178 lines
18 KiB
XML
178 lines
18 KiB
XML
<document id="F79CC2DC55EF52190FFE8BDC3772A9EA" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC10FB179CFD7CEAFFC98F856" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Xerotyphlops Hedges, Marion, Lipp, Marin & Vidal 2014" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="54" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="D4BC87BC1025201480F74BD8FB8F8DF1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="88AA84955EEF66E1530958FB7E53D2D4">
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<mods:title id="11BBE9BE17DCE8510154C5A9437A8644">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="2C7F974297D87141B9E0627D4B135B48" type="personal">
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<mods:namePart id="274B073AA55C3DD235D3DE66F53B7088">Pyron, Robert Alexander</mods:namePart>
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</mods:name>
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<mods:name id="4E6E4E39FDAA08EC175E7412CFB9F243" type="personal">
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<mods:roleTerm id="98AA024EB4A299420F5A606A7E275F9D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="31A508D9903FAEA6FDED80DB2E4A7F94">Wallach, Van</mods:namePart>
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<mods:typeOfResource id="E674CBDB0D0D376AA8CA4091E4F8C48B">text</mods:typeOfResource>
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<mods:title id="2244471F3D2D51A300339E24DC7D04AF">Zootaxa</mods:title>
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<mods:part id="98D776AD1C044C499D8A8711C502C97A">
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<mods:date id="04F0CCB650EB96FCA8AD6916C03BC259">2014</mods:date>
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<mods:detail id="B7CB336F16D208BC0DC1D703A1BC03BE" type="volume">
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<mods:number id="82B8FB6FAA33C414516046C82A48F998">3829</mods:number>
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<mods:classification id="AE627015E052E15B71B0B45E379CB047">journal article</mods:classification>
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<mods:identifier id="08FEFB47EFF35C09FD1280992DC73528" type="CLB-Dataset">36820</mods:identifier>
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<mods:identifier id="2BD9360FE944F286D9F9321786C73AB7" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<mods:identifier id="C602DBB56DF552D1045E93AAEBD1284B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B587DDC10FB179CFD7CEAFFC98F856" ID-DOI="http://doi.org/10.5281/zenodo.6127992" ID-GBIF-Taxon="127668689" ID-Zenodo-Dep="6127992" LSID="urn:lsid:plazi:treatment:03B587DDC10FB179CFD7CEAFFC98F856" httpUri="http://treatment.plazi.org/id/03B587DDC10FB179CFD7CEAFFC98F856" lastPageNumber="54" pageId="53" pageNumber="54">
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<subSubSection id="C3066540C10FB179CFD7CEAFFCDAFB19" box="[151,867,1185,1211]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="8BA336CBC10FB179CFD7CEAFFCDAFB19" blockId="53.[151,867,1185,1211]" box="[151,867,1185,1211]" pageId="53" pageNumber="54">
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<heading id="D0EB81A7C10FB179CFD7CEAFFCDAFB19" bold="true" box="[151,867,1185,1211]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
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<taxonomicName id="4C1C4D48C10FB179CFD7CEAFFCDAFB19" authority="Hedges, Marion, Lipp, Marin & Vidal, 2014" authorityName="Hedges, Marion, Lipp, Marin & Vidal" authorityYear="2014" box="[151,867,1185,1211]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10FB179CFD7CEAFFCDAFB19" bold="true" box="[151,867,1185,1211]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CFD7CEAFFE8FFB19" bold="true" box="[151,310,1185,1211]" italics="true" pageId="53" pageNumber="54">Xerotyphlops</emphasis>
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<bibRefCitation id="EF8D4B3AC10FB179CE7ECEAFFCDAFB19" author="Hedges" box="[318,867,1185,1211]" pageId="53" pageNumber="54" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article" year="2014">Hedges, Marion, Lipp, Marin & Vidal, 2014</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CFD7CEE9FAA6FA86" pageId="53" pageNumber="54" type="materials_examined">
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<paragraph id="8BA336CBC10FB179CFD7CEE9FD60FAA2" blockId="53.[151,1437,1255,2037]" box="[151,729,1255,1280]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CFD7CEE9FE8BFAA2" bold="true" box="[151,306,1255,1280]" pageId="53" pageNumber="54">
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<typeStatus id="54A78869C10FB179CFD7CEE9FF6BFAA2" box="[151,210,1255,1280]" pageId="53" pageNumber="54">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C10FB179CE79CEE9FD60FAA2" authority="Merrem, 1820" authorityName="Merrem" authorityYear="1820" box="[313,729,1255,1280]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vermicularis">
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<emphasis id="B968EAD9C10FB179CE79CEE9FD88FAA2" box="[313,561,1255,1280]" italics="true" pageId="53" pageNumber="54">Typhlops vermicularis</emphasis>
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<bibRefCitation id="EF8D4B3AC10FB179CD79CEE6FD60FAA2" author="Merrem" box="[569,729,1256,1280]" pageId="53" pageNumber="54" refString="Merrem, B. (1820) Versuch eines Systems der Amphibien. Johannes Christin Krieger, Marburg, 191 pp." type="book" year="1820">Merrem, 1820</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC10FB179CF87CF02FAA6FA86" blockId="53.[151,1437,1255,2037]" box="[199,1311,1292,1317]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CF02FE3CFA87" bold="true" box="[199,389,1292,1317]" pageId="53" pageNumber="54">Species content.</emphasis>
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<emphasis id="B968EAD9C10FB179CECCCF02FB8EFA86" box="[396,1079,1292,1317]" italics="true" pageId="53" pageNumber="54">
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<taxonomicName id="4C1C4D48C10FB179CECCCF02FD6FFA81" authority="Xer." authorityName="Xer." box="[396,726,1292,1317]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="etheridgei">
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Xerotyphlops etheridgei,
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<emphasis id="B968EAD9C10FB179CDE6CF02FD6FFA81" bold="true" box="[678,726,1292,1315]" italics="true" pageId="53" pageNumber="54">Xer.</emphasis>
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</taxonomicName>
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socotranus,
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<emphasis id="B968EAD9C10FB179CC29CF02FB96FA87" bold="true" box="[873,1071,1292,1317]" italics="true" pageId="53" pageNumber="54">
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Xer.
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<taxonomicName id="4C1C4D48C10FB179CCDFCF02FB96FA87" box="[927,1071,1292,1317]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vermicularis">vermicularis</taxonomicName>
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</emphasis>
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,
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</emphasis>
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and
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<emphasis id="B968EAD9C10FB179CB2DCF03FB4BFA87" box="[1133,1266,1292,1317]" italics="true" pageId="53" pageNumber="54">
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Xer.
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<taxonomicName id="4C1C4D48C10FB179CBE1CF02FB4BFA87" box="[1185,1266,1292,1317]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="wilsoni">wilsoni</taxonomicName>
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</emphasis>
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(?).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CF87CF21FDAEF977" pageId="53" pageNumber="54" type="diagnosis">
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<paragraph id="8BA336CBC10FB179CF87CF21FCD8F9CA" blockId="53.[151,1437,1255,2037]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CF21FEF8FAEA" bold="true" box="[199,321,1327,1352]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C10FB179CE09CF21FE64FAEA" box="[329,477,1327,1352]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10FB179CE09CF21FE64FAEA" box="[329,477,1327,1352]" italics="true" pageId="53" pageNumber="54">Xerotyphlops</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by the combination of the following characters: small- to moderate-sized (total length
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<quantity id="4CE49B2EC10FB179CDA3CF5AFCCDFACE" box="[739,884,1364,1389]" metricMagnitude="-1" metricUnit="m" metricValue="2.485" metricValueMax="4.05" metricValueMin="0.9199999999999999" pageId="53" pageNumber="54" unit="mm" value="248.5" valueMax="405.0" valueMin="92.0">92–405 mm</quantity>
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), stout- to moderate-bodied (length/width ratio 34–56) snakes with 20–30 scale rows (usually with reduction), 346–435 total middorsals, short to moderate tail (1.1–3.1% total length) with 7–13 subcaudals (length/width ratio 1.0–1.5), and apical spine. Dorsal and lateral head profiles bluntly rounded, narrow rostral (0.27–0.41 head width), inferior nasal suture in contact with second supralabial, preocular in contact with second and third supralabials, eye moderate with distinct pupil, T-III or T-V
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<collectingCountry id="F30B765BC10FB179CFD7CC06FF06F982" box="[151,191,1544,1568]" name="Cyprus" pageId="53" pageNumber="54">SIP</collectingCountry>
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, and postoculars 2–3. Lateral tongue papillae present; left lung absent, tracheal, cardiac and right lungs multicameral (with 22–29 + 5–9 + 2–9 chambers, respectively); testes segmented; hemipenis eversible, lacking retrocloacal sacs; and rectal caecum moderate (1.4–2.2% SVL).
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</paragraph>
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<paragraph id="8BA336CBC10FB179CF87CC7AFDAEF977" blockId="53.[151,1437,1255,2037]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CC7AFE5EF92F" bold="true" box="[199,487,1652,1677]" pageId="53" pageNumber="54">Phylogenetic definition.</emphasis>
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Includes the MRCA of
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<taxonomicName id="4C1C4D48C10FB179CC55CC7AFBFCF92F" box="[789,1093,1652,1677]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vermicularis">
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<emphasis id="B968EAD9C10FB179CC55CC7AFBFCF92F" box="[789,1093,1652,1677]" italics="true" pageId="53" pageNumber="54">Xerotyphlops vermicularis</emphasis>
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</taxonomicName>
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and
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<emphasis id="B968EAD9C10FB179CBC8CC7BFA84F92E" box="[1160,1341,1653,1676]" italics="true" pageId="53" pageNumber="54">Xer. socotranus</emphasis>
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and all descendants thereof, and all species more closely related to
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<emphasis id="B968EAD9C10FB179CC0CCC97FBB6F912" box="[844,1039,1687,1712]" italics="true" pageId="53" pageNumber="54">
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Xer.
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<taxonomicName id="4C1C4D48C10FB179CCC2CC99FBB6F912" box="[898,1039,1687,1712]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vermicularis">vermicularis</taxonomicName>
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</emphasis>
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than to the
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<typeStatus id="54A78869C10FB179CBDECC96FB77F912" box="[1182,1230,1688,1712]" pageId="53" pageNumber="54">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CF87CCD1FC10F95A" box="[199,937,1759,1784]" pageId="53" pageNumber="54" type="etymology">
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<paragraph id="8BA336CBC10FB179CF87CCD1FC10F95A" blockId="53.[151,1437,1255,2037]" box="[199,937,1759,1784]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CCD1FEF7F95A" bold="true" box="[199,334,1759,1784]" pageId="53" pageNumber="54">Etymology.</emphasis>
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Named after the desert (xeric) habitats of the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CF87CD0AFD18F8C6" pageId="53" pageNumber="54" type="distribution">
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<paragraph id="8BA336CBC10FB179CF87CD0AFD18F8C6" blockId="53.[151,1437,1255,2037]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CD0AFED9F8BF" bold="true" box="[199,352,1796,1821]" pageId="53" pageNumber="54">Distribution.</emphasis>
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The
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<typeStatus id="54A78869C10FB179CEE4CD0BFE6DF8BF" box="[420,468,1797,1821]" pageId="53" pageNumber="54">type</typeStatus>
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species (
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<taxonomicName id="4C1C4D48C10FB179CD06CD0AFCCDF8BF" box="[582,884,1796,1821]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vermicularis">
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<emphasis id="B968EAD9C10FB179CD06CD0AFCCDF8BF" box="[582,884,1796,1821]" italics="true" pageId="53" pageNumber="54">Xerotyphlops vermicularis</emphasis>
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</taxonomicName>
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) occurs from southeastern Europe through the Middle East. The other three species are more localized, with
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<emphasis id="B968EAD9C10FB179CC28CD27FBA9F8E2" box="[872,1040,1831,1856]" italics="true" pageId="53" pageNumber="54">
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Xer.
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<taxonomicName id="4C1C4D48C10FB179CCDECD29FBA9F8E2" box="[926,1040,1831,1856]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="etheridgei">etheridgei</taxonomicName>
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</emphasis>
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in
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<collectingCountry id="F30B765BC10FB179CB79CD29FB0EF8E2" box="[1081,1207,1831,1856]" name="Mauritania" pageId="53" pageNumber="54">Mauritania</collectingCountry>
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,
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<emphasis id="B968EAD9C10FB179CB85CD27FACFF8E2" box="[1221,1398,1833,1856]" italics="true" pageId="53" pageNumber="54">Xer. socotranus</emphasis>
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on
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<collectingCountry id="F30B765BC10FB179CFD7CD42FF56F8C6" box="[151,239,1868,1892]" name="Yemen" pageId="53" pageNumber="54">Socotra</collectingCountry>
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, and
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<emphasis id="B968EAD9C10FB179CE6ACD43FE17F8C7" box="[298,430,1868,1893]" italics="true" pageId="53" pageNumber="54">
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Xer.
|
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<taxonomicName id="4C1C4D48C10FB179CE1DCD42FE17F8C7" box="[349,430,1868,1893]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="wilsoni">wilsoni</taxonomicName>
|
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</emphasis>
|
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in southwestern
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<collectingCountry id="F30B765BC10FB179CD2CCD43FD24F8C6" box="[620,669,1869,1892]" name="Iran" pageId="53" pageNumber="54">Iran</collectingCountry>
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.
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</paragraph>
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</subSubSection>
|
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<subSubSection id="C3066540C10FB179CF87CD61FC98F856" pageId="53" pageNumber="54" type="discussion">
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<paragraph id="8BA336CBC10FB179CF87CD61FC98F856" blockId="53.[151,1437,1255,2037]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CD61FE82F82A" bold="true" box="[199,315,1903,1928]" pageId="53" pageNumber="54">Remarks.</emphasis>
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Assignment of
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<taxonomicName id="4C1C4D48C10FB179CEAFCD61FD64F82A" box="[495,733,1903,1928]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="wilsoni">
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<emphasis id="B968EAD9C10FB179CEAFCD61FD64F82A" box="[495,733,1903,1928]" italics="true" pageId="53" pageNumber="54">Xerotyphlops wilsoni</emphasis>
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</taxonomicName>
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is tentative, as it is only known from the lost
|
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<typeStatus id="54A78869C10FB179CBA7CD61FAF5F82A" box="[1255,1356,1903,1928]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
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, which has not been examined, though other putative specimens have been sighted near the
|
||
<typeStatus id="54A78869C10FB179CB13CD9BFB3AF80F" box="[1107,1155,1941,1965]" pageId="53" pageNumber="54">type</typeStatus>
|
||
locality (B. Safaei, pers. comm.). Little morphological data is known for this species from the brief description (Table 2). This group likely contains additional cryptic species (
|
||
<bibRefCitation id="EF8D4B3AC10FB179CD66CDD2FCACF856" author="Kornilios" box="[550,789,2012,2037]" pageId="53" pageNumber="54" refString="Kornilios, P., Ilgaz, C., Kumlutas, Y., Lymberakis, P., Moravec, J., Sindaco, R., Rastegar-Pouyani, N., Afroosheh, M., Giokas, S., Fraguedakis-Tsolis, S. & Chondropoulos, B. (2012) Neogene climatic oscillations shape the biogeography and evolutionary history of the Eurasian blindsnake. Molecular Phylogenetics and Evolution, 62, 856 - 873. http: // dx. doi. org / 10.1016 / j. ympev. 2011.11.035" type="journal article" year="2012">
|
||
Kornilios
|
||
<emphasis id="B968EAD9C10FB179CDD9CDD3FD68F857" box="[665,721,2012,2037]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |