treatments-xml/data/03/B9/7F/03B97F78FFC4FFDBFF94FD9FF6F7F9CF.xml

<document id="3C33FD90BA6B2EA1964490EC427ED42D" ID-DOI="10.5281/zenodo.5340349" ID-ISSN="0374-1036" ID-Zenodo-Dep="5340349" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630186132833" checkinUser="felipe" docAuthor="Ježek, Jan &amp; Tkoč, Michal" docDate="2012" docId="03B97F78FFC4FFDBFF94FD9FF6F7F9CF" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.545-557.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Gondwanoscurus socotrensis Ježek &amp; Tkoč 2012, sp. nov." docType="treatment" docVersion="5" lastPageNumber="554" masterDocId="FF800700FFC7FFD2FE1EFFA5F75CFFFC" masterDocTitle="A new species of the genus Gondwanoscurus, and two new records of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Socotra Island" masterLastPageNumber="557" masterPageNumber="545" pageNumber="548" updateTime="1699179815582" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A90B189AE6AA483547C62D297531B27D">A new species of the genus Gondwanoscurus, and two new records of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Socotra Island</mods:title>
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<mods:name id="661683222E5F71FB8E5413D99F6866FB" type="personal">
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<mods:namePart id="A85167F89ACA83CF6DD8C3EF53E3C054">Ježek, Jan</mods:namePart>
<mods:affiliation id="042B7C436CA17F6BD97B17F58586B310">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jan. jezek @ quick. cz</mods:affiliation>
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<mods:namePart id="D64B7036F293B552D70D6DB40B4CA626">Tkoč, Michal</mods:namePart>
<mods:affiliation id="8F745BAA07B6C83EC7683C0CC61A2420">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jan. jezek @ quick. cz &amp; Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Praha 2, Czech Republic; e-mail: michaltkoc @ gmail. com</mods:affiliation>
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<mods:title id="3FC56023FEDC62F2AFE4C2A138A8173B">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="9D78CFD1B9F09B2F1E0D550B961DC6E0">2012</mods:date>
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<heading id="D0E77902FFC4FFD1FF94FD9FF474FDAF" bold="true" box="[394,808,570,595]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="548" reason="3">
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<taxonomicName id="4C10B5EDFFC4FFD1FF94FD9FF597FDAF" ID-CoL="9J4PK" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[394,715,570,595]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC4FFD1FF94FD9FF597FDAF" bold="true" box="[394,715,570,595]" italics="true" pageId="3" pageNumber="548">Gondwanoscurus socotrensis</emphasis>
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<taxonomicNameLabel id="A257AF07FFC4FFD1FCCCFD9EF474FDAF" box="[722,808,571,595]" pageId="3" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BAFCE6EFFC4FFD1FC3BFDFCF5D1FD90" blockId="3.[549,653,601,620]" box="[549,653,601,620]" pageId="3" pageNumber="548">
(
<figureCitation id="132BD2EBFFC4FFD1FC35FDFCF5DAFD90" box="[555,646,601,620]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStartId-0="4.[151,190,1161,1180]" captionStartId-1="5.[113,152,1137,1156]" captionStartId-2="5.[111,150,1545,1564]" captionStartId-3="6.[151,190,1516,1535]" captionStartId-4="7.[113,152,1470,1489]" captionStartId-5="8.[151,190,1444,1463]" captionTargetBox-0="[207,1078,228,1132]" captionTargetBox-1="[166,1033,232,1114]" captionTargetBox-2="[113,1085,1237,1528]" captionTargetBox-3="[299,991,234,1472]" captionTargetBox-4="[163,1038,231,1441]" captionTargetBox-5="[243,1032,242,1398]" captionTargetId-0="figure-12@4.[151,1126,216,1140]" captionTargetId-1="figure-16@5.[125,1081,215,1126]" captionTargetId-2="figure-48@5.[113,1085,1237,1528]" captionTargetId-3="figure-12@6.[243,1033,219,1493]" captionTargetId-4="figure-16@7.[138,1062,208,1462]" captionTargetId-5="figure-12@8.[224,1053,220,1413]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="Figs. 1–2. Gondwanoscurus socotrensis sp. nov., wing (1 – male; 2 – female)." captionText-1="Figs. 3–6. Gondwanoscurus socotrensis sp. nov. 3 – head, male; 4 – frontal suture and facets, male; 5 – terminal lobes of labium, male; 6 – terminal (fourth) palpomere, female." captionText-2="Figs.7–9. Gondwanoscurus socotrensis sp. nov., female. 7 – genital chamber ventrally; 8 – same, laterally; 9 – same, anteriorly." captionText-3="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." captionText-4="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." captionText-5="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi-0="http://doi.org/10.5281/zenodo.5340351" figureDoi-1="http://doi.org/10.5281/zenodo.5340353" figureDoi-2="http://doi.org/10.5281/zenodo.5340355" figureDoi-3="http://doi.org/10.5281/zenodo.5340357" figureDoi-4="http://doi.org/10.5281/zenodo.5340359" figureDoi-5="http://doi.org/10.5281/zenodo.5340363" httpUri-0="https://zenodo.org/record/5340351/files/figure.png" httpUri-1="https://zenodo.org/record/5340353/files/figure.png" httpUri-2="https://zenodo.org/record/5340355/files/figure.png" httpUri-3="https://zenodo.org/record/5340357/files/figure.png" httpUri-4="https://zenodo.org/record/5340359/files/figure.png" httpUri-5="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">Figs. 1–33</figureCitation>
)
</paragraph>
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<subSubSection id="C30A9DE5FFC4FFD1FE6FFD26F7A8FCCA" pageId="3" pageNumber="548" type="materials_examined">
<paragraph id="8BAFCE6EFFC4FFD1FE6FFD26F371FD6A" blockId="3.[113,1090,642,822]" box="[113,1069,642,663]" pageId="3" pageNumber="548">
<emphasis id="B964127CFFC4FFD1FE6FFD26F7B6FD6A" bold="true" box="[113,234,642,662]" pageId="3" pageNumber="548">
<typeStatus id="54AB70CCFFC4FFD1FE6FFD26F7C2FD6A" box="[113,158,643,662]" pageId="3" pageNumber="548">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3078EFEFFC4FFD1FEF1FD21F671FD6B" box="[239,301,644,663]" name="Yemen" pageId="3" pageNumber="548">Yemen</collectingCountry>
, 
<collectingCountry id="F3078EFEFFC4FFD1FF2BFD26F6EEFD6A" box="[309,434,643,662]" name="Socotra Island" pageId="3" pageNumber="548">Socotra Island</collectingCountry>
, Al Haghier Mts., Scant Mt., 
<quantity id="4CE8638BFFC4FFD1FCB3FD26F5B2FD6B" box="[685,750,643,663]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="548" unit="m" value="1450.0">1450 m</quantity>
a.s.l., 
<geoCoordinate id="EE24A8A9FFC4FFD1FD3BFD26F420FD6B" box="[805,892,642,663]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="3" pageNumber="548" precision="92" value="12.576667">12°34.6′N</geoCoordinate>
<geoCoordinate id="EE24A8A9FFC4FFD1FD9CFD26F48BFD6B" box="[898,983,642,663]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="3" pageNumber="548" precision="92" value="54.025">54°01.5′E</geoCoordinate>
(
<figureCitation id="132BD2EBFFC4FFD1FDFCFD26F37EFD6A" box="[994,1058,643,662]" captionStart="Figs" captionStartId="10.[152,191,1487,1506]" captionTargetBox="[185,1097,219,1454]" captionTargetPageId="10" captionText="Figs. 34–35. Socotran localities. 34 – Yemen, Socotra Island, Al Haghier Mts., Scant Mt., 1450 m a.s.l., 12°34.6′N 54°01.5′E, type locality of Gondwanoscurus socotrensis sp. nov. 35 – Yemen, Socotra Island, Al Haghier Mts., wadi Madar, 1180–1230 m a.s.l., 12°33.2′N 54°00.4′E, a detail of the habitat, a typical dry shrubby slopes, biotopes suitable for light trapping. Photos by J. Suchomel (November 2010)." figureDoi="http://doi.org/10.5281/zenodo.5340365" httpUri="https://zenodo.org/record/5340365/files/figure.png" pageId="3" pageNumber="548">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="8BAFCE6EFFC4FFD1FE6FFD3BF7A8FCCA" blockId="3.[113,1090,642,822]" pageId="3" pageNumber="548">
<emphasis id="B964127CFFC4FFD1FE6FFD3BF7A4FD4D" bold="true" box="[113,248,669,689]" pageId="3" pageNumber="548">Type material.</emphasis>
<materialsCitation id="3B78C433FFC4FFD1FF1EFD3BF362FD37" collectingDate="2010-11-12" collectingDateMax="2010-11-13" collectingDateMin="2010-11-12" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Bezdek &amp; Cat. No. &amp; Inv. No." country="Yemen" elevation="1450" latitude="12.576667" location="Al Haghier Mts." longLatPrecision="129" longitude="54.025" municipality="Scant Mt." pageId="3" pageNumber="548" specimenCount="1" typeStatus="holotype">
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: J, 
<collectingCountry id="F3078EFEFFC4FFD1FF84FD3BF684FD4D" box="[410,472,670,689]" name="Yemen" pageId="3" pageNumber="548">Yemen</collectingCountry>
, 
<collectingCountry id="F3078EFEFFC4FFD1FFFCFD3BF53DFD4D" box="[482,609,670,689]" name="Socotra Island" pageId="3" pageNumber="548">Socotra Island</collectingCountry>
, 
<location id="8ECF98B5FFC4FFD1FC75FD3BF5A2FD4D" LSID="urn:lsid:plazi:treatment:03B97F78FFC4FFDBFF94FD9FF6F7F9CF:8ECF98B5FFC4FFD1FC75FD3BF5A2FD4D" box="[619,766,670,689]" country="Yemen" latitude="12.576667" longLatPrecision="129" longitude="54.025" municipality="Scant Mt." name="Al Haghier Mts." pageId="3" pageNumber="548">Al Haghier Mts.</location>
, 
<collectingMunicipality id="6BCB5414FFC4FFD1FD16FD3BF43CFD4D" box="[776,864,670,689]" pageId="3" pageNumber="548">Scant Mt.</collectingMunicipality>
, 
<elevation id="003D295DFFC4FFD1FD74FD3BF482FD4D" box="[874,990,670,689]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="548" unit="m" value="1450.0">
<quantity id="4CE8638BFFC4FFD1FD74FD3BF4F1FD4D" box="[874,941,670,689]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="548" unit="m" value="1450.0">1450 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="EE24A8A9FFC4FFD1FDF7FD3BF31CFD4D" box="[1001,1088,669,689]" degrees="12" direction="north" minutes="34.6" orientation="latitude" pageId="3" pageNumber="548" precision="92" value="12.576667">12°34.6′N</geoCoordinate>
<geoCoordinate id="EE24A8A9FFC4FFD1FE6FFD1DF794FD30" box="[113,200,696,716]" degrees="54" direction="east" minutes="01.5" orientation="longitude" pageId="3" pageNumber="548" precision="92" value="54.025">54°01.5′E</geoCoordinate>
. 
<date id="FFAEE8AEFFC4FFD1FECCFD1CF60AFD37" box="[210,342,696,716]" pageId="3" pageNumber="548" value="2010-11-12" valueMax="2010-11-13" valueMin="2010-11-12">
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</date>
, at light, 
<collectorName id="26E5ABB8FFC4FFD1FFB4FD1CF6AEFD37" box="[426,498,696,716]" pageId="3" pageNumber="548">J. Hájek</collectorName>
and 
<collectorName id="26E5ABB8FFC4FFD1FC00FD1CF52FFD37" box="[542,627,696,716]" pageId="3" pageNumber="548">J. Bezděk</collectorName>
leg. (
<collectionCode id="ED0156ABFFC4FFD1FCB8FD1CF5BAFD30" box="[678,742,697,716]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="548">NMPC</collectionCode>
, slide 
<collectorName id="26E5ABB8FFC4FFD1FD01FD1CF436FD30" box="[799,874,697,716]" pageId="3" pageNumber="548">Cat. No.</collectorName>
34568, 
<collectorName id="26E5ABB8FFC4FFD1FDAFFD1CF4A5FD30" box="[945,1017,697,716]" pageId="3" pageNumber="548">Inv. No.</collectorName>
19769)
</materialsCitation>
. 
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<typeStatus id="54AB70CCFFC4FFD1FE6FFD76F7B6FD1A" box="[113,234,723,742]" pageId="3" pageNumber="548" type="paratype">PARATYPES</typeStatus>
: 14 JJ and 
<specimenCount id="9D1605E7FFC4FFD1FF48FD76F6CCFD1A" box="[342,400,717,742]" pageId="3" pageNumber="548" type="female">13 ♀♀</specimenCount>
, same data as holotype (
<collectionCode id="ED0156ABFFC4FFD1FC7CFD76F5FFFD1A" box="[610,675,723,742]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="548">NMPC</collectionCode>
), slides 
<collectorName id="26E5ABB8FFC4FFD1FCF9FD76F46CFD1B" box="[743,816,723,743]" pageId="3" pageNumber="548">Cat. No.</collectorName>
34569−34595, 
<collectorName id="26E5ABB8FFC4FFD1FDA8FD71F4A0FD1B" box="[950,1020,724,743]" pageId="3" pageNumber="548">Inv. No.</collectorName>
19770– 19796 (
<typeStatus id="54AB70CCFFC4FFD1FEADFD4BF7A5FCFD" box="[179,249,750,769]" pageId="3" pageNumber="548" type="allotype">allotype</typeStatus>
34583/19784)
</materialsCitation>
; 
<materialsCitation id="3B78C433FFC4FFD1FF9DFD4BF7ACFCCA" collectingDate="2010-11-12" collectingDateMax="2010-11-14" collectingDateMin="2010-11-12" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Bezdek &amp; Cat. No. &amp; Inv. No." country="Yemen" elevation="1205" latitude="12.553333" location="Al Haghier Mts." longLatPrecision="129" longitude="54.006668" municipality="wadi Madar" pageId="3" pageNumber="548" specimenCount="23" specimenCount-female="23" typeStatus="paratype">
<location id="8ECF98B5FFC4FFD1FF9DFD4BF54FFCFD" LSID="urn:lsid:plazi:treatment:03B97F78FFC4FFDBFF94FD9FF6F7F9CF:8ECF98B5FFC4FFD1FF9DFD4BF54FFCFD" box="[387,531,750,769]" country="Yemen" latitude="12.553333" longLatPrecision="129" longitude="54.006668" municipality="wadi Madar" name="Al Haghier Mts." pageId="3" pageNumber="548">Al Haghier Mts.</location>
, 
<collectingMunicipality id="6BCB5414FFC4FFD1FC02FD4BF5D9FCFD" box="[540,645,750,769]" pageId="3" pageNumber="548">wadi Madar</collectingMunicipality>
, 
<elevation id="003D295DFFC4FFD1FC91FD4BF46AFCFD" box="[655,822,750,769]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.23" metricValueMin="1.18" pageId="3" pageNumber="548" unit="m" value="1205.0" valueMax="1230.0" valueMin="1180.0">
<quantity id="4CE8638BFFC4FFD1FC91FD4BF45BFCFD" box="[655,775,750,769]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.23" metricValueMin="1.18" pageId="3" pageNumber="548" unit="m" value="1205.0" valueMax="1230.0" valueMin="1180.0">1180−1230 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="EE24A8A9FFC4FFD1FD5FFD4BF4CBFCFD" box="[833,919,749,769]" degrees="12" direction="north" minutes="33.2" orientation="latitude" pageId="3" pageNumber="548" precision="92" value="12.553333">12°33.2′N</geoCoordinate>
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, 
<date id="FFAEE8AEFFC4FFD1FDE1FD4BF7E8FCE7" pageId="3" pageNumber="548" value="2010-11-12" valueMax="2010-11-14" valueMin="2010-11-12">
<collectingDate id="EFEA1146FFC4FFD1FDE1FD4BF7E8FCE7" pageId="3" pageNumber="548" value="2010-11-12" valueMax="2010-11-14" valueMin="2010-11-12">12.−14. xi.2010</collectingDate>
</date>
, at light, 3 JJ and 
<specimenCount id="9D1605E7FFC4FFD1FF7AFCACF6C3FCE7" box="[356,415,770,796]" pageId="3" pageNumber="548" type="female">23 ♀♀</specimenCount>
, 
<collectorName id="26E5ABB8FFC4FFD1FFB2FCACF6AFFCE7" box="[428,499,776,796]" pageId="3" pageNumber="548">J. Hájek</collectorName>
and 
<collectorName id="26E5ABB8FFC4FFD1FC01FCACF528FCE7" box="[543,628,776,796]" pageId="3" pageNumber="548">J. Bezděk</collectorName>
leg. (
<collectionCode id="ED0156ABFFC4FFD1FCB8FCACF5BAFCE0" box="[678,742,777,796]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="548">NMPC</collectionCode>
, slides 
<collectorName id="26E5ABB8FFC4FFD1FD39FCACF42EFCE0" box="[807,882,777,796]" pageId="3" pageNumber="548">Cat. No.</collectorName>
34596−34621, 
<collectorName id="26E5ABB8FFC4FFD1FDE4FCACF31EFCE0" box="[1018,1090,777,796]" pageId="3" pageNumber="548">Inv. No.</collectorName>
19797–19822)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A9DE5FFC4FFDBFE6FFCEBF7C0FB88" lastPageId="9" lastPageNumber="554" pageId="3" pageNumber="548" type="description">
<paragraph id="8BAFCE6EFFC4FFD1FE6FFCEBF446FABB" blockId="3.[113,1089,846,1549]" pageId="3" pageNumber="548">
<emphasis id="B964127CFFC4FFD1FE6FFCEBF7A7FC9A" bold="true" box="[113,251,846,870]" pageId="3" pageNumber="548">Description.</emphasis>
<emphasis id="B964127CFFC4FFD1FEE1FCF5F661FC9B" box="[255,317,848,871]" italics="true" pageId="3" pageNumber="548">Male.</emphasis>
Eyes contiguous (
<figureCitation id="132BD2EBFFC4FFD1FFE4FCEAF500FC9B" box="[506,604,847,871]" captionStart="Figs" captionStartId="5.[113,152,1137,1156]" captionTargetBox="[166,1033,232,1114]" captionTargetId="figure-16@5.[125,1081,215,1126]" captionTargetPageId="5" captionText="Figs. 3–6. Gondwanoscurus socotrensis sp. nov. 3 – head, male; 4 – frontal suture and facets, male; 5 – terminal lobes of labium, male; 6 – terminal (fourth) palpomere, female." figureDoi="http://doi.org/10.5281/zenodo.5340353" httpUri="https://zenodo.org/record/5340353/files/figure.png" pageId="3" pageNumber="548">Figs. 3, 4</figureCitation>
), touching for more than three facet diameters, facets hardly hexagonal or inconspicuously globular, eye-bridge of four facet rows, ratio of facet diameter to the width of basis of scape 1: 3, vertex pyramidal, rounded, vertex ratio (width versus high) 2.4: 1. Only one row of sockets of side setae above dorsal apices of eyes. Frontoclypeus with oval area of insertions of setulae, setula patch has dorso-ventral cleft. Antenna with 16 antennomeres. Scape cylindrical, 2.4 times as long as pedicel, pedicel almost globular (
<figureCitation id="132BD2EBFFC4FFD1FF3EFBAAF631FBDB" box="[288,365,1039,1063]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">Fig. 24</figureCitation>
); flagellomeres 2–13 with long necks, flagellomere 1 amphora-shaped (
<figureCitation id="132BD2EBFFC4FFD1FE67FB8AF794FBBB" box="[121,200,1071,1095]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">Fig. 24</figureCitation>
), flagellar nodes (2−12) conspicuously excentrically bulbous with great width of one side (
<figureCitation id="132BD2EBFFC4FFD1FEB2FBEAF65BFB9B" box="[172,263,1103,1127]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="3" pageNumber="548">Figs. 16</figureCitation>
, 
<figureCitation id="132BD2EBFFC4FFD1FF0DFBEAF605FB9A" box="[275,345,1102,1127]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">24, 25</figureCitation>
), last two flagellomeres with conical nodes, apical flagellomere with asymmetrical side protuberance and terminal long digit (
<figureCitation id="132BD2EBFFC4FFD1FCDEFBCAF452FB7B" box="[704,782,1135,1159]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="3" pageNumber="548">Fig. 16</figureCitation>
). Sensory filaments (ascoids) of flagellomeres multiple, arranged in ring (
<figureCitation id="132BD2EBFFC4FFD1FC59FB2AF5CBFB5A" box="[583,663,1166,1191]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">Fig. 25</figureCitation>
), threadlike, multibranched (3–6 arms). Length ratios of maxillary palpomeres 1.0: 2.0: 1.8: 2.4, palpomere 4 thinest, cylindrical, annulate (
<figureCitation id="132BD2EBFFC4FFD1FEC2FB6AF671FB1B" box="[220,301,1231,1255]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="3" pageNumber="548">Fig. 17</figureCitation>
), scales of maxillary palpomeres maculated (compare as well the scales of wing on 
<figureCitation id="132BD2EBFFC4FFD1FEF2FB4AF613FAF1" box="[236,335,1263,1293]" captionStart="Figs" captionStartId="4.[151,190,1161,1180]" captionTargetBox="[207,1078,228,1132]" captionTargetId="figure-12@4.[151,1126,216,1140]" captionTargetPageId="4" captionText="Figs. 1–2. Gondwanoscurus socotrensis sp. nov., wing (1 – male; 2 – female)." figureDoi="http://doi.org/10.5281/zenodo.5340351" httpUri="https://zenodo.org/record/5340351/files/figure.png" pageId="3" pageNumber="548">
Fig. 1, P 
<subScript id="1794CC2BFFC4FFD1FF59FB5BF613FAF1" attach="left" box="[327,335,1278,1293]" fontSize="6" pageId="3" pageNumber="548">1</subScript>
</figureCitation>
on 
<figureCitation id="132BD2EBFFC4FFD1FF69FB4AF69AFAFB" box="[375,454,1263,1287]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="3" pageNumber="548">Fig. 12</figureCitation>
, haltere on 
<figureCitation id="132BD2EBFFC4FFD1FC5CFB4AF5CCFAFB" box="[578,656,1263,1287]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="3" pageNumber="548">Fig. 19</figureCitation>
, surstylus on 
<figureCitation id="132BD2EBFFC4FFD1FD3CFB4AF42DFAFB" box="[802,881,1263,1287]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="3" pageNumber="548">Fig. 31</figureCitation>
). Terminal lobes of labium bulbous (
<figureCitation id="132BD2EBFFC4FFD1FF36FAAAF636FADA" box="[296,362,1294,1319]" captionStart="Figs" captionStartId="5.[113,152,1137,1156]" captionTargetBox="[166,1033,232,1114]" captionTargetId="figure-16@5.[125,1081,215,1126]" captionTargetPageId="5" captionText="Figs. 3–6. Gondwanoscurus socotrensis sp. nov. 3 – head, male; 4 – frontal suture and facets, male; 5 – terminal lobes of labium, male; 6 – terminal (fourth) palpomere, female." figureDoi="http://doi.org/10.5281/zenodo.5340353" httpUri="https://zenodo.org/record/5340353/files/figure.png" pageId="3" pageNumber="548">Fig. 5</figureCitation>
), with many sensory setae. Cibarium, labrum and epipharynx as in 
<figureCitation id="132BD2EBFFC4FFD1FE6FFA8AF7E3FABB" box="[113,191,1327,1351]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="3" pageNumber="548">Fig. 18</figureCitation>
. Thoracic sclerites and spiraculum as in 
<figureCitation id="132BD2EBFFC4FFD1FC6EFA8AF44AFABB" box="[624,790,1327,1351]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="3" pageNumber="548">Figs. 10 and 11</figureCitation>
.
</paragraph>
<paragraph id="8BAFCE6EFFC4FFD6FE8EFAEAF679FA5E" blockId="3.[113,1089,846,1549]" lastBlockId="4.[151,1127,1258,1538]" lastPageId="4" lastPageNumber="549" pageId="3" pageNumber="548">
Wings broadly lancet-shaped, 
<quantity id="4CE8638BFFC4FFD1FFC5FAEAF573FA9A" box="[475,559,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="548" unit="mm" value="2.9">2.9 mm</quantity>
long (
<typeStatus id="54AB70CCFFC4FFD1FC6BFAEAF586FA9B" box="[629,730,1359,1383]" pageId="3" pageNumber="548" type="holotype">holotype</typeStatus>
), 
<typeStatus id="54AB70CCFFC4FFD1FCF0FAEAF40AFA9B" box="[750,854,1359,1383]" pageId="3" pageNumber="548" type="paratype">paratypes</typeStatus>
2.4–3.0 mm, slightly clouded, rounded at apex, with well developed and enlarged cubital area (
<figureCitation id="132BD2EBFFC4FFD1FD62FACAF4E5FA7B" box="[892,953,1391,1415]" captionStart="Figs" captionStartId="4.[151,190,1161,1180]" captionTargetBox="[207,1078,228,1132]" captionTargetId="figure-12@4.[151,1126,216,1140]" captionTargetPageId="4" captionText="Figs. 1–2. Gondwanoscurus socotrensis sp. nov., wing (1 – male; 2 – female)." figureDoi="http://doi.org/10.5281/zenodo.5340351" httpUri="https://zenodo.org/record/5340351/files/figure.png" pageId="3" pageNumber="548">Fig. 1</figureCitation>
), membrane bare, with conspicuous infuscation patches at apices of longitudinal veins, in area between R 
<subScript id="1794CC2BFFC4FFD1FE9DFA1BF7D7FA31" attach="left" box="[131,139,1470,1485]" fontSize="6" pageId="3" pageNumber="548">1</subScript>
and C, at basis of R 
<subScript id="1794CC2BFFC4FFD1FF6AFA1BF6D0FA31" attach="left" box="[372,396,1470,1485]" fontSize="6" pageId="3" pageNumber="548">2+3</subScript>
, and between CuA 
<subScript id="1794CC2BFFC4FFD1FC7DFA1BF537FA31" attach="left" box="[611,619,1470,1485]" fontSize="6" pageId="3" pageNumber="548">2</subScript>
and hind margin of the wing. Radial fork complete, medial one incomplete (very faint connection of M 
<subScript id="1794CC2BFFC4FFD1FD0CFA7BF446FA11" attach="left" box="[786,794,1502,1517]" fontSize="6" pageId="3" pageNumber="548">1</subScript>
to M 
<subScript id="1794CC2BFFC4FFD1FD4DFA7BF407FA11" attach="left" box="[851,859,1502,1517]" fontSize="6" pageId="3" pageNumber="548">2</subScript>
). Following veins or their parts strengthened: R 
<subScript id="1794CC2BFFC4FFD1FF8BFA5BF6F1F9F1" attach="left" box="[405,429,1534,1549]" fontSize="6" pageId="3" pageNumber="548">2+3</subScript>
and R 
<subScript id="1794CC2BFFC4FFD1FFE8FA5BF6A2F9F1" attach="left" box="[502,510,1534,1549]" fontSize="6" pageId="3" pageNumber="548">2</subScript>
, CuA 
<subScript id="1794CC2BFFC4FFD1FC5EFA5BF514F9F1" attach="left" box="[576,584,1534,1549]" fontSize="6" pageId="3" pageNumber="548">1</subScript>
and CuA 
<subScript id="1794CC2BFFC4FFD1FCAFFA5BF5E5F9F1" attach="left" box="[689,697,1534,1549]" fontSize="6" pageId="3" pageNumber="548">2</subScript>
, A 
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distad. Some parts of basal cell are more sclerotized. Basal costal wing node distinct, Sc uninterrupted, straight. CuA 
<subScript id="1794CC2BFFC3FFD6FA41FB5FF33BFAF5" attach="left" box="[1119,1127,1274,1289]" fontSize="6" pageId="4" pageNumber="549">1</subScript>
basally without connection to M 
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and to CuA 
<subScript id="1794CC2BFFC3FFD6FC8CFABFF5C6FAD5" attach="right" box="[658,666,1306,1321]" fontSize="6" pageId="4" pageNumber="549">2</subScript>
. R 
<subScript id="1794CC2BFFC3FFD6FCA5FABFF59FFAD5" attach="left" box="[699,707,1306,1321]" fontSize="6" pageId="4" pageNumber="549">5</subScript>
extending distally and reaching wing margin slightly below wing apex. Veins r-r, r-m and m-m not developed. Wing 1.8 times as long as wide. Haltere 2.7 times as long as wide (
<figureCitation id="132BD2EBFFC3FFD6FCA4FAEFF456FA9E" box="[698,778,1354,1378]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="4" pageNumber="549">Fig. 19</figureCitation>
). Length ratios of femora, tibiae and first tarsomeres: P 
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= 1.7: 2.0: 1.0; P 
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= 1.9: 2.5: 1.2; P 
<subScript id="1794CC2BFFC3FFD6FD30FADFF46AFA75" attach="left" box="[814,822,1402,1417]" fontSize="6" pageId="4" pageNumber="549">3</subScript>
= 2.0: 2.9: 1.2. Fore claws as in 
<figureCitation id="132BD2EBFFC3FFD6FECFFA2FF67DFA5E" box="[209,289,1418,1442]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="4" pageNumber="549">Fig. 27</figureCitation>
.
</paragraph>
<caption id="DF6F9EE6FFC3FFD6FE89FB2CF46DFB60" ID-DOI="http://doi.org/10.5281/zenodo.5340351" ID-Zenodo-Dep="5340351" box="[151,817,1161,1180]" httpUri="https://zenodo.org/record/5340351/files/figure.png" pageId="4" pageNumber="549" startId="4.[151,190,1161,1180]" targetBox="[207,1078,228,1132]" targetPageId="4">
<paragraph id="8BAFCE6EFFC3FFD6FE89FB2CF46DFB60" blockId="4.[151,817,1161,1180]" box="[151,817,1161,1180]" pageId="4" pageNumber="549">
Figs. 1–2. 
<taxonomicName id="4C10B5EDFFC3FFD6FEEFFB2FF6B7FB60" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[241,491,1162,1180]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC3FFD6FEEFFB2FF6B7FB60" box="[241,491,1162,1180]" italics="true" pageId="4" pageNumber="549">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFC3FFD6FFEFFB2CF569FB60" box="[497,565,1161,1180]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
, wing (1 – male; 2 – female).
</paragraph>
</caption>
<paragraph id="8BAFCE6EFFC3FFDBFEABFA0FF650FDC8" blockId="4.[151,1127,1258,1538]" lastBlockId="9.[113,1089,219,1588]" lastPageId="9" lastPageNumber="554" pageId="4" pageNumber="549">
Basal apodeme of male genitalia straight and narrow in dorsal view (
<figureCitation id="132BD2EBFFC3FFD6FDADFA0FF359FA3E" box="[947,1029,1450,1474]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="4" pageNumber="549">Fig. 28</figureCitation>
), bent in lateral and diagonal view (proximal end rounded – 
<figureCitation id="132BD2EBFFC3FFD6FCCBFA6FF46EFA1E" box="[725,818,1482,1506]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="4" pageNumber="549">Figs. 20</figureCitation>
, 
<figureCitation id="132BD2EBFFC3FFD6FD20FA6FF401FA1E" box="[830,861,1482,1506]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="4" pageNumber="549">29</figureCitation>
). Aedeagus simple with internal structures of characteristic shape (
<figureCitation id="132BD2EBFFC3FFD6FC6EFA4FF592F9FE" box="[624,718,1514,1538]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="4" pageNumber="549">Figs. 20</figureCitation>
, 
<figureCitation id="132BD2EBFFC3FFD6FCC4FA4FF478F9FE" box="[730,804,1514,1538]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="4" pageNumber="549">28, 29</figureCitation>
). Gonocoxites long and thin, almost straight, gonostyli elongate, 1.2 times as long as gonocoxites, gradually tapering to apex, little bent, bifurcate, slightly tapered, digitiform, subequal in length (
<figureCitation id="132BD2EBFFCEFFDBFDA7FF5EF348FEE8" box="[953,1044,251,276]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">Figs. 20</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFA01FF59F7CCFEC8" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">28, 29</figureCitation>
). Epandrium with two irregularly formed fields of caudal insertions of setulae on both sides, hardly connected caudally, and two central circular openings (apertures) (
<figureCitation id="132BD2EBFFCEFFDBFDFEFE9EF7D3FE88" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">Figs. 30, 31</figureCitation>
). Caudal epandrial notch deep. Sclerotized remainders of 10th segment inside of epandrium developed and safely indicated, triangular from dorsal view (
<figureCitation id="132BD2EBFFCEFFDBFD4CFEDEF4F8FE68" box="[850,932,379,404]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">Fig. 30</figureCitation>
). Hypandrium little widened medially (
<figureCitation id="132BD2EBFFCEFFDBFF99FE3EF6BBFE48" box="[391,487,411,436]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">Figs. 20</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFFEBFE39F51DFE48" box="[501,577,412,436]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">28, 29</figureCitation>
). Epiproct linear, fold-shaped, hardly visible, hypoproct longly triangular, rounded, both parts with microtrichia (
<figureCitation id="132BD2EBFFCEFFDBFD70FE1EF498FE28" box="[878,964,443,468]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">Fig. 30</figureCitation>
). Surstylus cylindrical, 3.7 times as long as its basal diameter (with a small protuberance), bent from lateral view, straight from dorsal one (
<figureCitation id="132BD2EBFFCEFFDBFC10FE5EF5CDFDE8" box="[526,657,507,532]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">Figs. 30, 32</figureCitation>
), 23 retinacula subapically, not frazzled (
<figureCitation id="132BD2EBFFCEFFDBFE67FDBEF788FDC8" box="[121,212,539,564]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">Figs. 21</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFEFEFDB9F7A3FDC8" box="[224,255,540,564]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">32</figureCitation>
).
</paragraph>
<caption id="DF6F9EE6FFC2FFD7FE6FFBD4F5D3FB63" ID-DOI="http://doi.org/10.5281/zenodo.5340353" ID-Zenodo-Dep="5340353" httpUri="https://zenodo.org/record/5340353/files/figure.png" pageId="5" pageNumber="550" startId="5.[113,152,1137,1156]" targetBox="[166,1033,232,1114]" targetPageId="5">
<paragraph id="8BAFCE6EFFC2FFD7FE6FFBD4F5D3FB63" blockId="5.[113,1088,1137,1184]" pageId="5" pageNumber="550">
Figs. 3–6. 
<taxonomicName id="4C10B5EDFFC2FFD7FED3FBD6F694FB79" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[205,456,1139,1157]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC2FFD7FED3FBD6F694FB79" box="[205,456,1139,1157]" italics="true" pageId="5" pageNumber="550">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFC2FFD7FFD1FBD7F54FFB79" box="[463,531,1138,1157]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
3 – head, male; 4 – frontal suture and facets, male; 5 – terminal lobes of labium, male; 6 – terminal (fourth) palpomere, female.
</paragraph>
</caption>
<caption id="DF6F9EE6FFC2FFD7FE71F9ACF794F9CA" ID-DOI="http://doi.org/10.5281/zenodo.5340355" ID-Zenodo-Dep="5340355" httpUri="https://zenodo.org/record/5340355/files/figure.png" pageId="5" pageNumber="550" startId="5.[111,150,1545,1564]" targetBox="[113,1085,1237,1528]" targetPageId="5">
<paragraph id="8BAFCE6EFFC2FFD7FE71F9ACF794F9CA" blockId="5.[111,1087,1545,1590]" pageId="5" pageNumber="550">
Figs.7–9. 
<taxonomicName id="4C10B5EDFFC2FFD7FED9F9AFF6E3F9E0" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[199,447,1546,1564]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="550" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC2FFD7FED9F9AFF6E3F9E0" box="[199,447,1546,1564]" italics="true" pageId="5" pageNumber="550">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFC2FFD7FFDDF9ACF55AF9E0" box="[451,518,1545,1564]" pageId="5" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
, female. 7 – genital chamber ventrally; 8 – same, laterally; 9 – same, anteriorly.
</paragraph>
</caption>
<caption id="DF6F9EE6FFC1FFD4FE89FA49F478F9C8" ID-DOI="http://doi.org/10.5281/zenodo.5340357" ID-Zenodo-Dep="5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="6" pageNumber="551" startId="6.[151,190,1516,1535]" targetBox="[299,991,234,1472]" targetPageId="6">
<paragraph id="8BAFCE6EFFC1FFD4FE89FA49F478F9C8" blockId="6.[151,1127,1516,1589]" pageId="6" pageNumber="551">
Figs. 10–14. 
<taxonomicName id="4C10B5EDFFC1FFD4FF1DFA48F6A7FA03" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[259,507,1517,1535]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="551" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC1FFD4FF1DFA48F6A7FA03" box="[259,507,1517,1535]" italics="true" pageId="6" pageNumber="551">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFC1FFD4FFE1FA49F51CFA03" box="[511,576,1516,1535]" pageId="6" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P 
<subScript id="1794CC2BFFC1FFD4FC7DF9B6F535F9E3" attach="left" box="[611,617,1555,1567]" fontSize="5" pageId="6" pageNumber="551">1</subScript>
with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female.
</paragraph>
</caption>
<caption id="DF6F9EE6FFC0FFD5FE6FFA1BF530F9DD" ID-DOI="http://doi.org/10.5281/zenodo.5340359" ID-Zenodo-Dep="5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="7" pageNumber="552" startId="7.[113,152,1470,1489]" targetBox="[163,1038,231,1441]" targetPageId="7">
<paragraph id="8BAFCE6EFFC0FFD5FE6FFA1BF530F9DD" blockId="7.[113,1087,1470,1570]" pageId="7" pageNumber="552">
Figs. 15–23. 
<taxonomicName id="4C10B5EDFFC0FFD5FEC1FA65F684FA2E" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[223,472,1472,1490]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFC0FFD5FEC1FA65F684FA2E" box="[223,472,1472,1490]" italics="true" pageId="7" pageNumber="552">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFC0FFD5FFC2FA1AF543FA2E" box="[476,543,1471,1490]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female.
</paragraph>
</caption>
<caption id="DF6F9EE6FFCFFFDAFE89FA01F54EF9DE" ID-DOI="http://doi.org/10.5281/zenodo.5340363" ID-Zenodo-Dep="5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="8" pageNumber="553" startId="8.[151,190,1444,1463]" targetBox="[243,1032,242,1398]" targetPageId="8">
<paragraph id="8BAFCE6EFFCFFFDAFE89FA01F54EF9DE" blockId="8.[151,1127,1444,1570]" pageId="8" pageNumber="553">
Figs. 24–33. 
<taxonomicName id="4C10B5EDFFCFFFDAFF16FA00F55FFA4B" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[264,515,1445,1463]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFCFFFDAFF16FA00F55FFA4B" box="[264,515,1445,1463]" italics="true" pageId="8" pageNumber="553">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFCFFFDAFC17FA01F511FA4B" box="[521,589,1444,1463]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P 
<subScript id="1794CC2BFFCFFFDAFF43FA43F63FFA0E" attach="left" box="[349,355,1510,1522]" fontSize="5" pageId="8" pageNumber="553">1</subScript>
, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female.
</paragraph>
</caption>
<paragraph id="8BAFCE6EFFCEFFDBFE8EFD98F536FD4F" blockId="9.[113,1089,219,1588]" pageId="9" pageNumber="554">
<emphasis id="B964127CFFCEFFDBFE8EFD98F7BBFDA8" box="[144,231,573,596]" italics="true" pageId="9" pageNumber="554">Female.</emphasis>
Eyes contiguous (
<figureCitation id="132BD2EBFFCEFFDBFFB3FD9EF6A0FDAF" box="[429,508,571,595]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">Fig. 15</figureCitation>
), vertex pyramidal, vertex ratio (width and high) 2.6: 1, frontoclypeal area of insertions of setulae as in male, however, near tentorial pits expanding to small obtuse corners. Sensory filaments (ascoids) of flagellomeres multiple (
<figureCitation id="132BD2EBFFCEFFDBFDFBFDDEF368FD6F" box="[997,1076,635,659]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">Fig. 26</figureCitation>
). Palpomere 4 annulate, scales maculate (
<figureCitation id="132BD2EBFFCEFFDBFC05FD3EF501FD4F" box="[539,605,667,691]" captionStart="Figs" captionStartId="5.[113,152,1137,1156]" captionTargetBox="[166,1033,232,1114]" captionTargetId="figure-16@5.[125,1081,215,1126]" captionTargetPageId="5" captionText="Figs. 3–6. Gondwanoscurus socotrensis sp. nov. 3 – head, male; 4 – frontal suture and facets, male; 5 – terminal lobes of labium, male; 6 – terminal (fourth) palpomere, female." figureDoi="http://doi.org/10.5281/zenodo.5340353" httpUri="https://zenodo.org/record/5340353/files/figure.png" pageId="9" pageNumber="554">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BAFCE6EFFCEFFDBFE8EFD1EF65CFC2F" blockId="9.[113,1089,219,1588]" pageId="9" pageNumber="554">
Wings narrowly lancet-shaped, 3.0 mm long (
<typeStatus id="54AB70CCFFCEFFDBFC9EFD1EF584FD2F" box="[640,728,699,723]" pageId="9" pageNumber="554" type="allotype">allotype</typeStatus>
<quantity id="4CE8638BFFCEFFDBFCC1FD1EF469FD2F" box="[735,821,699,723]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="554" unit="mm" value="2.5">2.5 mm</quantity>
, 
<typeStatus id="54AB70CCFFCEFFDBFD5FFD19F4FBFD28" box="[833,935,700,724]" pageId="9" pageNumber="554" type="paratype">paratypes</typeStatus>
2.2–3.0 mm), slightly clouded, rounded at apex, cubital area not enlarged (
<figureCitation id="132BD2EBFFCEFFDBFD1AFD7EF41AFD08" box="[772,838,731,756]" captionStart="Figs" captionStartId="4.[151,190,1161,1180]" captionTargetBox="[207,1078,228,1132]" captionTargetId="figure-12@4.[151,1126,216,1140]" captionTargetPageId="4" captionText="Figs. 1–2. Gondwanoscurus socotrensis sp. nov., wing (1 – male; 2 – female)." figureDoi="http://doi.org/10.5281/zenodo.5340351" httpUri="https://zenodo.org/record/5340351/files/figure.png" pageId="9" pageNumber="554">Fig. 2</figureCitation>
), membrane bare, with conspicuous infuscation patches at apices of longitudinal veins, in area between R 
<subScript id="1794CC2BFFCEFFDBFDF4FCAEF4AEFCE6" attach="left" box="[1002,1010,779,794]" fontSize="6" pageId="9" pageNumber="554">1</subScript>
and C, at basis of R 
<subScript id="1794CC2BFFCEFFDBFEEBFC8EF651FCC6" attach="left" box="[245,269,811,826]" fontSize="6" pageId="9" pageNumber="554">2+3</subScript>
. Radial fork complete, longly triangular, medial one with almost very imperceptible wide connection of M 
<subScript id="1794CC2BFFCEFFDBFFA8FCEEF6E2FCA6" attach="left" box="[438,446,843,858]" fontSize="6" pageId="9" pageNumber="554">1</subScript>
to M 
<subScript id="1794CC2BFFCEFFDBFFEAFCEEF6A0FCA6" attach="left" box="[500,508,843,858]" fontSize="6" pageId="9" pageNumber="554">2</subScript>
. Following veins or their parts strengthened: Sc, CuA 
<subScript id="1794CC2BFFCEFFDBFA27FCEEF31DFCA6" attach="right" box="[1081,1089,843,858]" fontSize="6" pageId="9" pageNumber="554">1</subScript>
and CuA 
<subScript id="1794CC2BFFCEFFDBFECCFCCEF786FC86" attach="right" box="[210,218,875,890]" fontSize="6" pageId="9" pageNumber="554">2</subScript>
, A 
<subScript id="1794CC2BFFCEFFDBFEE7FCCEF65DFC86" attach="left" box="[249,257,875,890]" fontSize="6" pageId="9" pageNumber="554">1</subScript>
distad. Some parts of basal cell are more sclerotized. Basal costal wing node distinct, Sc uninterrupted, straight. CuA 
<subScript id="1794CC2BFFCEFFDBFC03FC2EF579FC66" attach="left" box="[541,549,907,922]" fontSize="6" pageId="9" pageNumber="554">1</subScript>
basally without connection to M 
<subScript id="1794CC2BFFCEFFDBFD99FC2EF4D3FC66" attach="left" box="[903,911,907,922]" fontSize="6" pageId="9" pageNumber="554">3</subScript>
and to CuA 
<subScript id="1794CC2BFFCEFFDBFA0FFC2EF345FC66" attach="left" box="[1041,1049,907,922]" fontSize="6" pageId="9" pageNumber="554">2</subScript>
. R 
<subScript id="1794CC2BFFCEFFDBFA27FC2EF31DFC66" attach="left" box="[1081,1089,907,922]" fontSize="6" pageId="9" pageNumber="554">5</subScript>
extending distally and reaching wing margin slightly below wing apex. Wing 2.1 times as long as wide.
</paragraph>
<paragraph id="8BAFCE6EFFCEFFDBFE8EFC7EF7C0FB88" blockId="9.[113,1089,219,1588]" pageId="9" pageNumber="554">
Genitalia as figured (
<figureCitation id="132BD2EBFFCEFFDBFF6BFC7EF6BCFC08" box="[373,480,987,1012]" captionStart="Figs" captionStartId="5.[111,150,1545,1564]" captionTargetBox="[113,1085,1237,1528]" captionTargetId="figure-48@5.[113,1085,1237,1528]" captionTargetPageId="5" captionText="Figs.7–9. Gondwanoscurus socotrensis sp. nov., female. 7 – genital chamber ventrally; 8 – same, laterally; 9 – same, anteriorly." figureDoi="http://doi.org/10.5281/zenodo.5340355" httpUri="https://zenodo.org/record/5340355/files/figure.png" pageId="9" pageNumber="554">Figs. 7−9</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFFF2FC79F568FC08" box="[492,564,988,1012]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="9" pageNumber="554">13−14</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFC5EFC79F5D4FC08" box="[576,648,988,1012]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">22−23</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFC8AFC79F5EEFC08" box="[660,690,988,1012]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">33</figureCitation>
). Subgenital plate bilobed (
<figureCitation id="132BD2EBFFCEFFDBFDFEFC7EF360FC08" box="[992,1084,987,1012]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="9" pageNumber="554">Figs. 14</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFE6FFC59F7E6FBE8" box="[113,186,1020,1044]" captionStart="Figs" captionStartId="7.[113,152,1470,1489]" captionTargetBox="[163,1038,231,1441]" captionTargetId="figure-16@7.[138,1062,208,1462]" captionTargetPageId="7" captionText="Figs. 15–23. Gondwanoscurus socotrensis sp. nov. 15 – head, female; 16 – apical antennomeres, male; 17 – maxilla and palpus maxillaris, male; 18 – cibarium, labrum and epipharynx, male; 19 – haltere, male; 20 – aedeagus and gonopod diagonally, male; 21 – end of surstylus ventrally (some retinaculi omitted), male; 22 – subgenital plate (from a slide, pressed), female; 23 – same laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340359" httpUri="https://zenodo.org/record/5340359/files/figure.png" pageId="9" pageNumber="554">22−23</figureCitation>
), with deep caudal concavity, with microtrichia, many scales and setae; complicated sclerotized forms of genital chamber with wartlike structures (
<figureCitation id="132BD2EBFFCEFFDBFD07FBBEF4DFFBC8" box="[793,899,1051,1076]" captionStart="Figs" captionStartId="5.[111,150,1545,1564]" captionTargetBox="[113,1085,1237,1528]" captionTargetId="figure-48@5.[113,1085,1237,1528]" captionTargetPageId="5" captionText="Figs.7–9. Gondwanoscurus socotrensis sp. nov., female. 7 – genital chamber ventrally; 8 – same, laterally; 9 – same, anteriorly." figureDoi="http://doi.org/10.5281/zenodo.5340355" httpUri="https://zenodo.org/record/5340355/files/figure.png" pageId="9" pageNumber="554">Figs. 7−9</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFD91FBB9F4F1FBC8" box="[911,941,1052,1076]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="9" pageNumber="554">14</figureCitation>
). Cerci short, triangular, rounded caudaly, setose (
<figureCitation id="132BD2EBFFCEFFDBFFE5FB9EF50AFBA8" box="[507,598,1083,1108]" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="9" pageNumber="554">Figs. 13</figureCitation>
, 
<figureCitation id="132BD2EBFFCEFFDBFC7CFB99F5DCFBA8" box="[610,640,1084,1108]" captionStart="Figs" captionStartId="8.[151,190,1444,1463]" captionTargetBox="[243,1032,242,1398]" captionTargetId="figure-12@8.[224,1053,220,1413]" captionTargetPageId="8" captionText="Figs. 24–33. Gondwanoscurus socotrensis sp. nov. 24 – basal antennomeres, male; 25 – sensory area of a middle antennomere in detail, male; 26 – apical antennomeres (some ascoids on the empty sockets intentionally not figured), female; 27 – claw of P1, male; 28 – aedeagus and gonopods dorsally, male; 29 – aedeagus and gonopod laterally, male; 30 – epandrium and surstyli (ends with retinacula omitted), male; 31 – same laterally, male; 32 – surstylus laterally, male; 33 – cercus laterally, female." figureDoi="http://doi.org/10.5281/zenodo.5340363" httpUri="https://zenodo.org/record/5340363/files/figure.png" pageId="9" pageNumber="554">33</figureCitation>
), connected by wrinkled membrane (
<figureCitation id="132BD2EBFFCEFFDBFA0BFB9EF7D3FB88" captionStart="Figs" captionStartId="6.[151,190,1516,1535]" captionTargetBox="[299,991,234,1472]" captionTargetId="figure-12@6.[243,1033,219,1493]" captionTargetPageId="6" captionText="Figs. 10–14.Gondwanoscurus socotrensis sp. nov.10 – lateral view of thoracic sclerites, male; 11 – thoracal spiracle, male; 12 – connection of tibia and first tarsomere of P1 with a botka laterally, male; 13 – ovipositor ventrally (from a slide), female; 14 – subgenital plate and genital chamber ventrally, female." figureDoi="http://doi.org/10.5281/zenodo.5340357" httpUri="https://zenodo.org/record/5340357/files/figure.png" pageId="9" pageNumber="554">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A9DE5FFCEFFDBFE6FFBDEF7A8FA4F" pageId="9" pageNumber="554" type="diagnosis">
<paragraph id="8BAFCE6EFFCEFFDBFE6FFBDEF7A8FA4F" blockId="9.[113,1089,219,1588]" pageId="9" pageNumber="554">
<emphasis id="B964127CFFCEFFDBFE6FFBDEF632FB6F" bold="true" box="[113,366,1147,1171]" pageId="9" pageNumber="554">Differential diagnosis.</emphasis>
<taxonomicName id="4C10B5EDFFCEFFDBFF69FBD8F5EEFB68" authority="Ježek &amp; Tkoč, 2012" authorityName="Ježek &amp; Tkoč" authorityYear="2012" box="[375,690,1149,1172]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="socotrensis" status="sp. nov.">
<emphasis id="B964127CFFCEFFDBFF69FBD8F5EEFB68" box="[375,690,1149,1172]" italics="true" pageId="9" pageNumber="554">Gondwanoscurus socotrensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257AF07FFCEFFDBFCA5FBD9F44CFB6F" box="[699,784,1148,1171]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
(J) has vertex ratio 2.4: 1 (width and high); the end of R 
<subScript id="1794CC2BFFCEFFDBFFAEFB0EF6E4FB46" attach="left" box="[432,440,1195,1210]" fontSize="6" pageId="9" pageNumber="554">4</subScript>
is above rounded wing apex; scales of palp segments, wings, legs, halteres and surstyli are maculated; hypandrium widened medially; gonostyli elongate, gradually tapering to apex, bifurcate apically, rami digitiform, subequal in length; aedeagus simple, with one short pointed part inside of a sheath. 
<taxonomicName id="4C10B5EDFFCEFFDBFCB3FB58F409FAE8" authorityName="Jezek" authorityYear="2002" box="[685,853,1277,1300]" class="Insecta" family="Psychodidae" genus="Gondwanoscurus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="malaysiensis">
<emphasis id="B964127CFFCEFFDBFCB3FB58F409FAE8" box="[685,853,1277,1300]" italics="true" pageId="9" pageNumber="554">G. malaysiensis</emphasis>
</taxonomicName>
differs by vertex ratio 1.8: 1; the end of R 
<subScript id="1794CC2BFFCEFFDBFF23FA8EF619FAC6" attach="left" box="[317,325,1323,1338]" fontSize="6" pageId="9" pageNumber="554">4</subScript>
is in pointed apex of wing; scales of palpomeres, wings, legs and surstyli are without maculation; hypandrium narrow; gonostyli with broad semiglobular basal portion, cylindrical medial portion, and abruptly narrowed at the beginning very narrow terminal portion; aedeagal complex composed from three free long pointed parts, trowel-shaped in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C30A9DE5FFCEFFDBFE6FFA19F655FA0F" pageId="9" pageNumber="554" type="etymology">
<paragraph id="8BAFCE6EFFCEFFDBFE6FFA19F655FA0F" blockId="9.[113,1089,219,1588]" pageId="9" pageNumber="554">
<emphasis id="B964127CFFCEFFDBFE6FFA19F7AEFA28" bold="true" box="[113,242,1468,1492]" pageId="9" pageNumber="554">Etymology.</emphasis>
The new species name (adjective) is based on the name of the island where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="C30A9DE5FFCEFFDBFE6FFA5EF342F9EF" box="[113,1054,1531,1556]" pageId="9" pageNumber="554" type="biology_ecology">
<paragraph id="8BAFCE6EFFCEFFDBFE6FFA5EF342F9EF" blockId="9.[113,1089,219,1588]" box="[113,1054,1531,1556]" pageId="9" pageNumber="554">
<emphasis id="B964127CFFCEFFDBFE6FFA5EF546F9EF" bold="true" box="[113,538,1531,1556]" pageId="9" pageNumber="554">Biology and collecting circumstances.</emphasis>
Unknown. The individuals were caught at light.
</paragraph>
</subSubSection>
<subSubSection id="C30A9DE5FFCEFFDBFE6FF9BEF6F7F9CF" box="[113,427,1563,1588]" pageId="9" pageNumber="554" type="distribution">
<paragraph id="8BAFCE6EFFCEFFDBFE6FF9BEF6F7F9CF" blockId="9.[113,1089,219,1588]" box="[113,427,1563,1588]" pageId="9" pageNumber="554">
<emphasis id="B964127CFFCEFFDBFE6FF9BEF658F9CF" bold="true" box="[113,260,1563,1587]" pageId="9" pageNumber="554">Distribution.</emphasis>
<collectingCountry id="F3078EFEFFCEFFDBFF14F9B9F6FBF9CF" box="[266,423,1563,1588]" name="Socotra Island" pageId="9" pageNumber="554">Socotra Island</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>