageId="4" pageNumber="418">C. adustum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13512A13FFE5FFA9FCC0FA891884FA8B" box="[853,920,1303,1329]" captionStart="FIGURE 7" captionStartId="9.[151,250,1567,1591]" captionTargetBox="[157,1431,181,1544]" captionTargetId="figure-20@9.[157,1431,181,1544]" captionTargetPageId="9" captionText="FIGURE 7. Cordielytrum adustum. A Dorsal habitus, B ventral sclerites, C lateral view, D frontal close-up. Scale bars: 0.5 mm (A, C), 0.25 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.5531526" httpUri="https://zenodo.org/record/5531526/files/figure.png" pageId="4" pageNumber="418">Fig. 7</figureCitation>
|
||
) by its larger size, the more discrete light and dark patches of coloration on its elytra, and its more strongly transverse pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C370651DFFE5FFA9FF52FAC01BE7F803" pageId="4" pageNumber="418" type="description">
|
||
<paragraph id="8BD53696FFE5FFA9FF52FAC019C2FA27" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52FAC01A4DFAC2" bold="true" box="[199,337,1374,1400]" pageId="4" pageNumber="418">Description</emphasis>
|
||
. Body small, compact, subovate, convex; mottled tan and brown with a slight sheen. Length (anterior of pronotum to apex of elytra)
|
||
<quantity id="4C929B73FFE5FFA9FDA1FA1D19C6FA27" box="[564,730,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.6799999999999997" metricValueMax="3.13" metricValueMin="2.23" pageId="4" pageNumber="418" unit="mm" value="2.6799999999999997" valueMax="3.13" valueMin="2.23">2.23–3.13 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD53696FFE5FFA9FF52FA391A5DF9EB" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52FA391A1BFA7B" bold="true" box="[199,263,1447,1473]" pageId="4" pageNumber="418">Head</emphasis>
|
||
completely covered with light tan, depressed, ovoid scales; pronounced superantennal carinae, with a small number of recumbent, apically expanded, ligulate setae; antennae stout, antennomeres 1–10 densely squamous, apical antennomere with simple setae, antennomeres 1–4 and 11 slightly longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with moderately long ligulate scales pointing toward middle.
|
||
</paragraph>
|
||
<paragraph id="8BD53696FFE5FFA9FF52F9C51BE8F95B" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52F9C51A5CF9CF" bold="true" box="[199,320,1627,1653]" pageId="4" pageNumber="418">Pronotum</emphasis>
|
||
covered with densely matted setae, short, erect setae protruding from deep cavities within matted setation, dorsally truncate, bifurcate; two shallow anterolateral depressions; medial cavity shallow, broad, extending about ½ total length and about 1/3 width of pronotum, borders rounded; two posterolateral cavities on each side shallow.
|
||
</paragraph>
|
||
<paragraph id="8BD53696FFE5FFA9FF52F9751FABF8CB" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52F9751A01F8BF" bold="true" box="[199,285,1771,1797]" pageId="4" pageNumber="418">Elytra:</emphasis>
|
||
Setation silvery tan-gray with large distinct black patches, background color dark brown; surface densely covered in small scoop-shaped scales; basal ¼ of anterolateral margin with row of very long bristly setae; five highly prominent longitudinal carinae extending length of each elytron, including one at suture, each carina with one row of very short truncate setae; ~4 rows of deep, open depressions between carinae.
|
||
</paragraph>
|
||
<paragraph id="8BD53696FFE5FFA9FF52F8E51BE7F803" blockId="4.[151,1437,1014,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52F8E51A9FF82E" bold="true" box="[199,387,1914,1941]" pageId="4" pageNumber="418">Ventral Surface</emphasis>
|
||
: Ventral part of pro-, meso, and, metathorax, legs, and abdominal ventrites with scales similar to elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C370651DFFE5FFA9FF52F85D1E7FF867" box="[199,1379,1987,2013]" pageId="4" pageNumber="418" type="etymology">
|
||
<paragraph id="8BD53696FFE5FFA9FF52F85D1E7FF867" blockId="4.[151,1437,1014,2013]" box="[199,1379,1987,2013]" pageId="4" pageNumber="418">
|
||
<emphasis id="B91EEA84FFE5FFA9FF52F85D1A54F867" bold="true" box="[199,328,1987,2013]" pageId="4" pageNumber="418">Etymology</emphasis>
|
||
: The name “inca” is from the beach la Caleta del Inca near the collection site of the
|
||
<typeStatus id="54D18834FFE5FFA9FB6CF85D1E43F867" box="[1273,1375,1987,2013]" pageId="4" pageNumber="418" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |