532 lines
76 KiB
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532 lines
76 KiB
XML
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<mods:title id="E538CF6177991A80C3025F3F3153EA8F">Filling gaps in the knowledge of Goniodorididae taxa (Mollusca, Gastropoda, Nudibranchia) with description of seven new species</mods:title>
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<mods:namePart id="91BAD96BC00F3A8B12212438167B130F">Paz-Sedano, Sofia</mods:namePart>
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<mods:namePart id="E615E14734BEF6FBDEF4D6DA85166565">Cobb, Gary</mods:namePart>
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<mods:affiliation id="980634E95FD36859F41CF219A9539445">Nudibranch Central FB Group, Buderim, Queensland, Australia</mods:affiliation>
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<mods:namePart id="FC0FFF63EBC2C67F6A5B3EC0F83B375D">Gosliner, Terrence M.</mods:namePart>
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<mods:namePart id="70C59A08366E9ED433E6106112636FB4">Pola, Marta</mods:namePart>
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<mods:affiliation id="9E51C39408D298D4956398909D5D13C4">Department of Biology, Faculty of Sciences, Autonomous University of Madrid, Madrid, Spain & Biodiversity and Global Change Research Center (CIBC-UAM), Madrid, Spain</mods:affiliation>
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<mods:title id="BAE9BA2B8FAC67863EAAD6163BBF09FA">Zootaxa</mods:title>
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<taxonomicName id="4C1C4D76FFC5EE1F00F94D86FE10FF6C" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[151,469,151,178]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="533" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
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<emphasis id="B968EAE7FFC5EE1F00F94D86FE10FF6C" bold="true" box="[151,469,151,178]" italics="true" pageId="10" pageNumber="533">Ceratodoris trypomandyas</emphasis>
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<taxonomicNameLabel id="A25B579CFFC5EE1F01B34D86FDF9FF6F" box="[477,572,151,177]" pageId="10" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C306657EFFC5EE1F00F94DAAFBF6FF24" pageId="10" pageNumber="533" type="description">
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<paragraph id="8BA336F5FFC5EE1F00F94DAAFE7CFF08" blockId="10.[151,1075,151,250]" box="[151,441,187,214]" pageId="10" pageNumber="533">
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(
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<figureCitation id="13272A70FFC5EE1F00F14DAAFEE5FF0B" box="[159,288,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1930]" captionTargetBox="[271,1316,364,1880]" captionTargetId="figure-42@2.[271,1316,364,1880]" captionTargetPageId="2" captionText="FIGURE 1. Living animals. A. Trapania franae sp. nov. (QM MO 86031). B‒C. Bermudella lahainensis sp. nov., B. CASIZ 180297 and C. CASIZ 192539; D. Bermudella mica. E. Ceratodoris plebeia (MNCN 15.05/94478). F‒G. Ceratodoris trypomandyas sp. nov., F. QM MO 86033 and G. AM C. 270712. H. Murphydoris polkadotsa sp. nov. (QM MO 86038). (Photos A, F–H by G. Cobb., B‒C by C. Pittman, D by P. Pérez, E by S. T. Huang)" figureDoi="http://doi.org/10.5281/zenodo.11064314" httpUri="https://zenodo.org/record/11064314/files/figure.png" pageId="10" pageNumber="533">Figs. 1F‒G</figureCitation>
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,
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<figureCitation id="13272A70FFC5EE1F01434DAAFE9AFF0B" box="[301,351,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[365,1223,192,1766]" captionTargetId="figure-19@3.[354,1234,181,1776]" captionTargetPageId="3" captionText="FIGURE 2. Buccal bulb and reproductive system. A‒B. Trapania franae sp. nov. (QM MO 86031), A. Buccal bulb. B. Reproductive system. C‒D. Bermudella lahainensis sp. nov. (CASIZ 199240). C. Buccal bulb. D. Reproductive system. E‒F. Bermudella mica (MNCN 15.05/94474). E. Buccal bulb. F. Reproductive system. G–H. Bermudella plebeia (MNCN 15.05/94477). G. Buccal bulb. H. Reproductive system. I‒J. Ceratodoris trypomandyas sp. nov. (QM MO 86034). I. Buccal bulb. J. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064316" httpUri="https://zenodo.org/record/11064316/files/figure.png" pageId="10" pageNumber="533">2I‒J</figureCitation>
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,
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<figureCitation id="13272A70FFC5EE1F01054DAAFE74FF08" box="[363,433,187,214]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">5A‒D</figureCitation>
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)
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Zoobank:
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urn:lsid:zoobank.org:act:
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<taxonomicName id="4C1C4D76FFC5EE1F00F94C35FEDBFEE2" ID-CoL="7SDR8" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[151,286,292,316]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="533" phylum="Mollusca" rank="species" species="plana">Okenia plana</taxonomicName>
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—
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<bibRefCitation id="EF8D4B04FFC5EE1F01594C35FE1FFEE2" author="Rudman, W. B." box="[311,474,292,316]" pageId="10" pageNumber="533" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman (2004)</bibRefCitation>
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: 49
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</treatmentCitation>
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; Figs. 20C, 23, 24B, 27D, 29F, 30A.
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<paragraph id="8BA336F5FFC5EE1F00F94C53FDA9FE84" blockId="10.[151,886,292,377]" box="[151,620,322,346]" pageId="10" pageNumber="533">
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<treatmentCitationGroup id="AB0C11DBFFC5EE1F00F94C53FDA9FE84" box="[151,620,322,346]" pageId="10" pageNumber="533">
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<taxonomicName id="4C1C4D76FFC5EE1F00F94C53FEE4FE84" ID-CoL="7SDR8" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[151,289,322,346]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="533" phylum="Mollusca" rank="species" species="plana">
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<emphasis id="B968EAE7FFC5EE1F00F94C53FEE4FE84" box="[151,289,322,346]" italics="true" pageId="10" pageNumber="533">Okenia plana</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0ABD10E4FFC5EE1F01544C53FDADFE84" author="Debelius, H. & Kuiter, R. H." box="[314,616,322,346]" page="24" pageId="10" pageNumber="533" year="2007">
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<bibRefCitation id="EF8D4B04FFC5EE1F01544C53FD86FE84" author="Debelius, H. & Kuiter, R. H." box="[314,579,322,346]" pageId="10" pageNumber="533" refId="ref15860" refString="Debelius, H. & Kuiter, R. H. (2007) Nudibranchs of the World. IKAN-Unterwasserarchiv, Frankfurt, 360 pp." type="book" year="2007">Debelius & Kuiter (2007)</bibRefCitation>
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: 24
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</treatmentCitation>
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.
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<paragraph id="8BA336F5FFC5EE1F00F94C70FD99FEA7" blockId="10.[151,886,292,377]" box="[151,604,353,377]" pageId="10" pageNumber="533">
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<taxonomicName id="4C1C4D76FFC5EE1F00F94C70FE85FEA7" ID-CoL="BG9C8" box="[151,320,353,377]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="533" phylum="Mollusca" rank="species" species="undetermined">
|
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<emphasis id="B968EAE7FFC5EE1F00F94C70FED4FEA7" box="[151,273,353,377]" italics="true" pageId="10" pageNumber="533">Ceratodoris</emphasis>
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spD
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</taxonomicName>
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||
—
|
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<bibRefCitation id="EF8D4B04FFC5EE1F01344C70FD92FEA7" author="Paz-Sedano, S. & Moles, J. & Smirnoff, D. & Gosliner, T. M. & Pola, M." box="[346,599,353,377]" pageId="10" pageNumber="533" pagination="107990" refId="ref17463" refString="Paz-Sedano, S., Moles, J., Smirnoff, D., Gosliner, T. M. & Pola, M. (2024) A combined phylogenetic strategy illuminates the evolution of Goniodorididae nudibranchs (Mollusca, Gastropoda, Heterobranchia). Molecular Phylogenetics and Evolution, 192, 107990. https: // doi. org / 10.1016 / j. ympev. 2023.107990" type="journal article" year="2024">
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Paz-Sedano
|
||
<emphasis id="B968EAE7FFC5EE1F01B64C73FDCEFEA7" box="[472,523,353,377]" italics="true" pageId="10" pageNumber="533">et al.</emphasis>
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(2024)
|
||
</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<emphasis id="B968EAE7FFC5EE1F00F94CB6FEFAFE1F" bold="true" box="[151,319,423,449]" pageId="10" pageNumber="533">Type material</emphasis>
|
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.
|
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<materialsCitation id="3B743CA8FFC5EE1F01224CB6FD09FE3B" collectingDate="2020-11-03" collectionCode="QM" collectorName="La Balsa Park & G. Cobb" country="Australia" elevation="2" location="La Balsa Park" municipality="Mooloolah River" pageId="10" pageNumber="533" specimenCode="MO 86033" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<emphasis id="B968EAE7FFC5EE1F01224CB6FE7CFE1F" bold="true" box="[332,441,423,449]" pageId="10" pageNumber="533">
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<typeStatus id="54A78857FFC5EE1F01224CB6FE7CFE1F" box="[332,441,423,449]" pageId="10" pageNumber="533" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<collectionCode id="ED0DAE30FFC5EE1F01AB4CB6FE37FE1F" box="[453,498,423,449]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="10" pageNumber="533" type="Museum">QM</collectionCode>
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,
|
||
<collectorName id="26E95323FFC5EE1F02104CB9FCD9FE1F" box="[638,796,423,450]" pageId="10" pageNumber="533">
|
||
<location id="8EC3602EFFC5EE1F02104CB9FCD9FE1F" LSID="urn:lsid:plazi:treatment:03B587E3FFC5EE1900F94D86FEC3FB87:8EC3602EFFC5EE1F02104CB9FCD9FE1F" box="[638,796,423,450]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="10" pageNumber="533" stateProvince="Queensland">La Balsa Park</location>
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC7AC8FFFC5EE1F03484CB6FC2DFE1F" box="[806,1000,423,449]" pageId="10" pageNumber="533">Mooloolah River</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49D8F817FFC5EE1F039C4CB6FBBCFE1F" box="[1010,1145,423,449]" country="Australia" name="Queensland" pageId="10" pageNumber="533">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B7665FFC5EE1F04ED4CB6FB28FE1F" box="[1155,1261,423,449]" name="Australia" pageId="10" pageNumber="533">Australia</collectingCountry>
|
||
,
|
||
<elevation id="0031D1C6FFC5EE1F04984CB6FA5DFE1F" box="[1270,1432,423,450]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">
|
||
<quantity id="4CE49B10FFC5EE1F04984CB6FA94FE1C" box="[1270,1361,423,450]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity>
|
||
depth
|
||
</elevation>
|
||
,
|
||
<date id="FFA21035FFC5EE1F00F94CDAFEB3FE3B" box="[151,374,459,485]" pageId="10" pageNumber="533" value="2020-11-03">
|
||
<collectingDate id="EFE6E9DDFFC5EE1F00F94CDAFEB3FE3B" box="[151,374,459,485]" pageId="10" pageNumber="533" value="2020-11-03">03 November 2020</collectingDate>
|
||
</date>
|
||
, col. by
|
||
<collectorName id="26E95323FFC5EE1F01B44CDAFDF8FE3B" box="[474,573,459,485]" pageId="10" pageNumber="533">G. Cobb</collectorName>
|
||
, 96% EtOH
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B743CA8FFC5EE1F02B74CDAFEF6FDF3" collectingDate="2020-11-03" collectionCode="QM, SEM" collectorName="La Balsa Park & G. Cobb" country="Australia" elevation="2" location="La Balsa Park" municipality="Mooloolah River" pageId="10" pageNumber="533" specimenCode="MO 86034" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<emphasis id="B968EAE7FFC5EE1F02B74CDAFC94FE3B" bold="true" box="[729,849,459,485]" pageId="10" pageNumber="533">
|
||
<typeStatus id="54A78857FFC5EE1F02B74CDAFC94FE3B" box="[729,849,459,485]" pageId="10" pageNumber="533" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="ED0DAE30FFC5EE1F030F4CDAFC4BFE3B" box="[865,910,459,485]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="10" pageNumber="533" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBBA9E8EFFC5EE1F03F94CDAFBD3FE3B" box="[919,1046,459,485]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="10" pageNumber="533" type="Museum">MO 86034</specimenCode>
|
||
,
|
||
<collectorName id="26E95323FFC5EE1F044D4CDDFB03FE3B" box="[1059,1222,459,486]" pageId="10" pageNumber="533">
|
||
<location id="8EC3602EFFC5EE1F044D4CDDFB03FE3B" LSID="urn:lsid:plazi:treatment:03B587E3FFC5EE1900F94D86FEC3FB87:8EC3602EFFC5EE1F044D4CDDFB03FE3B" box="[1059,1222,459,486]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="10" pageNumber="533" stateProvince="Queensland">La Balsa Park</location>
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC7AC8FFFC5EE1F04BD4CDAFA5DFE3B" box="[1235,1432,459,485]" pageId="10" pageNumber="533">Mooloolah River</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49D8F817FFC5EE1F00F94CFEFEDBFDD7" box="[151,286,495,521]" country="Australia" name="Queensland" pageId="10" pageNumber="533">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B7665FFC5EE1F01454CFEFE50FDD7" box="[299,405,495,521]" name="Australia" pageId="10" pageNumber="533">Australia</collectingCountry>
|
||
,
|
||
<elevation id="0031D1C6FFC5EE1F01CE4CFEFD82FDD7" box="[416,583,495,522]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">
|
||
<quantity id="4CE49B10FFC5EE1F01CE4CFEFE3BFDD4" box="[416,510,495,522]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity>
|
||
depth
|
||
</elevation>
|
||
,
|
||
<date id="FFA21035FFC5EE1F023D4CFEFCF5FDD7" box="[595,816,495,521]" pageId="10" pageNumber="533" value="2020-11-03">
|
||
<collectingDate id="EFE6E9DDFFC5EE1F023D4CFEFCF5FDD7" box="[595,816,495,521]" pageId="10" pageNumber="533" value="2020-11-03">03 November 2020</collectingDate>
|
||
</date>
|
||
, col. by
|
||
<collectorName id="26E95323FFC5EE1F03FC4CFEFC30FDD7" box="[914,1013,495,521]" pageId="10" pageNumber="533">G. Cobb</collectorName>
|
||
, 96% EtOH, dissected (
|
||
<collectionCode id="ED0DAE30FFC5EE1F056A4CFEFA86FDD7" box="[1284,1347,495,521]" pageId="10" pageNumber="533">SEM</collectionCode>
|
||
: radula, labial cuticle)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B743CA8FFC5EE1F01524F02FDD2FD8F" collectingDate="2020-11-03" collectionCode="AM" collectorName="La Balsa Park & G. Cobb" country="Australia" elevation="2" location="La Balsa Park" municipality="Mooloolah River" pageId="10" pageNumber="533" specimenCode="C270711" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<collectionCode id="ED0DAE30FFC5EE1F01524F02FEACFDF3" box="[316,361,531,557]" country="Australia" name="Australian Museum" pageId="10" pageNumber="533" type="Museum">AM</collectionCode>
|
||
<specimenCode id="DBBA9E8EFFC5EE1F01014F02FE1CFDF3" box="[367,473,531,557]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="10" pageNumber="533" type="Museum">C270711</specimenCode>
|
||
,
|
||
<collectorName id="26E95323FFC5EE1F018D4F05FD45FDF3" box="[483,640,531,558]" pageId="10" pageNumber="533">
|
||
<location id="8EC3602EFFC5EE1F018D4F05FD45FDF3" LSID="urn:lsid:plazi:treatment:03B587E3FFC5EE1900F94D86FEC3FB87:8EC3602EFFC5EE1F018D4F05FD45FDF3" box="[483,640,531,558]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="10" pageNumber="533" stateProvince="Queensland">La Balsa Park</location>
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC7AC8FFFC5EE1F02E44F02FC8EFDF3" box="[650,843,531,557]" pageId="10" pageNumber="533">Mooloolah River</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49D8F817FFC5EE1F033B4F02FC19FDF3" box="[853,988,531,557]" country="Australia" name="Queensland" pageId="10" pageNumber="533">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B7665FFC5EE1F03884F02FB95FDF3" box="[998,1104,531,557]" name="Australia" pageId="10" pageNumber="533">Australia</collectingCountry>
|
||
,
|
||
<elevation id="0031D1C6FFC5EE1F04374F02FB3FFDF3" box="[1113,1274,531,558]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">
|
||
<quantity id="4CE49B10FFC5EE1F04374F02FB71FDF0" box="[1113,1204,531,558]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity>
|
||
depth
|
||
</elevation>
|
||
,
|
||
<date id="FFA21035FFC5EE1F056A4F02FF17FD8F" pageId="10" pageNumber="533" value="2020-11-03">
|
||
<collectingDate id="EFE6E9DDFFC5EE1F056A4F02FF17FD8F" pageId="10" pageNumber="533" value="2020-11-03">03 November 2020</collectingDate>
|
||
</date>
|
||
, col. by
|
||
<collectorName id="26E95323FFC5EE1F01404F26FE4BFD8F" box="[302,398,567,593]" pageId="10" pageNumber="533">G. Cobb</collectorName>
|
||
, 96% EtOH
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B743CA8FFC5EE1F024F4F26FB01FDAB" collectingDate="2020-11-03" collectionCode="AM, SEM" collectorName="La Balsa Park & G. Cobb" country="Australia" elevation="2" location="La Balsa Park" municipality="Mooloolah River" pageId="10" pageNumber="533" specimenCode="C270712" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<collectionCode id="ED0DAE30FFC5EE1F024F4F26FD8BFD8F" box="[545,590,567,593]" country="Australia" name="Australian Museum" pageId="10" pageNumber="533" type="Museum">AM</collectionCode>
|
||
<specimenCode id="DBBA9E8EFFC5EE1F023A4F26FD7BFD8F" box="[596,702,567,593]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="10" pageNumber="533" type="Museum">C270712</specimenCode>
|
||
,
|
||
<collectorName id="26E95323FFC5EE1F02A64F29FCA3FD8F" box="[712,870,567,594]" pageId="10" pageNumber="533">
|
||
<location id="8EC3602EFFC5EE1F02A64F29FCA3FD8F" LSID="urn:lsid:plazi:treatment:03B587E3FFC5EE1900F94D86FEC3FB87:8EC3602EFFC5EE1F02A64F29FCA3FD8F" box="[712,870,567,594]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="10" pageNumber="533" stateProvince="Queensland">La Balsa Park</location>
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6BC7AC8FFFC5EE1F031E4F26FBF7FD8F" box="[880,1074,567,593]" pageId="10" pageNumber="533">Mooloolah River</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49D8F817FFC5EE1F04524F26FB06FD8F" box="[1084,1219,567,593]" country="Australia" name="Queensland" pageId="10" pageNumber="533">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B7665FFC5EE1F04A04F26FAFDFD8F" box="[1230,1336,567,593]" name="Australia" pageId="10" pageNumber="533">Australia</collectingCountry>
|
||
,
|
||
<elevation id="0031D1C6FFC5EE1F052F4F26FF1DFDAB" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">
|
||
<quantity id="4CE49B10FFC5EE1F052F4F26FA59FD8C" box="[1345,1436,567,594]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="10" pageNumber="533" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity>
|
||
depth
|
||
</elevation>
|
||
,
|
||
<date id="FFA21035FFC5EE1F008D4F4AFE7AFDAB" box="[227,447,603,629]" pageId="10" pageNumber="533" value="2020-11-03">
|
||
<collectingDate id="EFE6E9DDFFC5EE1F008D4F4AFE7AFDAB" box="[227,447,603,629]" pageId="10" pageNumber="533" value="2020-11-03">03 November 2020</collectingDate>
|
||
</date>
|
||
, col. by
|
||
<collectorName id="26E95323FFC5EE1F02734F4AFDBBFDAB" box="[541,638,603,629]" pageId="10" pageNumber="533">G. Cobb</collectorName>
|
||
, 96% EtOH, dissected (
|
||
<collectionCode id="ED0DAE30FFC5EE1F03E74F4AFC0DFDAB" box="[905,968,603,629]" pageId="10" pageNumber="533">SEM</collectionCode>
|
||
: radula, labial cuticle)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC5EE1F00A94F6EFB98FC64" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A94F6EFE02FD47" bold="true" box="[199,455,639,665]" pageId="10" pageNumber="533">External morphology</emphasis>
|
||
(
|
||
<figureCitation id="13272A70FFC5EE1F01B84F6EFD9CFD47" box="[470,601,639,665]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1930]" captionTargetBox="[271,1316,364,1880]" captionTargetId="figure-42@2.[271,1316,364,1880]" captionTargetPageId="2" captionText="FIGURE 1. Living animals. A. Trapania franae sp. nov. (QM MO 86031). B‒C. Bermudella lahainensis sp. nov., B. CASIZ 180297 and C. CASIZ 192539; D. Bermudella mica. E. Ceratodoris plebeia (MNCN 15.05/94478). F‒G. Ceratodoris trypomandyas sp. nov., F. QM MO 86033 and G. AM C. 270712. H. Murphydoris polkadotsa sp. nov. (QM MO 86038). (Photos A, F–H by G. Cobb., B‒C by C. Pittman, D by P. Pérez, E by S. T. Huang)" figureDoi="http://doi.org/10.5281/zenodo.11064314" httpUri="https://zenodo.org/record/11064314/files/figure.png" pageId="10" pageNumber="533">Figs. 1F‒G</figureCitation>
|
||
). Preserved specimens
|
||
<quantity id="4CE49B10FFC5EE1F03304F6EFC7EFD44" box="[862,955,639,666]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="10" pageNumber="533" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
|
||
in length, live
|
||
<quantity id="4CE49B10FFC5EE1F040D4F6EFB06FD44" box="[1123,1219,639,666]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="10" pageNumber="533" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
|
||
. Body rounded and very flat, with developed notal border. Dorsal part of body bearing thin, relatively small, and elongated papillae. Five papillae border each side of dorsum. One in front of rhinophores, three between rhinophores and gill, and one behind gill. One additional papilla located in middorsal part of body. All papillae have same shape and size. Rhinophores nonretractile, elongated, slender, bearing 9–12 lamellae each oriented to posterior part. Rhinophoral sheaths absent. Gill composed of nine small, plumose branches forming a semicircle around anus. One middorsal branch larger, four smaller branches on each side. Posteriormost gill branches tiny. One oral tentacle on each side of mouth, semicircular, almost fused with body. Reproductive opening located on right lateral side, in first third of body. Mantle covered by very dense network of spicules, forming a rhomboid pattern.
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC5EE1F00A94ED2FB00FBF8" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A94ED2FEA8FC03" bold="true" box="[199,365,963,989]" pageId="10" pageNumber="533">Color pattern</emphasis>
|
||
(
|
||
<figureCitation id="13272A70FFC5EE1F01134ED2FDC4FC03" box="[381,513,963,989]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1930]" captionTargetBox="[271,1316,364,1880]" captionTargetId="figure-42@2.[271,1316,364,1880]" captionTargetPageId="2" captionText="FIGURE 1. Living animals. A. Trapania franae sp. nov. (QM MO 86031). B‒C. Bermudella lahainensis sp. nov., B. CASIZ 180297 and C. CASIZ 192539; D. Bermudella mica. E. Ceratodoris plebeia (MNCN 15.05/94478). F‒G. Ceratodoris trypomandyas sp. nov., F. QM MO 86033 and G. AM C. 270712. H. Murphydoris polkadotsa sp. nov. (QM MO 86038). (Photos A, F–H by G. Cobb., B‒C by C. Pittman, D by P. Pérez, E by S. T. Huang)" figureDoi="http://doi.org/10.5281/zenodo.11064314" httpUri="https://zenodo.org/record/11064314/files/figure.png" pageId="10" pageNumber="533">Figs. 1F‒G</figureCitation>
|
||
). Background of body translucent. Light brown patches covering body, delimited with a network of hyaline white lines. Rhinophores same color as body. Papillae translucent, may have some brown patches. Gill branches translucent. A dense network of spicules visible, providing a bright tone.
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC5EE1F00A9493EFE7CFA6C" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A9493EFE53FB97" bold="true" box="[199,406,1071,1097]" pageId="10" pageNumber="533">Foregut anatomy</emphasis>
|
||
(
|
||
<figureCitation id="13272A70FFC5EE1F01C9493EFDC5FB97" box="[423,512,1071,1097]" captionStart="FIGURE 2" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[365,1223,192,1766]" captionTargetId="figure-19@3.[354,1234,181,1776]" captionTargetPageId="3" captionText="FIGURE 2. Buccal bulb and reproductive system. A‒B. Trapania franae sp. nov. (QM MO 86031), A. Buccal bulb. B. Reproductive system. C‒D. Bermudella lahainensis sp. nov. (CASIZ 199240). C. Buccal bulb. D. Reproductive system. E‒F. Bermudella mica (MNCN 15.05/94474). E. Buccal bulb. F. Reproductive system. G–H. Bermudella plebeia (MNCN 15.05/94477). G. Buccal bulb. H. Reproductive system. I‒J. Ceratodoris trypomandyas sp. nov. (QM MO 86034). I. Buccal bulb. J. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064316" httpUri="https://zenodo.org/record/11064316/files/figure.png" pageId="10" pageNumber="533">Figs. 2I</figureCitation>
|
||
,
|
||
<figureCitation id="13272A70FFC5EE1F0263493EFD91FB94" box="[525,596,1071,1098]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">5A‒D</figureCitation>
|
||
). Buccal bulb thick, muscular (
|
||
<figureCitation id="13272A70FFC5EE1F03D6493EFBCDFB97" box="[952,1032,1071,1097]" captionStart="FIGURE 2" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[365,1223,192,1766]" captionTargetId="figure-19@3.[354,1234,181,1776]" captionTargetPageId="3" captionText="FIGURE 2. Buccal bulb and reproductive system. A‒B. Trapania franae sp. nov. (QM MO 86031), A. Buccal bulb. B. Reproductive system. C‒D. Bermudella lahainensis sp. nov. (CASIZ 199240). C. Buccal bulb. D. Reproductive system. E‒F. Bermudella mica (MNCN 15.05/94474). E. Buccal bulb. F. Reproductive system. G–H. Bermudella plebeia (MNCN 15.05/94477). G. Buccal bulb. H. Reproductive system. I‒J. Ceratodoris trypomandyas sp. nov. (QM MO 86034). I. Buccal bulb. J. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064316" httpUri="https://zenodo.org/record/11064316/files/figure.png" pageId="10" pageNumber="533">Fig. 2I</figureCitation>
|
||
). Large and elongated buccal pump located dorsally and expands posteriorly. Rounded radular sac located ventrally. Esophagus begins behind buccal pump. Very small, oval salivary glands located at junction of esophagus with buccal bulb. Nervous system surrounds this junction, covering salivary glands. Esophagus continues posteriorly and inserts into digestive-hermaphroditic gland. Thin, weak labial cuticle surrounds lips and expands within buccal pump. Labial cuticle with honey-comb elements (
|
||
<figureCitation id="13272A70FFC5EE1F016149F2FEAAFB20" box="[271,367,1251,1278]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Fig. 5A</figureCitation>
|
||
). Radular formula 23‒24 × 1.1.0.1.1. Inner lateral tooth large, with single cusp, and wide rectangular base (
|
||
<figureCitation id="13272A70FFC5EE1F010C4816FE2CFAFC" box="[354,489,1287,1314]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Figs. 5B–C</figureCitation>
|
||
). Cusp large and pointed with smooth masticatory margin (
|
||
<figureCitation id="13272A70FFC5EE1F04F94816FAD6FAFC" box="[1175,1299,1287,1314]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Fig. 5B–C</figureCitation>
|
||
). Base ends in somewhat prominent wing, with rounded external edge. Outer lateral tooth much smaller (
|
||
<figureCitation id="13272A70FFC5EE1F04C0483AFAF6FA98" box="[1198,1331,1323,1350]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Figs. 5B–C</figureCitation>
|
||
). Base of outer tooth rectangular, top with large and pointed cusps. Cusps located externally larger than inner cusps, form a circle where inner cusps remain inside (
|
||
<figureCitation id="13272A70FFC5EE1F02384862FD19FA50" box="[598,732,1395,1422]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Figs. 5B–D</figureCitation>
|
||
). Outer teeth may have four to six cusps. External cusps larger than inner ones (
|
||
<figureCitation id="13272A70FFC5EE1F013C4886FE6EFA6C" box="[338,427,1431,1458]" captionStart="FIGURE 5" captionStartId="11.[151,250,1727,1751]" captionTargetBox="[232,1355,181,1702]" captionTargetId="figure-19@11.[232,1355,181,1702]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis. A‒D. Ceratodoris trypomandyas sp. nov. A. Detail of labial cuticle elements (QM MO 86034). B. Frontal view of radula (QM MO 86034). C. Detail of inner and outer lateral teeth (QM MO 86034). D. Detail of outer lateral teeth (AM C. 270712). E‒H. Murphydoris polkadotsa sp. nov. (QM MO 86038). E. Frontal view of radula. F. Detail of inner lateral teeth. G. Detail of outer lateral teeth. H. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="10" pageNumber="533">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC5EE1F00A948AAFAECF92B" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A948AAFE06FA0B" bold="true" box="[199,451,1467,1493]" pageId="10" pageNumber="533">Reproductive system</emphasis>
|
||
(
|
||
<figureCitation id="13272A70FFC5EE1F01BA48AAFDE2FA0B" box="[468,551,1467,1493]" captionStart="FIGURE 2" captionStartId="3.[151,250,1800,1824]" captionTargetBox="[365,1223,192,1766]" captionTargetId="figure-19@3.[354,1234,181,1776]" captionTargetPageId="3" captionText="FIGURE 2. Buccal bulb and reproductive system. A‒B. Trapania franae sp. nov. (QM MO 86031), A. Buccal bulb. B. Reproductive system. C‒D. Bermudella lahainensis sp. nov. (CASIZ 199240). C. Buccal bulb. D. Reproductive system. E‒F. Bermudella mica (MNCN 15.05/94474). E. Buccal bulb. F. Reproductive system. G–H. Bermudella plebeia (MNCN 15.05/94477). G. Buccal bulb. H. Reproductive system. I‒J. Ceratodoris trypomandyas sp. nov. (QM MO 86034). I. Buccal bulb. J. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064316" httpUri="https://zenodo.org/record/11064316/files/figure.png" pageId="10" pageNumber="533">Fig. 2J</figureCitation>
|
||
). Reproductive system located in anterior third of body. Thin hermaphroditic duct begins at ovotestis, located inside digestive-hermaphroditic gland. Hermaphroditic duct expands into small, kidney shape ampulla. On opposite side of hermaphroditic duct, a postampullary duct emerges and joins prostate. Oviduct connects near union of postampullary duct with prostate. Short, oviduct enters inside female gland mass. Prostate very large, wide, and elongate, continues as thin and long vas deferens. Vas deferens expands to short ejaculatory duct. Penial spines were not appreciated. Vagina thin, half width than vas deferens, very long. Vagina connects with large, rounded bursa copulatrix. From bursa copulatrix arises a short and thin duct that connects with tiny, oval receptaculum seminis. At base of receptaculum seminis arises a thin and long uterine duct. Vagina and receptaculum seminis completely attached, resembling a single duct before separation during dissection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306657EFFC5EE1F00A94BEEFCCAF8E3" pageId="10" pageNumber="533" type="etymology">
|
||
<paragraph id="8BA336F5FFC5EE1F00A94BEEFCCAF8E3" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A94BEEFE8DF8C7" bold="true" box="[199,328,1791,1817]" pageId="10" pageNumber="533">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName id="4C1C4D76FFC5EE1F01A54BEEFDBCF8C7" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[459,633,1791,1817]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="10" pageNumber="533" phylum="Mollusca" rank="species" species="trypomandyas">“trypomandyas</taxonomicName>
|
||
” has Greek origin. Trypo means holes and mandyas mantle, referring to the light brown patches that resemble holes on the body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306657EFFC5EE1F00A94A56FC61F813" pageId="10" pageNumber="533" type="distribution">
|
||
<paragraph id="8BA336F5FFC5EE1F00A94A56FABBF8BF" blockId="10.[151,1437,423,1998]" box="[199,1406,1863,1890]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A94A56FE9CF8BF" bold="true" box="[199,345,1863,1889]" pageId="10" pageNumber="533">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F30B7665FFC5EE1F01094A56FE0BF8BF" box="[359,462,1863,1889]" name="Australia" pageId="10" pageNumber="533">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B04FFC5EE1F01B24A56FD45F8BF" author="Rudman, W. B." box="[476,640,1863,1889]" pageId="10" pageNumber="533" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B04FFC5EE1F02E54A56FC59F8BF" author="Debelius, H. & Kuiter, R. H." box="[651,924,1863,1889]" pageId="10" pageNumber="533" refId="ref15860" refString="Debelius, H. & Kuiter, R. H. (2007) Nudibranchs of the World. IKAN-Unterwasserarchiv, Frankfurt, 360 pp." type="book" year="2007">Debelius & Kuiter 2007</bibRefCitation>
|
||
; present study) and
|
||
<collectingCountry id="F30B7665FFC5EE1F04124A56FB7AF8BF" box="[1148,1215,1863,1889]" name="China" pageId="10" pageNumber="533">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B04FFC5EE1F04A34A56FAB7F8BF" author="Rudman, W. B." box="[1229,1394,1863,1889]" pageId="10" pageNumber="533" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC5EE1F00A94A7AFC61F813" blockId="10.[151,1437,423,1998]" pageId="10" pageNumber="533">
|
||
<emphasis id="B968EAE7FFC5EE1F00A94A7AFE46F85B" bold="true" box="[199,387,1899,1925]" pageId="10" pageNumber="533">Natural history</emphasis>
|
||
.
|
||
<bibRefCitation id="EF8D4B04FFC5EE1F01FB4A7AFD8FF85B" author="Rudman, W. B." box="[405,586,1899,1925]" pageId="10" pageNumber="533" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman (2004)</bibRefCitation>
|
||
found specimens on brown algae and on different species of bryozoans, including
|
||
<taxonomicName id="4C1C4D76FFC5EE1F01624A9EFD5CF877" authority="(Bosc, 1802)" baseAuthorityName="Bosc" baseAuthorityYear="1802" box="[268,665,1935,1961]" class="Gymnolaemata" family="Membraniporidae" genus="Jellyella" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="533" phylum="Bryozoa" rank="species" species="tuberculata">
|
||
<emphasis id="B968EAE7FFC5EE1F01624A9EFE32F877" box="[268,503,1935,1961]" italics="true" pageId="10" pageNumber="533">Jellyella tuberculata</emphasis>
|
||
(Bosc, 1802)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D76FFC5EE1F02C74A9EFB8DF877" authority="(Moll, 1803)" baseAuthorityName="Moll" baseAuthorityYear="1803" box="[681,1096,1935,1961]" class="Gymnolaemata" family="Cryptosulidae" genus="Cryptosula" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="533" phylum="Bryozoa" rank="species" species="pallasiana">
|
||
<emphasis id="B968EAE7FFC5EE1F02C74A9EFC63F877" box="[681,934,1935,1961]" italics="true" pageId="10" pageNumber="533">Cryptosula pallasiana</emphasis>
|
||
(Moll, 1803)
|
||
</taxonomicName>
|
||
. The author documented
|
||
<specimenCount id="9D1AFD7CFFC5EE1F051A4A81FEC4F813" count="1" pageId="10" pageNumber="533" type="generic">one specimen</specimenCount>
|
||
preying on
|
||
<taxonomicName id="4C1C4D76FFC5EE1F01E94AA2FC65F813" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[391,928,1971,1997]" class="Gymnolaemata" family="Membraniporidae" genus="Membranipora" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="533" phylum="Bryozoa" rank="species" species="membranacea">
|
||
<emphasis id="B968EAE7FFC5EE1F01E94AA2FD12F813" box="[391,727,1971,1997]" italics="true" pageId="10" pageNumber="533">Membranipora membranacea</emphasis>
|
||
(Linnaeus, 1767)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF63667DFFC4EE1E00F94BAEFB47F8B9" ID-DOI="http://doi.org/10.5281/zenodo.11064322" ID-Zenodo-Dep="11064322" httpUri="https://zenodo.org/record/11064322/files/figure.png" pageId="11" pageNumber="534" startId="11.[151,250,1727,1751]" targetBox="[232,1355,181,1702]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BA336F5FFC4EE1E00F94BAEFB47F8B9" blockId="11.[151,1437,1727,1895]" pageId="11" pageNumber="534">
|
||
<emphasis id="B968EAE7FFC4EE1E00F94BAEFEC9F909" bold="true" box="[151,268,1727,1751]" pageId="11" pageNumber="534">FIGURE 5</emphasis>
|
||
. Scanning electron micrographs and light microscope photographs of cuticles, radulae, and penis.
|
||
<emphasis id="B968EAE7FFC4EE1E048B4BAEFAD3F909" bold="true" box="[1253,1302,1727,1751]" pageId="11" pageNumber="534">A‒D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1C4D76FFC4EE1E054C4BAEFEE2F925" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC4EE1E054C4BAEFEE2F925" italics="true" pageId="11" pageNumber="534">Ceratodoris trypomandyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC4EE1E01414BF2FE45F924" box="[303,384,1763,1786]" pageId="11" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B968EAE7FFC4EE1E01E94BF2FE5CF925" bold="true" box="[391,409,1763,1787]" pageId="11" pageNumber="534">A</emphasis>
|
||
. Detail of labial cuticle elements (QM MO 86034).
|
||
<emphasis id="B968EAE7FFC4EE1E03D94BF2FC0DF925" bold="true" box="[951,968,1763,1787]" pageId="11" pageNumber="534">B</emphasis>
|
||
. Frontal view of radula (QM MO 86034).
|
||
<emphasis id="B968EAE7FFC4EE1E05EA4BF2FA53F925" bold="true" box="[1412,1430,1763,1787]" pageId="11" pageNumber="534">C</emphasis>
|
||
. Detail of inner and outer lateral teeth (QM MO 86034).
|
||
<emphasis id="B968EAE7FFC4EE1E02B04A16FD35F8C1" bold="true" box="[734,752,1799,1823]" pageId="11" pageNumber="534">D</emphasis>
|
||
. Detail of outer lateral teeth (AM C. 270712).
|
||
<emphasis id="B968EAE7FFC4EE1E04B64A16FACFF8C1" bold="true" box="[1240,1290,1799,1823]" pageId="11" pageNumber="534">E‒H</emphasis>
|
||
.
|
||
<taxonomicName id="4C1C4D76FFC4EE1E05764A16FEC3F89D" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" class="Gastropoda" family="Goniodorididae" genus="Murphydoris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="species" species="polkadotsa" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC4EE1E05764A16FEC3F89D" italics="true" pageId="11" pageNumber="534">Murphydoris polkadotsa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC4EE1E01634A3AFE98F89C" box="[269,349,1835,1858]" pageId="11" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(QM MO 86038).
|
||
<emphasis id="B968EAE7FFC4EE1E02704A3AFDEAF89D" bold="true" box="[542,559,1835,1859]" pageId="11" pageNumber="534">E</emphasis>
|
||
. Frontal view of radula.
|
||
<emphasis id="B968EAE7FFC4EE1E03444A3AFCFCF89D" bold="true" box="[810,825,1835,1859]" pageId="11" pageNumber="534">F</emphasis>
|
||
. Detail of inner lateral teeth.
|
||
<emphasis id="B968EAE7FFC4EE1E040A4A3AFBBDF89D" bold="true" box="[1124,1144,1835,1859]" pageId="11" pageNumber="534">G</emphasis>
|
||
. Detail of outer lateral teeth.
|
||
<emphasis id="B968EAE7FFC4EE1E00F94A5EFF6EF8B9" bold="true" box="[151,171,1871,1895]" pageId="11" pageNumber="534">H</emphasis>
|
||
. Detail of penial spines. Scale bars: A, G: 5 μm; B, E: 50 μm; C: 30 μm; D: 10 μm; F, H: 20 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C306657EFFC4EE1900A94AA0FEC3FB87" lastPageId="12" lastPageNumber="535" pageId="11" pageNumber="534" type="discussion">
|
||
<paragraph id="8BA336F5FFC4EE1900A94AA0FB9BFE9F" blockId="11.[151,1436,1969,2032]" lastBlockId="12.[151,1437,151,1113]" lastPageId="12" lastPageNumber="535" pageId="11" pageNumber="534">
|
||
<emphasis id="B968EAE7FFC4EE1E00A94AA0FEF1F815" bold="true" box="[199,308,1969,1995]" pageId="11" pageNumber="534">Remarks</emphasis>
|
||
. Within the genus
|
||
<taxonomicName id="4C1C4D76FFC4EE1E02644AA0FD54F815" authorityName="Gray" authorityYear="1850" box="[522,657,1969,1995]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAE7FFC4EE1E02644AA0FD54F815" box="[522,657,1969,1995]" italics="true" pageId="11" pageNumber="534">Ceratodoris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1C4D76FFC4EE1E02CE4AA3FC59F815" authority="(Baba, 1960)" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[672,924,1969,1996]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC4EE1E02CE4AA3FCC7F815" box="[672,770,1969,1996]" italics="true" pageId="11" pageNumber="534">C. plana</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B04FFC4EE1E037F4AA0FC56F815" author="Baba, K." box="[785,915,1969,1995]" pageId="11" pageNumber="534" pagination="79 - 83" refId="ref15223" refString="Baba, K. (1960) The genera Okenia, Goniodoridella and Goniodoris from Japan (Nudibranchia Goniodorididae). Publications of the Seto marine Biological Laboratory, 8 (1), 79 - 83. https: // doi. org / 10.5134 / 174698" type="journal article" year="1960">Baba, 1960</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D76FFC4EE1E03BD4AA3FA9FF815" authority="(Bouchet & Ortea, 1983)" baseAuthorityName="Bouchet & Ortea" baseAuthorityYear="1983" box="[979,1370,1969,1996]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B968EAE7FFC4EE1E03BD4AA3FBFDF815" box="[979,1080,1969,1996]" italics="true" pageId="11" pageNumber="534">C. pilosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B04FFC4EE1E04264AA0FA94F815" author="Bouchet, P. & Ortea, J." box="[1096,1361,1969,1995]" pageId="11" pageNumber="534" pagination="227 - 233" refId="ref15337" refString="Bouchet, P. & Ortea, J. (1983) Hopkinsia pilosa, new species feeding on Bryozoa in the South Pacific (Mollusca, Opisthobranchia). Venus, 42, 227 - 233." type="journal article" year="1983">Bouchet & Ortea, 1983</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
share with
|
||
<taxonomicName id="4C1C4D76FFC4EE1E00BE4AC7FE55F831" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[208,400,2005,2032]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="534" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC4EE1E00BE4AC7FE55F831" box="[208,400,2005,2032]" italics="true" pageId="11" pageNumber="534">C. trypomandyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC4EE1E01F84AC7FE2AF82E" box="[406,495,2006,2032]" pageId="11" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
|
||
a flattened body bearing dorsal thin papillae, with brownish speckles on translucent whitish background.
|
||
<taxonomicName id="4C1C4D76FFC3EE1901EB4D86FC41FF6F" authority="(Quoy & Gaimard, 1832)" baseAuthorityName="Quoy & Gaimard" baseAuthorityYear="1832" box="[389,900,151,177]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="eolida">
|
||
<emphasis id="B968EAE7FFC3EE1901EB4D86FD9EFF6F" box="[389,603,151,177]" italics="true" pageId="12" pageNumber="535">Ceratodoris eolida</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B04FFC3EE1902054D86FCBEFF6F" author="Quoy, J. R. C. & Gaimard, J. P." box="[619,891,151,177]" pageId="12" pageNumber="535" pagination="1 - 390" refId="ref17779" refString="Quoy, J. R. C. & Gaimard, J. P. (1832 - 1835) Voyage de la corvette l'Astrolabe: execute par ordre du roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. Zoologie. 1, 2 (1), 2 (2), 3 (1), 3 (2), 4 & Atlas (Mollusques). In: Dumont d'Urville, J. (1834), Voyage de Decouvertes de l'Astrolabe. J. Tastu, Editeur-Imprimeur, Paris, pp. i - l + 1 - 264 (1830), pp. 1 - 321 (1832), pp. 321 - 686 (1833), pp. 1 - 366 (1834), pp. 367 - 954 (1835), pp. 1 - 390 (1833), pls. 1 - 93 (1833). https: // doi. org / 10.5962 / bhl. title. 2132" type="journal article" year="1832">Quoy & Gaimard, 1832</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has a similar body shape but it has white body and few red patches (
|
||
<bibRefCitation id="EF8D4B04FFC3EE1901EE4DAAFD40FF0B" author="Quoy, J. R. C. & Gaimard, J. P." box="[384,645,187,213]" pageId="12" pageNumber="535" pagination="1 - 390" refId="ref17779" refString="Quoy, J. R. C. & Gaimard, J. P. (1832 - 1835) Voyage de la corvette l'Astrolabe: execute par ordre du roi, pendant les annees 1826 - 1827 - 1828 - 1829, sous le commandement de M. J. Dumont d'Urville. Zoologie. 1, 2 (1), 2 (2), 3 (1), 3 (2), 4 & Atlas (Mollusques). In: Dumont d'Urville, J. (1834), Voyage de Decouvertes de l'Astrolabe. J. Tastu, Editeur-Imprimeur, Paris, pp. i - l + 1 - 264 (1830), pp. 1 - 321 (1832), pp. 321 - 686 (1833), pp. 1 - 366 (1834), pp. 367 - 954 (1835), pp. 1 - 390 (1833), pls. 1 - 93 (1833). https: // doi. org / 10.5962 / bhl. title. 2132" type="journal article" year="1832">Quoy & Gaimard 1832</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1C4D76FFC3EE1902F94DAAFCA1FF0B" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[663,868,187,213]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE1902F94DAAFCA1FF0B" box="[663,868,187,213]" italics="true" pageId="12" pageNumber="535">Ceratodoris plana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D76FFC3EE1903F64DAAFC3EFF0B" baseAuthorityName="Bouchet & Ortea" baseAuthorityYear="1983" box="[920,1019,187,213]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B968EAE7FFC3EE1903F64DAAFC3EFF0B" box="[920,1019,187,213]" italics="true" pageId="12" pageNumber="535">C. pilosa</emphasis>
|
||
</taxonomicName>
|
||
differ by having five pairs of papillae and several dozen papillae, respectively (
|
||
<bibRefCitation id="EF8D4B04FFC3EE19020C4DCEFCA3FF27" author="Bouchet, P. & Ortea, J." box="[610,870,223,249]" pageId="12" pageNumber="535" pagination="227 - 233" refId="ref15337" refString="Bouchet, P. & Ortea, J. (1983) Hopkinsia pilosa, new species feeding on Bryozoa in the South Pacific (Mollusca, Opisthobranchia). Venus, 42, 227 - 233." type="journal article" year="1983">Bouchet & Ortea 1983</bibRefCitation>
|
||
). Moreover,
|
||
<taxonomicName id="4C1C4D76FFC3EE19039B4DCEFB93FF27" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[1013,1110,223,249]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE19039B4DCEFB93FF27" box="[1013,1110,223,249]" italics="true" pageId="12" pageNumber="535">C. plana</emphasis>
|
||
</taxonomicName>
|
||
has one papilla in the central part of the body while
|
||
<taxonomicName id="4C1C4D76FFC3EE1901C84C12FDCBFEC3" baseAuthorityName="Bouchet & Ortea" baseAuthorityYear="1983" box="[422,526,259,285]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="pilosa">
|
||
<emphasis id="B968EAE7FFC3EE1901C84C12FDCBFEC3" box="[422,526,259,285]" italics="true" pageId="12" pageNumber="535">C. pilosa</emphasis>
|
||
</taxonomicName>
|
||
has five (
|
||
<bibRefCitation id="EF8D4B04FFC3EE1902E84C12FC51FEC3" author="Bouchet, P. & Ortea, J." box="[646,916,259,285]" pageId="12" pageNumber="535" pagination="227 - 233" refId="ref15337" refString="Bouchet, P. & Ortea, J. (1983) Hopkinsia pilosa, new species feeding on Bryozoa in the South Pacific (Mollusca, Opisthobranchia). Venus, 42, 227 - 233." type="journal article" year="1983">Bouchet & Ortea 1983</bibRefCitation>
|
||
). The new species
|
||
<taxonomicName id="4C1C4D76FFC3EE19041A4C12FAF2FEC3" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[1140,1335,259,285]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC3EE19041A4C12FAF2FEC3" box="[1140,1335,259,285]" italics="true" pageId="12" pageNumber="535">C. trypomandyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC3EE19052F4C12FA58FEC3" box="[1345,1437,259,285]" pageId="12" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
|
||
resembles
|
||
<emphasis id="B968EAE7FFC3EE1901604C36FEB0FE9F" box="[270,373,295,321]" italics="true" pageId="12" pageNumber="535">
|
||
<taxonomicName id="4C1C4D76FFC3EE1901604C36FEB7FE9F" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[270,370,295,321]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">C. plana</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
having five pairs of papillae and one additional middorsal papilla.
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC3EE1900A94C5AFC9FFCCB" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="535">
|
||
After its original description,
|
||
<taxonomicName id="4C1C4D76FFC3EE1902754C5AFDBBFEBB" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[539,638,331,357]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE1902754C5AFDBBFEBB" box="[539,638,331,357]" italics="true" pageId="12" pageNumber="535">C. plana</emphasis>
|
||
</taxonomicName>
|
||
was redescribed by
|
||
<bibRefCitation id="EF8D4B04FFC3EE1903054C5AFBDAFEBB" author="Gosliner, T. M." box="[875,1055,331,357]" pageId="12" pageNumber="535" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner (2004)</bibRefCitation>
|
||
using specimens from
|
||
<collectingRegion id="49D8F817FFC3EE1905454C5AFA58FEBB" box="[1323,1437,331,357]" country="United States of America" name="California" pageId="12" pageNumber="535">California</collectingRegion>
|
||
(
|
||
<collectingCountry id="F30B7665FFC3EE1900F14C7EFF1FFE57" box="[159,218,367,393]" name="United States of America" pageId="12" pageNumber="535">USA</collectingCountry>
|
||
) and the
|
||
<collectingCountry id="F30B7665FFC3EE19013E4C7EFE14FE57" box="[336,465,367,393]" name="Philippines" pageId="12" pageNumber="535">Philippines</collectingCountry>
|
||
, and by
|
||
<bibRefCitation id="EF8D4B04FFC3EE1902504C7EFD36FE57" author="Rudman, W. B." box="[574,755,367,393]" pageId="12" pageNumber="535" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman (2004)</bibRefCitation>
|
||
using specimens from
|
||
<collectingCountry id="F30B7665FFC3EE1904654C7EFBB7FE57" box="[1035,1138,367,393]" name="Australia" pageId="12" pageNumber="535">Australia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30B7665FFC3EE1904DF4C7EFB3DFE57" box="[1201,1272,367,393]" name="China" pageId="12" pageNumber="535">China</collectingCountry>
|
||
. Both authors described the species having five pairs of papillae and a single middorsal one. However, the color pattern of the specimens of each author did not completely match.
|
||
<bibRefCitation id="EF8D4B04FFC3EE1902B34CA6FC4AFE0F" author="Gosliner, T. M." box="[733,911,439,465]" pageId="12" pageNumber="535" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner (2004)</bibRefCitation>
|
||
described the species as “translucent yellowish white with the exception of small brown spots scattered over the surface of the notum and occasional opaque white spots”.
|
||
<bibRefCitation id="EF8D4B04FFC3EE1900834CEEFE5AFDC7" author="Rudman, W. B." box="[237,415,511,537]" pageId="12" pageNumber="535" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman (2004)</bibRefCitation>
|
||
described a variation of color, depending on the locality. Specimens from
|
||
<collectingCountry id="F30B7665FFC3EE1904994CEEFA9BFDC7" box="[1271,1374,511,537]" name="Australia" pageId="12" pageNumber="535">Australia</collectingCountry>
|
||
were described as “translucent white with closely packed, relatively large, brown patches, […] each brown patch is edged with a thin opaque white line. Sometimes the opaque white pigment is absent.” Specimens from
|
||
<collectingCountry id="F30B7665FFC3EE1904B44F56FA9BFDBF" box="[1242,1374,583,609]" name="Hong Kong" pageId="12" pageNumber="535">Hong Kong</collectingCountry>
|
||
were described having small dark brown spots instead large brown patches. In addition to these subtle variations in the color pattern, the anatomy of the reproductive system of the specimens used by
|
||
<bibRefCitation id="EF8D4B04FFC3EE1904694F9EFB7CFD77" author="Gosliner, T. M." box="[1031,1209,655,681]" pageId="12" pageNumber="535" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner (2004)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8D4B04FFC3EE1904834F9EFA59FD77" author="Rudman, W. B." box="[1261,1436,655,681]" pageId="12" pageNumber="535" pagination="1 - 70" refId="ref18018" refString="Rudman, W. B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 695.1.1" type="journal article" year="2004">Rudman (2004)</bibRefCitation>
|
||
also disagreed. The receptaculum seminis joins at the base of the bursa copulatrix, together with a long uterine duct in Rudman’s specimens. In Gosliner’s specimens, an elongate and narrow duct arises from the base of the bursa copulatrix and connects with a pyriform receptaculum seminis.
|
||
</paragraph>
|
||
<paragraph id="8BA336F5FFC3EE1900A94E0EFEC3FB87" blockId="12.[151,1437,151,1113]" pageId="12" pageNumber="535">
|
||
These differences in external and internal anatomy support the possibility that several different species have been described under the name of
|
||
<taxonomicName id="4C1C4D76FFC3EE1902794E52FDBDFC83" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[535,632,835,861]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE1902794E52FDBDFC83" box="[535,632,835,861]" italics="true" pageId="12" pageNumber="535">C. plana</emphasis>
|
||
</taxonomicName>
|
||
due to their external similarity. The existence of several similar species of
|
||
<taxonomicName id="4C1C4D76FFC3EE1900D84E76FEF8FC5F" authorityName="Gray" authorityYear="1850" box="[182,317,871,897]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAE7FFC3EE1900D84E76FEF8FC5F" box="[182,317,871,897]" italics="true" pageId="12" pageNumber="535">Ceratodoris</emphasis>
|
||
</taxonomicName>
|
||
is also supported by molecular analyses (
|
||
<bibRefCitation id="EF8D4B04FFC3EE19037D4E76FBDBFC5F" author="Paz-Sedano, S. & Moles, J. & Smirnoff, D. & Gosliner, T. M. & Pola, M." box="[787,1054,871,897]" pageId="12" pageNumber="535" pagination="107990" refId="ref17463" refString="Paz-Sedano, S., Moles, J., Smirnoff, D., Gosliner, T. M. & Pola, M. (2024) A combined phylogenetic strategy illuminates the evolution of Goniodorididae nudibranchs (Mollusca, Gastropoda, Heterobranchia). Molecular Phylogenetics and Evolution, 192, 107990. https: // doi. org / 10.1016 / j. ympev. 2023.107990" type="journal article" year="2024">
|
||
Paz-Sedano
|
||
<emphasis id="B968EAE7FFC3EE1903CE4E76FC1CFC5F" box="[928,985,871,897]" italics="true" pageId="12" pageNumber="535">et al.</emphasis>
|
||
2024
|
||
</bibRefCitation>
|
||
). In the original description,
|
||
<bibRefCitation id="EF8D4B04FFC3EE19050D4E76FF27FC7B" author="Baba, K." pageId="12" pageNumber="535" pagination="79 - 83" refId="ref15223" refString="Baba, K. (1960) The genera Okenia, Goniodoridella and Goniodoris from Japan (Nudibranchia Goniodorididae). Publications of the Seto marine Biological Laboratory, 8 (1), 79 - 83. https: // doi. org / 10.5134 / 174698" type="journal article" year="1960">Baba (1960)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C1C4D76FFC3EE1901344E9AFE7FFC7B" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[346,442,907,933]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE1901344E9AFE7FFC7B" box="[346,442,907,933]" italics="true" pageId="12" pageNumber="535">C. plana</emphasis>
|
||
</taxonomicName>
|
||
as having a plentifully sprinkled with minute spots of chocolate brown on the translucent whitish ground, which is most consistent with Gosliner´s (2004) description. Moreover,
|
||
<taxonomicName id="4C1C4D76FFC3EE1904FE4EBEFB36FC17" baseAuthorityName="Baba" baseAuthorityYear="1960" box="[1168,1267,943,969]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="plana">
|
||
<emphasis id="B968EAE7FFC3EE1904FE4EBEFB36FC17" box="[1168,1267,943,969]" italics="true" pageId="12" pageNumber="535">C. plana</emphasis>
|
||
</taxonomicName>
|
||
was described having yellow rhinophores, and the tips of the papillae densely whitish.
|
||
<taxonomicName id="4C1C4D76FFC3EE1903A54EC2FB3CFC33" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[971,1273,979,1005]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC3EE1903A54EC2FB3CFC33" box="[971,1273,979,1005]" italics="true" pageId="12" pageNumber="535">Ceratodoris trypomandyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC3EE19056F4EC2FA9EFC33" box="[1281,1371,979,1005]" pageId="12" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
|
||
lacks yellow color and the papillae have brownish patches.
|
||
<bibRefCitation id="EF8D4B04FFC3EE1902984EE6FC47FBCF" author="Baba, K." box="[758,898,1015,1041]" pageId="12" pageNumber="535" pagination="79 - 83" refId="ref15223" refString="Baba, K. (1960) The genera Okenia, Goniodoridella and Goniodoris from Japan (Nudibranchia Goniodorididae). Publications of the Seto marine Biological Laboratory, 8 (1), 79 - 83. https: // doi. org / 10.5134 / 174698" type="journal article" year="1960">Baba (1960)</bibRefCitation>
|
||
did not give details of the reproductive system. Rudman´s (2004) description matches the new species
|
||
<taxonomicName id="4C1C4D76FFC3EE190293490AFC78FBEB" authority="Paz-Sedano & Cobb & Gosliner & Pola, 2024" authorityName="Paz-Sedano & Cobb & Gosliner & Pola" authorityYear="2024" box="[765,957,1051,1077]" class="Gastropoda" family="Goniodorididae" genus="Ceratodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="535" phylum="Mollusca" rank="species" species="trypomandyas" status="sp. nov.">
|
||
<emphasis id="B968EAE7FFC3EE190293490AFC78FBEB" box="[765,957,1051,1077]" italics="true" pageId="12" pageNumber="535">C. trypomandyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B579CFFC3EE1903AD490AFBD8FBEB" box="[963,1053,1051,1077]" pageId="12" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, who also studied specimens from
|
||
<collectingCountry id="F30B7665FFC3EE1900F9492EFEC4FB87" box="[151,257,1087,1113]" name="Australia" pageId="12" pageNumber="535">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |