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treatments-xml/data/03/BA/58/03BA5863DA58A329B2A38E90C4B5B8C1.xml
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<document id="20ED8B28D778FA2C040EE1F243AF0067" ID-CLB-Dataset="24684" ID-DOI="10.11646/zootaxa.4706.2.9" ID-GBIF-Dataset="62014a8c-1b33-4609-8f33-656de4f3fdb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3567473" ID-ZooBank="28FD7782-BE38-4412-A9B3-F2E6B5914119" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575876454148" checkinUser="plazi" docAuthor="Huang, Gui-Qiang &amp; Wang, Fa-Lei" docDate="2019" docId="03BA5863DA58A329B2A38E90C4B5B8C1" docLanguage="en" docName="zootaxa.4706.2.9.pdf" docOrigin="Zootaxa 4706 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala qiului Huang &amp; Wang 2019, new species" docType="treatment" docVersion="8" lastPageNumber="370" masterDocId="FF83201BDA59A32DB2348E71C062B94A" masterDocTitle="Two new and one newly recorded species of Anomala Samouelle, 1819 (Coleoptera Scarabaeidae: Rutelinae) from Yunnan, China" masterLastPageNumber="374" masterPageNumber="366" pageNumber="367" updateTime="1698773612123" updateUser="ExternalLinkService">
<mods:mods id="504CB0651DC40406231E1A6CBEAE6ABF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="67136E3DF360F39B49F01150DF416294">
<mods:title id="8D5E2D05E010EC6BF281A5447243415A">Two new and one newly recorded species of Anomala Samouelle, 1819 (Coleoptera Scarabaeidae: Rutelinae) from Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name id="156EBF482AC0C6253B78FACD9C5973FD" type="personal">
<mods:role id="9FDAC5632F559B5142A7EA334789EA68">
<mods:roleTerm id="CF7480A27C8D482DD8DBF6227F835CD5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F57AFB76A505B6A9A73937C5F069F00F">Huang, Gui-Qiang</mods:namePart>
</mods:name>
<mods:name id="D7DECC9729E3CE8DEBA9D150AB221FC6" type="personal">
<mods:role id="54DBE31C88C4E3446700E421E97EBC8B">
<mods:roleTerm id="3017B9E241DB82DAF4E4BBEFCBB33932">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6F908D49E649C218AA9DCBEF00B799A">Wang, Fa-Lei</mods:namePart>
</mods:name>
<mods:typeOfResource id="C30E8AE21AE20C0907A0E57F54BAD31A">text</mods:typeOfResource>
<mods:relatedItem id="2288B7BA9BC6C4A8568FC5972B2FF51C" type="host">
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<mods:title id="5619F7C52BBB76C41CA990E83D421BCD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4C7EEA758C552859B068D6125C22A458">
<mods:date id="CE19DD53A2D8CDE1937EB864B270DB20">2019</mods:date>
<mods:detail id="4C12CBCE9A463F87AB11CACD97381BD7" type="pubDate">
<mods:number id="93A102E8CB36AFCF1F338100817F7B4C">2019-12-09</mods:number>
</mods:detail>
<mods:detail id="8D79B036951755B8F5D28BADB3029C6A" type="volume">
<mods:number id="011B46416C2FD48DB9B1F23293CFC26B">4706</mods:number>
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<mods:number id="29BC21A573098DED66D769BE898F71CE">2</mods:number>
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</mods:extent>
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<mods:classification id="0DD3FE04E7DB3C54703DD8BF58AE2560">journal article</mods:classification>
<mods:identifier id="97BDA8243A30523F870E6447DB94C564" type="CLB-Dataset">24684</mods:identifier>
<mods:identifier id="FA7C65BECD712DB1090CF0871904FDD3" type="DOI">10.11646/zootaxa.4706.2.9</mods:identifier>
<mods:identifier id="07C51BF4BD20A5EBD890FF079604563A" type="GBIF-Dataset">62014a8c-1b33-4609-8f33-656de4f3fdb5</mods:identifier>
<mods:identifier id="076B3B7A902E0AEE3A98CE7D8888E8A5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BE7155A7807BA0B8E8ED5CE0424C8A5F" type="Zenodo-Dep">3567473</mods:identifier>
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<treatment id="03BA5863DA58A329B2A38E90C4B5B8C1" ID-DOI="http://doi.org/10.5281/zenodo.5931354" ID-GBIF-Taxon="160676584" ID-Zenodo-Dep="5931354" LSID="urn:lsid:plazi:treatment:03BA5863DA58A329B2A38E90C4B5B8C1" httpUri="http://treatment.plazi.org/id/03BA5863DA58A329B2A38E90C4B5B8C1" lastPageId="4" lastPageNumber="370" pageId="1" pageNumber="367">
<subSubSection id="C309BAFEDA58A32CB2A38E90C2A2B9B1" box="[151,704,225,252]" pageId="1" pageNumber="367" type="nomenclature">
<paragraph id="8BACE975DA58A32CB2A38E90C2A2B9B1" blockId="1.[151,704,225,288]" box="[151,704,225,252]" pageId="1" pageNumber="367">
<heading id="D0E45E19DA58A32CB2A38E90C2A2B9B1" bold="true" box="[151,704,225,252]" fontSize="11" level="1" pageId="1" pageNumber="367" reason="1">
<emphasis id="B9673567DA58A32CB2A38E90C2A2B9B1" bold="true" box="[151,704,225,252]" pageId="1" pageNumber="367">
<taxonomicName id="4C1392F6DA58A32CB2A38E90C241B9B6" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2019" box="[151,547,225,252]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA58A32CB2A38E90C137B9B6" bold="true" box="[151,341,225,252]" italics="true" pageId="1" pageNumber="367">Anomala qiului</emphasis>
Huang &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="A254881CDA58A32CB0048E93C2A2B9B1" box="[560,704,225,252]" pageId="1" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309BAFEDA58A32CB2A38F74C186B855" box="[151,484,261,288]" pageId="1" pageNumber="367" type="description">
<paragraph id="8BACE975DA58A32CB2A38F74C186B855" blockId="1.[151,704,225,288]" box="[151,484,261,288]" pageId="1" pageNumber="367">
(
<figureCitation id="1328F5F0DA58A32CB2AB8F74C168B855" box="[159,266,261,287]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1910,1935]" captionTargetBox="[195,1399,181,1874]" captionTargetId="figure@2.[180,1407,179,1882]" captionTargetPageId="2" captionText="FIGURES 1–9. Anomala qiului Huang &amp; Wang, new species (1–4), Anomala fuscicauda Lin, 1999 (5) and Anomala trochanterica Arrow, 1917 (6–9). 1, habitus, holotype, ♂, dorsal view; 2, habitus, holotype, ♂, ventral view; 3, habitus, paratype, ♀, dorsal view; 4, habitus, paratype, ♀, ventral view; 5, habitus, ♂ from Yunnan, China, dorsal view; 6, 7, habitus, ♂ from Yun- nan, China, dorsal view; 8, habitus, ♂ from Sichuan, China, dorsal view; 9, habitus, ♂ from Xizang, China, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3567475" httpUri="https://zenodo.org/record/3567475/files/figure.png" pageId="1" pageNumber="367">Figs. 1–4</figureCitation>
,
<figureCitation id="1328F5F0DA58A32CB3238F74C1D1B855" box="[279,435,261,288]" captionStart="FIGURES 10–28" captionStartId="3.[151,264,1826,1851]" captionTargetBox="[206,1368,226,1777]" captionTargetId="figure@3.[170,1416,181,1801]" captionTargetPageId="3" captionText="FIGURES 10–28. Anomala species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28). Anomala qiului Huang &amp; Wang, new species, holotype (10, 15, 20) and ♀ paratype (28), Anomala fuscicauda Lin, 1999 from Yunnan, China (11, 16, 21), Anomala trochanterica Arrow, 1917 from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24), Anomala merula Arrow, 1917 from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and Anomala validipes Arrow 1917 from Assam, India (26, captured from Arrow, 1917)." figureDoi="http://doi.org/10.5281/zenodo.3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="1" pageNumber="367">10, 15, 20, 28</figureCitation>
,
<figureCitation id="1328F5F0DA58A32CB38B8F74C1BEB855" box="[447,476,261,287]" captionStart="FIGURE 44" captionStartId="7.[151,250,1731,1756]" captionTargetBox="[151,1412,910,1705]" captionTargetId="figure@7.[151,1436,885,1705]" captionTargetPageId="7" captionText="FIGURE 44. Distribution map of eight species of Anomala in this study." figureDoi="http://doi.org/10.5281/zenodo.3567481" httpUri="https://zenodo.org/record/3567481/files/figure.png" pageId="1" pageNumber="367">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C309BAFEDA58A32CB2A38F3FC428B899" pageId="1" pageNumber="367" type="materials_examined">
<paragraph id="8BACE975DA58A32CB2A38F3FC428B899" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="367">
<materialsCitation id="3B7BE328DA58A32CB2A38F3FC2BAB8C1" ID-GBIF-Occurrence="2556817703" collectingDate="2018-04-02" collectionCode="MYNU" collectorName="Lu Qiu" country="China" county="Yingjiang County" elevation="640" location="Nabang" longitude="97.940636" municipality="Dehong" pageId="1" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B9673567DA58A32CB2A38F3FC1AEB822" bold="true" box="[151,460,333,360]" pageId="1" pageNumber="367">
Type material.
<typeStatus id="54A857D7DA58A32CB3628F3FC1A5B822" box="[342,455,334,360]" pageId="1" pageNumber="367" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D1522FCDA58A32CB3EC8F3FC18FB822" box="[472,493,334,360]" pageId="1" pageNumber="367" type="male">♂</specimenCount>
(
<collectionCode id="ED0271B0DA58A32CB0358F3CC23CB82D" box="[513,606,333,359]" pageId="1" pageNumber="367">MYNU</collectionCode>
):
<location id="8ECCBFAEDA58A32CB0418F3CC2ADB82D" LSID="urn:lsid:plazi:treatment:03BA5863DA58A329B2A38E90C4B5B8C1:8ECCBFAEDA58A32CB0418F3CC2ADB82D" box="[629,719,333,359]" country="China" county="Yingjiang County" longitude="97.940636" municipality="Dehong" name="Nabang" pageId="1" pageNumber="367" stateProvince="Yunnan">Nabang</location>
,
<collectingCounty id="62CD91F9DA58A32CB0EA8F3CC3CEB82D" box="[734,940,333,359]" pageId="1" pageNumber="367">Yingjiang County</collectingCounty>
,
<collectingMunicipality id="6BC8730FDA58A32CB1888F3CC47AB82D" box="[956,1048,333,359]" pageId="1" pageNumber="367">Dehong</collectingMunicipality>
,
<collectingRegion id="49D72797DA58A32CB6138F3FC4E2B822" box="[1063,1152,334,360]" country="China" name="Yunnan" pageId="1" pageNumber="367">Yunnan</collectingRegion>
,
<collectingCountry id="F304A9E5DA58A32CB6BB8F3CC4B7B82D" box="[1167,1237,333,359]" name="China" pageId="1" pageNumber="367">China</collectingCountry>
,
<geoCoordinate id="EE278FB2DA58A32CB6D18F3CC5F5B822" box="[1253,1431,333,360]" degrees="97" direction="east" minutes="56" orientation="longitude" pageId="1" pageNumber="367" precision="1" seconds="26.28" value="97.940636">97°56’26.28” E</geoCoordinate>
,
<geoCoordinate id="EE278FB2DA58A32CB2A38F00C12FB8C6" box="[151,333,369,396]" degrees="24" direction="north" minutes="75" orientation="latitude" pageId="1" pageNumber="367" seconds="04.56">24°75’04.56” N</geoCoordinate>
, alt.
<quantity id="4CEB4490DA58A32CB3B78F00C1AFB8C6" box="[387,461,369,396]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="367" unit="m" value="640.0">
<elevation id="003E0E46DA58A32CB3B78F00C1AFB8C6" box="[387,461,369,396]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="367" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="FFADCFB5DA58A32CB3ED8F00C228B8C1" box="[473,586,369,395]" pageId="1" pageNumber="367" value="2018-04-02">
<collectingDate id="EFE9365DDA58A32CB3ED8F00C228B8C1" box="[473,586,369,395]" pageId="1" pageNumber="367" value="2018-04-02">2.IV.2018</collectingDate>
</date>
,
<collectorName id="26E68CA3DA58A32CB0628F03C2C5B8C1" box="[598,679,369,396]" pageId="1" pageNumber="367">Lu Qiu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B7BE328DA58A32CB0D48F03C5FAB8C1" ID-GBIF-Occurrence="2465037471" box="[736,1432,369,396]" collectingDate="2018-04-02" collectionCode="CFLW" collectorName="Lu Qiu" country="China" county="Yingjiang County" elevation="640" location="Nabang" longitude="97.940636" municipality="Dehong" pageId="1" pageNumber="367" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B9673567DA58A32CB0D48F03C303B8C6" bold="true" box="[736,865,370,396]" pageId="1" pageNumber="367">
<typeStatus id="54A857D7DA58A32CB0D48F03C33EB8C6" box="[736,860,370,396]" pageId="1" pageNumber="367" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D1522FCDA58A32CB15D8F00C3EEB8C1" box="[873,908,369,395]" pageId="1" pageNumber="367" type="male">1♂</specimenCount>
and
<specimenCount id="9D1522FCDA58A32CB1F08F00C39EB8C1" box="[964,1020,369,395]" pageId="1" pageNumber="367" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED0271B0DA58A32CB63F8F00C43CB8C1" box="[1035,1118,369,395]" pageId="1" pageNumber="367">CFLW</collectionCode>
): same data as the holotype
</materialsCitation>
;
<materialsCitation id="3B7BE328DA58A32CB2A38FE4C49FB8E5" ID-GBIF-Occurrence="2465037453" box="[151,1277,405,432]" collectingDate="2013-05-09" collectionCode="CMZZ" collectorName="Xiao-Dong Yang" country="China" county="Yingjiang County" location="Nabang" municipality="Dehong" pageId="1" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D1522FCDA58A32CB2A38FE4C0D8B8E5" box="[151,186,405,431]" pageId="1" pageNumber="367" type="male">1♂</specimenCount>
(
<collectionCode id="ED0271B0DA58A32CB2FF8FE4C142B8E5" box="[203,288,405,431]" pageId="1" pageNumber="367">CMZZ</collectionCode>
):
<location id="8ECCBFAEDA58A32CB3018FE4C1EDB8E5" LSID="urn:lsid:plazi:treatment:03BA5863DA58A329B2A38E90C4B5B8C1:8ECCBFAEDA58A32CB3018FE4C1EDB8E5" box="[309,399,405,431]" country="China" county="Yingjiang County" municipality="Dehong" name="Nabang" pageId="1" pageNumber="367" stateProvince="Yunnan">Nabang</location>
,
<collectingCounty id="62CD91F9DA58A32CB3AF8FE4C204B8E5" box="[411,614,405,431]" pageId="1" pageNumber="367">Yingjiang County</collectingCounty>
,
<collectingMunicipality id="6BC8730FDA58A32CB0478FE4C2ADB8E5" box="[627,719,405,431]" pageId="1" pageNumber="367">Dehong</collectingMunicipality>
,
<collectingRegion id="49D72797DA58A32CB0EF8FE7C356B8FA" box="[731,820,406,432]" country="China" name="Yunnan" pageId="1" pageNumber="367">Yunnan</collectingRegion>
,
<collectingCountry id="F304A9E5DA58A32CB1748FE4C3E4B8E5" box="[832,902,405,431]" name="China" pageId="1" pageNumber="367">China</collectingCountry>
,
<date id="FFADCFB5DA58A32CB1A78FE4C399B8E5" box="[915,1019,405,431]" pageId="1" pageNumber="367" value="2013-05-09">
<collectingDate id="EFE9365DDA58A32CB1A78FE4C399B8E5" box="[915,1019,405,431]" pageId="1" pageNumber="367" value="2013-05-09">9.V.2013</collectingDate>
</date>
,
<collectorName id="26E68CA3DA58A32CB63C8FE4C4AAB8FA" box="[1032,1224,405,432]" pageId="1" pageNumber="367">Xiao-Dong Yang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B7BE328DA58A32CB73F8FE4C428B899" ID-GBIF-Occurrence="2465037451" collectingDate="2018-05-08" collectionCode="CCCC" collectorName="Xiao-Dong Yang" country="China" county="Yingjiang County" location="Nabang" municipality="Dehong" pageId="1" pageNumber="367" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D1522FCDA58A32CB73F8FE4C54CB8E5" box="[1291,1326,405,431]" pageId="1" pageNumber="367" type="female">1♀</specimenCount>
(
<collectionCode id="ED0271B0DA58A32CB70B8FE4C5F3B8E5" box="[1343,1425,405,431]" country="USA" httpUri="http://grbio.org/cool/3dng-x47a" name="Carthage College" pageId="1" pageNumber="367">CCCC</collectionCode>
):
<location id="8ECCBFAEDA58A32CB2A38FC8C093B899" LSID="urn:lsid:plazi:treatment:03BA5863DA58A329B2A38E90C4B5B8C1:8ECCBFAEDA58A32CB2A38FC8C093B899" box="[151,241,441,467]" country="China" county="Yingjiang County" municipality="Dehong" name="Nabang" pageId="1" pageNumber="367" stateProvince="Yunnan">Nabang</location>
,
<collectingCounty id="62CD91F9DA58A32CB2CF8FC8C1A7B899" box="[251,453,441,467]" pageId="1" pageNumber="367">Yingjiang County</collectingCounty>
,
<collectingMunicipality id="6BC8730FDA58A32CB3FB8FC8C249B899" box="[463,555,441,467]" pageId="1" pageNumber="367">Dehong</collectingMunicipality>
,
<collectingRegion id="49D72797DA58A32CB0018FCBC2ECB89E" box="[565,654,442,468]" country="China" name="Yunnan" pageId="1" pageNumber="367">Yunnan</collectingRegion>
,
<collectingCountry id="F304A9E5DA58A32CB0AC8FC8C2BCB899" box="[664,734,441,467]" name="China" pageId="1" pageNumber="367">China</collectingCountry>
,
<date id="FFADCFB5DA58A32CB0DD8FC8C333B899" box="[745,849,441,467]" pageId="1" pageNumber="367" value="2018-05-08">
<collectingDate id="EFE9365DDA58A32CB0DD8FC8C333B899" box="[745,849,441,467]" pageId="1" pageNumber="367" value="2018-05-08">8.V.2018</collectingDate>
</date>
,
<collectorName id="26E68CA3DA58A32CB1688FC8C478B89E" box="[860,1050,441,468]" pageId="1" pageNumber="367">Xiao-Dong Yang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309BAFEDA58A329B2F38FACC388B855" lastPageId="4" lastPageNumber="370" pageId="1" pageNumber="367" type="description">
<paragraph id="8BACE975DA58A32CB2F38FACC16BBCAD" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="367">
<emphasis id="B9673567DA58A32CB2F38FACC13BB8B2" bold="true" box="[199,345,477,504]" pageId="1" pageNumber="367">Description.</emphasis>
Male,
<typeStatus id="54A857D7DA58A32CB39D8FACC269B8BD" box="[425,523,477,503]" pageId="1" pageNumber="367" type="holotype">holotype</typeStatus>
(
<figureCitation id="1328F5F0DA58A32CB02F8FACC2E5B8BD" box="[539,647,477,503]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1910,1935]" captionTargetBox="[195,1399,181,1874]" captionTargetId="figure@2.[180,1407,179,1882]" captionTargetPageId="2" captionText="FIGURES 1–9. Anomala qiului Huang &amp; Wang, new species (1–4), Anomala fuscicauda Lin, 1999 (5) and Anomala trochanterica Arrow, 1917 (6–9). 1, habitus, holotype, ♂, dorsal view; 2, habitus, holotype, ♂, ventral view; 3, habitus, paratype, ♀, dorsal view; 4, habitus, paratype, ♀, ventral view; 5, habitus, ♂ from Yunnan, China, dorsal view; 6, 7, habitus, ♂ from Yun- nan, China, dorsal view; 8, habitus, ♂ from Sichuan, China, dorsal view; 9, habitus, ♂ from Xizang, China, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3567475" httpUri="https://zenodo.org/record/3567475/files/figure.png" pageId="1" pageNumber="367">Figs. 1–2</figureCitation>
,
<figureCitation id="1328F5F0DA58A32CB0A08FACC368B8BD" box="[660,778,477,504]" captionStart="FIGURES 10–28" captionStartId="3.[151,264,1826,1851]" captionTargetBox="[206,1368,226,1777]" captionTargetId="figure@3.[170,1416,181,1801]" captionTargetPageId="3" captionText="FIGURES 10–28. Anomala species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28). Anomala qiului Huang &amp; Wang, new species, holotype (10, 15, 20) and ♀ paratype (28), Anomala fuscicauda Lin, 1999 from Yunnan, China (11, 16, 21), Anomala trochanterica Arrow, 1917 from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24), Anomala merula Arrow, 1917 from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and Anomala validipes Arrow 1917 from Assam, India (26, captured from Arrow, 1917)." figureDoi="http://doi.org/10.5281/zenodo.3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="1" pageNumber="367">10, 15, 20</figureCitation>
).
<emphasis id="B9673567DA58A32CB12A8FACC3E7B8B2" bold="true" box="[798,901,477,504]" pageId="1" pageNumber="367">General.</emphasis>
Length
<quantity id="4CEB4490DA58A32CB1D08FACC429B8B2" box="[996,1099,477,504]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="367" unit="mm" value="13.6">13.6 mm</quantity>
, greatest width
<quantity id="4CEB4490DA58A32CB6CA8FACC53AB8B2" box="[1278,1368,477,504]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="1" pageNumber="367" unit="mm" value="9.4">9.4 mm</quantity>
; body elongate ovoid, moderately convex in profile, sub-lustrous.
<emphasis id="B9673567DA58A32CB1188C70C317BB56" bold="true" box="[812,885,513,540]" pageId="1" pageNumber="367">Color.</emphasis>
Whole body black, antennae brown.
<emphasis id="B9673567DA58A32CB7248C73C535BB56" bold="true" box="[1296,1367,514,540]" pageId="1" pageNumber="367">Head.</emphasis>
Clypeus sub-trapezoidal, rounded apically, with dense and fine punctures, anterior margin slightly reflexed towards dorsum; clypeofrontal suture extremely weak; frons with dense punctures (sparse at base), the punctures large on disc, small near eyes; interocular distance 0.63 times as wide as the maximum transverse width of head; antenna sparsely covered with yellow hairs, antennal club shorter than antennomeres 2–6.
<emphasis id="B9673567DA58A32CB6188CE3C4CEBBE6" bold="true" box="[1068,1196,658,684]" pageId="1" pageNumber="367">Pronotum.</emphasis>
1.5 times as broad as long, sides expanded in middle; basal part of the sides with short marginal line, basal marginal line not interrupted in the middle, gradually disappearing towards the sides; anterior angles obtuse, posterior angles rounded apically; disc densely covered with punctures but densest at sides; punctures on sides moderately large, gradually smaller towards anterior angles; with a distinct longitudinal smooth area from basal margin to the center of disc; with several long yellow setae along lateral margins.
<emphasis id="B9673567DA58A32CB0188D34C2C9BA2A" bold="true" box="[556,683,837,864]" pageId="1" pageNumber="367">Scutellum.</emphasis>
Tongue-shaped, 1.2 times as wide as long; with sparse punctures, punctures only dense basally; posterior margin smooth.
<emphasis id="B9673567DA58A32CB1218D1BC30ABACE" bold="true" box="[789,872,874,900]" pageId="1" pageNumber="367">Elytra.</emphasis>
Approximately 1.17 times as long wide, humeral umbone and apical protuberance prominent and smooth, strial punctures dense, large and deep, forming several moderately deep longitudinal striae, intervals between striae smooth; elytra with large, dense and irregular punctures near base of suture, with small, sparse and rounded punctures close to sides of scutellum; epipleuron starting apical of level of humeral umbone, flat and gradually expanded at basal 1/2, ending at apical 1/4.
<emphasis id="B9673567DA58A32CB6D28D88C502BD5E" bold="true" box="[1254,1376,1017,1044]" pageId="1" pageNumber="367">Pygidium.</emphasis>
Subtriangular and moderately convex in profile, with transverse punctures on disc, punctures gradually sparser towards sides; posterior marginal line complete, with brown setae at posterior margin.
<emphasis id="B9673567DA58A32CB62C8A30C575BD16" bold="true" box="[1048,1303,1089,1116]" pageId="1" pageNumber="367">Abdominal ventrites.</emphasis>
Abdominal ventrites 1–3 with fine punctures, the punctures denser in center and at sides; abdominal ventrite 4 with moderately dense transverse punctures, gradually finer from center to sides, denser in center and at sides; abdominal ventrite 5 with dense, fine punctures; abdominal ventrites 1–3 with a transverse rows of sparse, short yellow-brown setae near middle, the setae slightly denser at sides; abdominal ventrite 4 with a row of short yellowish brown setae near apical margin; abdominal ventrite 5 with a row of yellowish brown setae near apex.
<emphasis id="B9673567DA58A32CB60F8A87C418BC5A" bold="true" box="[1083,1146,1270,1296]" pageId="1" pageNumber="367">Legs.</emphasis>
All tarsi short, metatarsus rather strong; protarsal inner claw and mesotarsal outer claw cleft, mesotarsal inner and metatarsal inner and outer claws simple and long; protibia bidentate, apical tooth produced and rounded apically, slightly extending beyond apex of first segment of tarsus, basal tooth short and obtuse, inner spur short; mesotibia slightly fusiform, metatibia very strong and fusiform; both mesotibia and metatibia densely covered with yellow hairs at inner sides, with two terminal spurs, the upper one 1.65 times as long as the lower one.
<emphasis id="B9673567DA58A32CB14D8BD8C38CBC8E" bold="true" box="[889,1006,1449,1476]" pageId="1" pageNumber="367">Genitalia.</emphasis>
As in
<figureCitation id="1328F5F0DA58A32CB6028BD8C484BC89" box="[1078,1254,1449,1476]" captionStart="FIGURES 10–28" captionStartId="3.[151,264,1826,1851]" captionTargetBox="[206,1368,226,1777]" captionTargetId="figure@3.[170,1416,181,1801]" captionTargetPageId="3" captionText="FIGURES 10–28. Anomala species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28). Anomala qiului Huang &amp; Wang, new species, holotype (10, 15, 20) and ♀ paratype (28), Anomala fuscicauda Lin, 1999 from Yunnan, China (11, 16, 21), Anomala trochanterica Arrow, 1917 from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24), Anomala merula Arrow, 1917 from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and Anomala validipes Arrow 1917 from Assam, India (26, captured from Arrow, 1917)." figureDoi="http://doi.org/10.5281/zenodo.3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="1" pageNumber="367">Figs. 10, 15, 20</figureCitation>
. See differential diagnosis.
</paragraph>
<paragraph id="8BACE975DA58A32CB2F38B80C2A0BF65" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="367">
<emphasis id="B9673567DA58A32CB2F38B80C12FBF46" bold="true" box="[199,333,1521,1548]" pageId="1" pageNumber="367">Variability.</emphasis>
Body length
<quantity id="4CEB4490DA58A32CB3D68B80C2E4BF46" box="[482,646,1521,1548]" metricMagnitude="-2" metricUnit="m" metricValue="1.305" metricValueMax="1.3599999999999999" metricValueMin="1.25" pageId="1" pageNumber="367" unit="mm" value="13.05" valueMax="13.6" valueMin="12.5">12.5–13.6 mm</quantity>
, greatest width
<quantity id="4CEB4490DA58A32CB1018B80C3D9BF46" box="[821,955,1521,1548]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.4" metricValueMin="9.1" pageId="1" pageNumber="367" unit="mm" value="9.25" valueMax="9.4" valueMin="9.1">9.1–9.4 mm</quantity>
in males. The
<typeStatus id="54A857D7DA58A32CB6698B80C4DDBF41" box="[1117,1215,1521,1547]" pageId="1" pageNumber="367" type="holotype">holotype</typeStatus>
and male
<typeStatus id="54A857D7DA58A32CB7058B83C5FEBF46" box="[1329,1436,1522,1548]" pageId="1" pageNumber="367" type="paratype">paratypes</typeStatus>
show no differences in the shape of the aedeagus.
</paragraph>
<paragraph id="8BACE975DA58A32CB2F3884BC512BFD1" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="367">
<emphasis id="B9673567DA58A32CB2F3884BC14BBF1E" bold="true" box="[199,297,1594,1620]" pageId="1" pageNumber="367">Females</emphasis>
(
<figureCitation id="1328F5F0DA58A32CB30D8848C1C5BF19" box="[313,423,1593,1619]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1910,1935]" captionTargetBox="[195,1399,181,1874]" captionTargetId="figure@2.[180,1407,179,1882]" captionTargetPageId="2" captionText="FIGURES 1–9. Anomala qiului Huang &amp; Wang, new species (1–4), Anomala fuscicauda Lin, 1999 (5) and Anomala trochanterica Arrow, 1917 (6–9). 1, habitus, holotype, ♂, dorsal view; 2, habitus, holotype, ♂, ventral view; 3, habitus, paratype, ♀, dorsal view; 4, habitus, paratype, ♀, ventral view; 5, habitus, ♂ from Yunnan, China, dorsal view; 6, 7, habitus, ♂ from Yun- nan, China, dorsal view; 8, habitus, ♂ from Sichuan, China, dorsal view; 9, habitus, ♂ from Xizang, China, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3567475" httpUri="https://zenodo.org/record/3567475/files/figure.png" pageId="1" pageNumber="367">Figs. 3, 4</figureCitation>
,
<figureCitation id="1328F5F0DA58A32CB3808848C1B6BF19" box="[436,468,1593,1619]" captionStart="FIGURES 10–28" captionStartId="3.[151,264,1826,1851]" captionTargetBox="[206,1368,226,1777]" captionTargetId="figure@3.[170,1416,181,1801]" captionTargetPageId="3" captionText="FIGURES 10–28. Anomala species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28). Anomala qiului Huang &amp; Wang, new species, holotype (10, 15, 20) and ♀ paratype (28), Anomala fuscicauda Lin, 1999 from Yunnan, China (11, 16, 21), Anomala trochanterica Arrow, 1917 from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24), Anomala merula Arrow, 1917 from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and Anomala validipes Arrow 1917 from Assam, India (26, captured from Arrow, 1917)." figureDoi="http://doi.org/10.5281/zenodo.3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="1" pageNumber="367">28</figureCitation>
). Similar to male, but body larger, body length 19.7–20.0 mm, greatest width
<quantity id="4CEB4490DA58A32CB77C8848C0A4BF32" metricMagnitude="-3" metricUnit="m" metricValue="9.85" metricValueMax="9.9" metricValueMin="9.8" pageId="1" pageNumber="367" unit="mm" value="9.85" valueMax="9.9" valueMin="9.8">9.8–9.9 mm</quantity>
; antennal club shorter, apical margin of pronotum wider (compare with basal margin); apical tooth of protibia more rounded apically, distinctly extending beyond apex of second segment of tarsus, basal tooth right-angled.
</paragraph>
<paragraph id="8BACE975DA58A329B2F388D4C388B855" blockId="1.[151,1437,333,2016]" lastBlockId="4.[151,1436,153,395]" lastPageId="4" lastPageNumber="370" pageId="1" pageNumber="367">
<emphasis id="B9673567DA58A32CB2F388D4C1ADBF8A" bold="true" box="[199,463,1701,1728]" pageId="1" pageNumber="367">Differential diagnosis.</emphasis>
Around Himalaya region (
<collectingCountry id="F304A9E5DA58A32CB13588D4C32ABFF5" box="[769,840,1701,1727]" name="China" pageId="1" pageNumber="367">China</collectingCountry>
,
<collectingCountry id="F304A9E5DA58A32CB16188D4C3F3BFF5" box="[853,913,1701,1727]" name="India" pageId="1" pageNumber="367">India</collectingCountry>
,
<collectingCountry id="F304A9E5DA58A32CB1A988D4C38CBFF5" box="[925,1006,1701,1727]" name="Bhutan" pageId="1" pageNumber="367">Bhutan</collectingCountry>
and
<collectingCountry id="F304A9E5DA58A32CB61388D7C4FABF8A" box="[1063,1176,1702,1728]" name="Myanmar" pageId="1" pageNumber="367">Myanmar</collectingCountry>
), four morphologically similar species,
<taxonomicName id="4C1392F6DA58A32CB37A88B8C2CDBFA9" authority="Lin, 1999" authorityName="Lin" authorityYear="1999" box="[334,687,1737,1763]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="fuscicauda">
<emphasis id="B9673567DA58A32CB37A88B8C256BFA9" box="[334,564,1737,1763]" italics="true" pageId="1" pageNumber="367">Anomala fuscicauda</emphasis>
<bibRefCitation id="EF829484DA58A32CB00988B8C2CDBFA9" author="Lin, P." box="[573,687,1737,1763]" pageId="1" pageNumber="367" pagination="157 - 176" refId="ref5108" refString="Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxonomia, 21, 157 - 176." type="journal article" year="1999">Lin, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1392F6DA58A32CB08F88BBC3A6BFA9" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[699,964,1737,1764]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="merula">
<emphasis id="B9673567DA58A32CB08F88BBC349BFA9" box="[699,811,1737,1764]" italics="true" pageId="1" pageNumber="367">A. merula</emphasis>
<bibRefCitation id="EF829484DA58A32CB10688BBC3A6BFA9" author="Arrow, G. J." box="[818,964,1737,1764]" pageId="1" pageNumber="367" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1392F6DA58A32CB1E588BBC57CBFA9" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[977,1310,1737,1764]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA58A32CB1E588BBC4EABFA9" box="[977,1160,1737,1764]" italics="true" pageId="1" pageNumber="367">A. trochanterica</emphasis>
<bibRefCitation id="EF829484DA58A32CB6A488BBC57CBFA9" author="Arrow, G. J." box="[1168,1310,1737,1764]" pageId="1" pageNumber="367" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1392F6DA58A32CB76C88BBC11BBE4D" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="validipes">
<emphasis id="B9673567DA58A32CB76C88BBC080BE4D" italics="true" pageId="1" pageNumber="367">A. validipes</emphasis>
<bibRefCitation id="EF829484DA58A32CB2DE889FC11BBE4D" author="Arrow, G. J." box="[234,377,1773,1800]" pageId="1" pageNumber="367" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
share following characters with the new species: pronotum attenuated anteriorly, causing the anterior somewhat tapered body shape, sides of pronotum expanded in middle, posterior angles rounded apically; all tarsi short, hind legs extremely strong (compare with fore and mid legs). However,
<taxonomicName id="4C1392F6DA58A32CB6438947C4BABE05" authority="Huang &amp; Wang, 2019" authorityName="Huang &amp; Wang" authorityYear="2019" box="[1143,1240,1845,1872]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA58A32CB6438947C4EABE1A" box="[1143,1160,1846,1872]" italics="true" pageId="1" pageNumber="367">A</emphasis>
.
<emphasis id="B9673567DA58A32CB6A38944C4BABE05" box="[1175,1240,1845,1871]" italics="true" pageId="1" pageNumber="367">qiului</emphasis>
</taxonomicName>
<emphasis id="B9673567DA58A32CB6D58947C50EBE1A" bold="true" box="[1249,1388,1845,1872]" pageId="1" pageNumber="367">
<taxonomicNameLabel id="A254881CDA58A32CB6D58947C50EBE1A" box="[1249,1388,1845,1872]" pageId="1" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
can be distinguished from the four similar species by extremely asymmetric parameres of male genitalia (
<figureCitation id="1328F5F0DA58A32CB7338928C5F2BE3E" box="[1287,1424,1881,1908]" captionStart="FIGURES 10–28" captionStartId="3.[151,264,1826,1851]" captionTargetBox="[206,1368,226,1777]" captionTargetId="figure@3.[170,1416,181,1801]" captionTargetPageId="3" captionText="FIGURES 10–28. Anomala species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28). Anomala qiului Huang &amp; Wang, new species, holotype (10, 15, 20) and ♀ paratype (28), Anomala fuscicauda Lin, 1999 from Yunnan, China (11, 16, 21), Anomala trochanterica Arrow, 1917 from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24), Anomala merula Arrow, 1917 from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and Anomala validipes Arrow 1917 from Assam, India (26, captured from Arrow, 1917)." figureDoi="http://doi.org/10.5281/zenodo.3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="1" pageNumber="367">Figs. 10–26</figureCitation>
). Except for male genitalia,
<taxonomicName id="4C1392F6DA58A32CB38E890FC27BBEDD" authority="Huang &amp; Wang, 2019" authorityName="Huang &amp; Wang" authorityYear="2019" box="[442,537,1917,1944]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA58A32CB38E890FC27BBEDD" box="[442,537,1917,1944]" italics="true" pageId="1" pageNumber="367">A. qiului</emphasis>
</taxonomicName>
<emphasis id="B9673567DA58A32CB02B890FC2CABED2" bold="true" box="[543,680,1917,1944]" pageId="1" pageNumber="367">
<taxonomicNameLabel id="A254881CDA58A32CB02B890FC2CABED2" box="[543,680,1917,1944]" pageId="1" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1392F6DA58A32CB10D890FC3C4BEDD" authorityName="Arrow" authorityYear="1917" box="[825,934,1917,1944]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="merula">
<emphasis id="B9673567DA58A32CB10D890FC3C4BEDD" box="[825,934,1917,1944]" italics="true" pageId="1" pageNumber="367">A. merula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1392F6DA58A32CB1EE890FC4EDBEDD" authorityName="Arrow" authorityYear="1917" box="[986,1167,1917,1944]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA58A32CB1EE890FC4EDBEDD" box="[986,1167,1917,1944]" italics="true" pageId="1" pageNumber="367">A. trochanterica</emphasis>
</taxonomicName>
by the disc of pronotum with deeper punctures, elytra with large and irregular dense punctures near base of suture, with deeper striae and more convex intervals. Besides, females of
<taxonomicName id="4C1392F6DA58A32CB0B889B7C28DBE95" authority="Huang &amp; Wang, 2019" authorityName="Huang &amp; Wang" authorityYear="2019" box="[652,751,1989,2016]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA58A32CB0B889B7C28DBE95" box="[652,751,1989,2016]" italics="true" pageId="1" pageNumber="367">A. qiului</emphasis>
</taxonomicName>
<emphasis id="B9673567DA58A32CB0CC89B7C3E1BEAA" bold="true" box="[760,899,1989,2016]" pageId="1" pageNumber="367">
<taxonomicNameLabel id="A254881CDA58A32CB0CC89B7C3E1BEAA" box="[760,899,1989,2016]" pageId="1" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C1392F6DA58A32CB68A89B7C515BE95" authorityName="Arrow" authorityYear="1917" box="[1214,1399,1989,2016]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA58A32CB68A89B7C515BE95" box="[1214,1399,1989,2016]" italics="true" pageId="1" pageNumber="367">A. trochanterica</emphasis>
</taxonomicName>
by two simple vaginal palps, whereas
<taxonomicName id="4C1392F6DA5DA329B0128EE8C2BCB9F9" authorityName="Arrow" authorityYear="1917" box="[550,734,153,179]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA5DA329B0128EE8C2BCB9F9" box="[550,734,153,179]" italics="true" pageId="4" pageNumber="370">A. trochanterica</emphasis>
</taxonomicName>
with two additional sclerites in the membrane connecting the vaginal palps.
<taxonomicName id="4C1392F6DA5DA329B30E8ECCC185B99D" authority="Huang &amp; Wang, 2019" authorityName="Huang &amp; Wang" authorityYear="2019" box="[314,487,189,215]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA5DA329B30E8ECCC185B99D" box="[314,487,189,215]" italics="true" pageId="4" pageNumber="370">Anomala qiului</emphasis>
</taxonomicName>
<emphasis id="B9673567DA5DA329B3DA8ECCC215B99D" bold="true" box="[494,631,188,215]" pageId="4" pageNumber="370">
<taxonomicNameLabel id="A254881CDA5DA329B3DA8ECCC215B99D" box="[494,631,188,215]" pageId="4" pageNumber="370" rank="species">new species</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1392F6DA5DA329B13F8ECCC3C1B99D" authorityName="Lin" authorityYear="1999" box="[779,931,189,215]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="fuscicauda">
<emphasis id="B9673567DA5DA329B13F8ECCC3C1B99D" box="[779,931,189,215]" italics="true" pageId="4" pageNumber="370">A. fuscicauda</emphasis>
</taxonomicName>
by body black (brown in
<taxonomicName id="4C1392F6DA5DA329B6F08ECCC53EB99D" authorityName="Lin" authorityYear="1999" box="[1220,1372,189,215]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="fuscicauda">
<emphasis id="B9673567DA5DA329B6F08ECCC53EB99D" box="[1220,1372,189,215]" italics="true" pageId="4" pageNumber="370">A. fuscicauda</emphasis>
</taxonomicName>
), and elytra with several moderately deep and regular longitudinal striae and smooth intervals.
<taxonomicName id="4C1392F6DA5DA329B6B48E90C54FB9B1" authority="Huang &amp; Wang, 2019" authorityName="Huang &amp; Wang" authorityYear="2019" box="[1152,1325,225,251]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA5DA329B6B48E90C54FB9B1" box="[1152,1325,225,251]" italics="true" pageId="4" pageNumber="370">Anomala qiului</emphasis>
</taxonomicName>
<emphasis id="B9673567DA5DA329B7018E90C0A1B855" bold="true" pageId="4" pageNumber="370">
<taxonomicNameLabel id="A254881CDA5DA329B7018E90C0A1B855" pageId="4" pageNumber="370" rank="species">new species</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C1392F6DA5DA329B3638F74C1B9B855" authorityName="Arrow" authorityYear="1917" box="[343,475,261,287]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="validipes">
<emphasis id="B9673567DA5DA329B3638F74C10AB855" box="[343,360,261,287]" italics="true" pageId="4" pageNumber="370">A</emphasis>
.
<emphasis id="B9673567DA5DA329B3428F74C1B9B855" box="[374,475,261,287]" italics="true" pageId="4" pageNumber="370">validipes</emphasis>
</taxonomicName>
by body black (reddish brown in
<taxonomicName id="4C1392F6DA5DA329B1628F74C3B8B855" authorityName="Arrow" authorityYear="1917" box="[854,986,261,287]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="validipes">
<emphasis id="B9673567DA5DA329B1628F74C305B855" box="[854,871,261,287]" italics="true" pageId="4" pageNumber="370">A</emphasis>
.
<emphasis id="B9673567DA5DA329B1418F74C3B8B855" box="[885,986,261,287]" italics="true" pageId="4" pageNumber="370">validipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF6CB9FDDA5BA32FB2A38907C53ABEB0" ID-DOI="http://doi.org/10.5281/zenodo.3567475" ID-Zenodo-Dep="3567475" httpUri="https://zenodo.org/record/3567475/files/figure.png" pageId="2" pageNumber="368" startId="2.[151,264,1910,1935]" targetBox="[195,1399,181,1874]" targetPageId="2">
<paragraph id="8BACE975DA5BA32FB2A38907C53ABEB0" blockId="2.[151,1437,1910,2043]" pageId="2" pageNumber="368">
<emphasis id="B9673567DA5BA32FB2A38907C15FBEC5" bold="true" box="[151,317,1910,1935]" pageId="2" pageNumber="368">FIGURES 1–9.</emphasis>
<taxonomicName id="4C1392F6DA5BA32FB3738906C2F4BEC5" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2019" box="[327,662,1911,1935]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA5BA32FB3738906C185BEC5" box="[327,487,1911,1935]" italics="true" pageId="2" pageNumber="368">Anomala qiului</emphasis>
Huang &amp; Wang
</taxonomicName>
,
<emphasis id="B9673567DA5BA32FB0908906C341BEC5" bold="true" box="[676,803,1910,1935]" pageId="2" pageNumber="368">
<taxonomicNameLabel id="A254881CDA5BA32FB0908906C341BEC5" box="[676,803,1910,1935]" pageId="2" pageNumber="368" rank="species">new species</taxonomicNameLabel>
</emphasis>
(1–4),
<taxonomicName id="4C1392F6DA5BA32FB1478906C4D7BEC5" authority="Lin, 1999" authorityName="Lin" authorityYear="1999" box="[883,1205,1910,1935]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="fuscicauda">
<emphasis id="B9673567DA5BA32FB1478906C424BEC5" box="[883,1094,1911,1935]" italics="true" pageId="2" pageNumber="368">Anomala fuscicauda</emphasis>
<bibRefCitation id="EF829484DA5BA32FB67B8907C4D7BEC5" author="Lin, P." box="[1103,1205,1910,1935]" pageId="2" pageNumber="368" pagination="157 - 176" refId="ref5108" refString="Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxonomia, 21, 157 - 176." type="journal article" year="1999">Lin, 1999</bibRefCitation>
</taxonomicName>
(5) and
<taxonomicName id="4C1392F6DA5BA32FB7268906C1EBBEF9" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA5BA32FB7268906C161BEF9" italics="true" pageId="2" pageNumber="368">Anomala trochanterica</emphasis>
<bibRefCitation id="EF829484DA5BA32FB33D89EAC1EBBEF9" author="Arrow, G. J." box="[265,393,1947,1971]" pageId="2" pageNumber="368" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
(6–9). 1, habitus, holotype, ♂, dorsal view; 2, habitus, holotype, ♂, ventral view; 3, habitus, paratype, ♀, dorsal view; 4, habitus, paratype, ♀, ventral view; 5, habitus, ♂ from Yunnan, China, dorsal view; 6, 7, habitus, ♂ from Yun- nan, China, dorsal view; 8, habitus, ♂ from Sichuan, China, dorsal view; 9, habitus, ♂ from Xizang, China, dorsal view.
</paragraph>
</caption>
<caption id="DF6CB9FDDA5AA32EB2A38953C13FBEA5" ID-DOI="http://doi.org/10.5281/zenodo.3567477" ID-Zenodo-Dep="3567477" httpUri="https://zenodo.org/record/3567477/files/figure.png" pageId="3" pageNumber="369" startId="3.[151,264,1826,1851]" targetBox="[206,1368,226,1777]" targetPageId="3">
<paragraph id="8BACE975DA5AA32EB2A38953C13FBEA5" blockId="3.[151,1436,1826,2031]" pageId="3" pageNumber="369">
<emphasis id="B9673567DA5AA32EB2A38953C136BE71" bold="true" box="[151,340,1826,1851]" pageId="3" pageNumber="369">FIGURES 10–28.</emphasis>
<taxonomicName id="4C1392F6DA5AA32EB36D8952C1D6BE71" authorityName="Samouelle" authorityYear="1819" box="[345,436,1827,1851]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B9673567DA5AA32EB36D8952C1D6BE71" box="[345,436,1827,1851]" italics="true" pageId="3" pageNumber="369">Anomala</emphasis>
</taxonomicName>
species, aedeagus in dorsal (10–14), ventral (15–19) and lateral (20–26) view, female genital palps (27, 28).
<taxonomicName id="4C1392F6DA5AA32EB2C38936C223BE2A" authority="Huang &amp; Wang" authorityName="Huang &amp; Wang" authorityYear="2019" box="[247,577,1863,1888]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="qiului" status="sp. nov.">
<emphasis id="B9673567DA5AA32EB2C38936C1F4BE15" box="[247,406,1863,1887]" italics="true" pageId="3" pageNumber="369">Anomala qiului</emphasis>
Huang &amp; Wang
</taxonomicName>
,
<emphasis id="B9673567DA5AA32EB0798936C2A9BE15" bold="true" box="[589,715,1862,1887]" pageId="3" pageNumber="369">
<taxonomicNameLabel id="A254881CDA5AA32EB0798936C2A9BE15" box="[589,715,1862,1887]" pageId="3" pageNumber="369" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype (10, 15, 20) and ♀ paratype (28),
<taxonomicName id="4C1392F6DA5AA32EB6AE8936C0A8BEC9" authority="Lin, 1999" authorityName="Lin" authorityYear="1999" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="fuscicauda">
<emphasis id="B9673567DA5AA32EB6AE8936C509BE15" box="[1178,1387,1863,1887]" italics="true" pageId="3" pageNumber="369">Anomala fuscicauda</emphasis>
<bibRefCitation id="EF829484DA5AA32EB7478937C0A8BEC9" author="Lin, P." pageId="3" pageNumber="369" pagination="157 - 176" refId="ref5108" refString="Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxonomia, 21, 157 - 176." type="journal article" year="1999">Lin, 1999</bibRefCitation>
</taxonomicName>
from Yunnan, China (11, 16, 21),
<taxonomicName id="4C1392F6DA5AA32EB01F891AC3FEBEC9" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[555,924,1899,1923]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="trochanterica">
<emphasis id="B9673567DA5AA32EB01F891AC374BEC9" box="[555,790,1899,1923]" italics="true" pageId="3" pageNumber="369">Anomala trochanterica</emphasis>
<bibRefCitation id="EF829484DA5AA32EB128891AC3FEBEC9" author="Arrow, G. J." box="[796,924,1899,1923]" pageId="3" pageNumber="369" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
from Sichuan (12, 17, 22, 27), same species from Xizang (13, 18, 23), same species from Yunnan (14, 19, 24),
<taxonomicName id="4C1392F6DA5AA32EB12189FEC42BBEED" authority="Arrow, 1917" authorityName="Arrow" authorityYear="1917" box="[789,1097,1935,1959]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="merula">
<emphasis id="B9673567DA5AA32EB12189FEC3A3BEED" box="[789,961,1935,1959]" italics="true" pageId="3" pageNumber="369">Anomala merula</emphasis>
<bibRefCitation id="EF829484DA5AA32EB1F389FEC42BBEED" author="Arrow, G. J." box="[967,1097,1935,1959]" pageId="3" pageNumber="369" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
</taxonomicName>
from Assam, India (25, redrawn from a photograph of syntype provided by Carsten Zorn) and
<taxonomicName id="4C1392F6DA5AA32EB13989C2C42EBE81" authority="Arrow 1917" authorityName="Arrow" authorityYear="1917" box="[781,1100,1971,1995]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="validipes">
<emphasis id="B9673567DA5AA32EB13989C2C3A9BE81" box="[781,971,1971,1995]" italics="true" pageId="3" pageNumber="369">Anomala validipes</emphasis>
<bibRefCitation id="EF829484DA5AA32EB1E489C2C42EBE81" author="Arrow, G. J." box="[976,1100,1971,1995]" pageId="3" pageNumber="369" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow 1917</bibRefCitation>
</taxonomicName>
from Assam, India (26, captured from
<bibRefCitation id="EF829484DA5AA32EB2F989A6C130BEA5" author="Arrow, G. J." box="[205,338,2007,2031]" pageId="3" pageNumber="369" refId="ref4783" refString="Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London, 387 pp." type="book" year="1917">Arrow, 1917</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="C309BAFEDA5DA329B2F38F58C253B82D" pageId="4" pageNumber="370" type="etymology">
<paragraph id="8BACE975DA5DA329B2F38F58C253B82D" blockId="4.[151,1436,153,395]" pageId="4" pageNumber="370">
<emphasis id="B9673567DA5DA329B2F38F58C12CB809" bold="true" box="[199,334,297,323]" pageId="4" pageNumber="370">Etymology.</emphasis>
The new species is named after Dr. Lu Qiu (DZṜ) from Southwest University,
<collectingRegion id="49D72797DA5DA329B6FD8F58C52AB809" box="[1225,1352,297,323]" country="China" name="Chongqing" pageId="4" pageNumber="370">Chongqing</collectingRegion>
,
<collectingCountry id="F304A9E5DA5DA329B7668F58C5FAB809" box="[1362,1432,297,323]" name="China" pageId="4" pageNumber="370">China</collectingCountry>
, who collected most of
<typeStatus id="54A857D7DA5DA329B3A28F3CC1A4B82D" box="[406,454,333,359]" pageId="4" pageNumber="370">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="C309BAFEDA5DA329B2F38F01C4B5B8C1" box="[199,1239,368,395]" pageId="4" pageNumber="370" type="distribution">
<paragraph id="8BACE975DA5DA329B2F38F01C4B5B8C1" blockId="4.[151,1436,153,395]" box="[199,1239,368,395]" pageId="4" pageNumber="370">
<emphasis id="B9673567DA5DA329B2F38F01C103B8C1" bold="true" box="[199,353,368,395]" pageId="4" pageNumber="370">Distribution.</emphasis>
The new species is only known from Nabang county,
<collectingRegion id="49D72797DA5DA329B1F48F00C4E0B8C1" box="[960,1154,369,395]" country="China" name="Yunnan" pageId="4" pageNumber="370">Yunnan province</collectingRegion>
,
<collectingCountry id="F304A9E5DA5DA329B6B98F00C4B6B8C1" box="[1165,1236,369,395]" name="China" pageId="4" pageNumber="370">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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