156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="10BFBAB3C62F667E6B44C7DF12E4E7E8" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC10CB17BCFD7CD00FEE0FD6E" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Malayotyphlops Hedges, Marion, Lipp, Marin & Vidal 2014" docType="treatment" docVersion="7" lastPageNumber="56" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="55" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="B71450778B638B045E56F883953EB74D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D13151A1640E4493586A534D371AD53E">
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<mods:title id="F88325AEAE17759D6F2D883812DC1064">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="8C5FDF004646FBE3CEC75B4B8C1942B8" type="personal">
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<mods:namePart id="D0673DD5567A5CE5986FF469AD27FBBE">Pyron, Robert Alexander</mods:namePart>
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<mods:name id="D99AC80F637C7FE195526A21D644FDBF" type="personal">
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<mods:roleTerm id="E1BB480E194F090173A3C43AD778BA46">Author</mods:roleTerm>
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<mods:namePart id="CD61821B45CC77812BB7819E8B27FF1E">Wallach, Van</mods:namePart>
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<mods:typeOfResource id="8ECEB0A684FCF25D24BA62A78024EF0E">text</mods:typeOfResource>
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<mods:title id="908206E4BAF94449091D70107F2AA8C9">Zootaxa</mods:title>
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<mods:part id="D2A9811FBF25C2AC52D978629B7FCC53">
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<mods:date id="CDE544A021C75BC2905E218C74B4FD4F">2014</mods:date>
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<mods:number id="6190AEE90731D98CE25D3CD64D949D35">3829</mods:number>
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<mods:identifier id="ECC86BBCAAF5005A93ED212E0F643441" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<treatment id="03B587DDC10CB17BCFD7CD00FEE0FD6E" ID-DOI="http://doi.org/10.5281/zenodo.6127996" ID-GBIF-Taxon="127668692" ID-Zenodo-Dep="6127996" LSID="urn:lsid:plazi:treatment:03B587DDC10CB17BCFD7CD00FEE0FD6E" httpUri="http://treatment.plazi.org/id/03B587DDC10CB17BCFD7CD00FEE0FD6E" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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<subSubSection id="C3066540C10CB17ACFD7CD00FC3DF88A" box="[151,900,1806,1832]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="8BA336CBC10CB17ACFD7CD00FC3DF88A" blockId="54.[151,900,1806,1832]" box="[151,900,1806,1832]" pageId="54" pageNumber="55">
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<heading id="D0EB81A7C10CB17ACFD7CD00FC3DF88A" bold="true" box="[151,900,1806,1832]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
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<taxonomicName id="4C1C4D48C10CB17ACFD7CD00FC3DF88A" authority="Hedges, Marion, Lipp, Marin & Vidal, 2014" authorityName="Hedges, Marion, Lipp, Marin & Vidal" authorityYear="2014" box="[151,900,1806,1832]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10CB17ACFD7CD00FC3DF88A" bold="true" box="[151,900,1806,1832]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACFD7CD00FEEEF88A" bold="true" box="[151,343,1806,1832]" italics="true" pageId="54" pageNumber="55">Malayotyphlops</emphasis>
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<bibRefCitation id="EF8D4B3AC10CB17ACE1ECD00FC3DF88A" author="Hedges" box="[350,900,1806,1832]" pageId="54" pageNumber="55" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article" year="2014">Hedges, Marion, Lipp, Marin & Vidal, 2014</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACFD7CD5BFC65F814" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="8BA336CBC10CB17ACFD7CD5BFD17F8CC" blockId="54.[151,1437,1877,2010]" box="[151,686,1877,1902]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACFD7CD5BFE8BF8CC" bold="true" box="[151,306,1877,1902]" pageId="54" pageNumber="55">
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<typeStatus id="54A78869C10CB17ACFD7CD5BFF6BF8CC" box="[151,210,1877,1902]" pageId="54" pageNumber="55">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C10CB17ACE79CD5BFD17F8CC" authority="Taylor, 1919" authorityName="Taylor" authorityYear="1919" box="[313,686,1877,1902]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="luzonensis">
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<emphasis id="B968EAD9C10CB17ACE79CD5BFDA0F8CC" box="[313,537,1877,1902]" italics="true" pageId="54" pageNumber="55">Typhlops luzonensis</emphasis>
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<bibRefCitation id="EF8D4B3AC10CB17ACD60CD5BFD17F8CC" author="Taylor" box="[544,686,1877,1902]" pageId="54" pageNumber="55" refString="Taylor, E. H. (1919) New or rare Philippine reptiles. The Philippine Journal of Science, Manila, 14 D, 105 - 125." type="journal article" year="1919">Taylor, 1919</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC10CB17ACF87CD77FC65F814" blockId="54.[151,1437,1877,2010]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87CD77FE29F830" bold="true" box="[199,400,1913,1938]" pageId="54" pageNumber="55">Species content.</emphasis>
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<emphasis id="B968EAD9C10CB17ACEE1CD77FD52F817" italics="true" pageId="54" pageNumber="55">
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<taxonomicName id="4C1C4D48C10CB17ACEE1CD77FCF0F830" authority="Mal." authorityName="Mal." box="[417,841,1913,1938]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="canlaonensis">Malayotyphlops canlaonensis, Mal.</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACC1BCD74FB98F830" authority="Mal." authorityName="Mal." box="[859,1057,1913,1938]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="castanotus">castanotus, Mal.</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACB73CD77FB6EF830" authority="Mal." authorityName="Mal." box="[1075,1239,1913,1938]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="collaris">collaris, Mal.</taxonomicName>
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hypogius, Mal.
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<taxonomicName id="4C1C4D48C10CB17ACFD7CD93FEF1F814" authority="Mal." authorityName="Mal." box="[151,328,1949,1974]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="koekkoeki">koekkoeki, Mal.</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACE0ECD93FE65F814" authority="Mal." authorityName="Mal." box="[334,476,1949,1974]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="kraalii">
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kraalii,
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<emphasis id="B968EAD9C10CB17ACEE7CD93FE65F814" bold="true" box="[423,476,1949,1974]" italics="true" pageId="54" pageNumber="55">Mal.</emphasis>
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</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACEA3CD93FD19F814" authority="Mal." authorityName="Mal." box="[483,672,1949,1974]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="luzonensis">
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luzonensis,
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<emphasis id="B968EAD9C10CB17ACD2BCD93FD19F814" bold="true" box="[619,672,1949,1974]" italics="true" pageId="54" pageNumber="55">Mal.</emphasis>
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</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACDE8CD93FD5DF814" box="[680,740,1949,1974]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ruber">ruber</taxonomicName>
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,
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</emphasis>
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and
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<emphasis id="B968EAD9C10CB17ACC62CD93FC65F814" box="[802,988,1949,1974]" italics="true" pageId="54" pageNumber="55">
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Mal.
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<taxonomicName id="4C1C4D48C10CB17ACC1DCD93FC60F814" box="[861,985,1949,1974]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ruficaudus">ruficaudus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17BCF87CDCFFE6FFDC2" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="diagnosis">
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<paragraph id="8BA336CBC10CB17BCF87CDCFFD3AFE57" blockId="54.[151,1437,1877,2010]" lastBlockId="55.[151,1436,151,717]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87CDCFFEF8F878" bold="true" box="[199,321,1985,2010]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C10CB17ACE0DCDCFFDB8F878" box="[333,513,1985,2010]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10CB17ACE0DCDCFFDB8F878" box="[333,513,1985,2010]" italics="true" pageId="54" pageNumber="55">Malayotyphlops</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by a T-III
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<collectingCountry id="F30B765BC10CB17ACB8DCDCCFB4CF878" box="[1229,1269,1986,2010]" name="Cyprus" pageId="54" pageNumber="55">SIP</collectingCountry>
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and eversible hemipenis lacking retrocloacal sacs, and by the following combination of characters: small- to moderate-sized (total length
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<quantity id="4CE49B2EC10DB17BCE6DCAB2FE70FF76" box="[301,457,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" metricValueMax="4.45" metricValueMin="1.09" pageId="55" pageNumber="56" unit="mm" value="277.0" valueMax="445.0" valueMin="109.0">109–445 mm</quantity>
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), moderate to stout-bodied (length/width ratio 26–58) snakes with 20–30 scale rows (with reduction), 280–460 total middorsals, short to moderate tail (1.4–3.6% total length) with 7–18 subcaudals (length/width ratio 0.8–1.6), and apical spine small. Dorsal and lateral head profiles rounded, narrow to moderate rostral (0.31–0.61 head width), inferior nasal suture in contact with first or second supralabial, preocular in contact with second and third supralabials, eye small with distinct pupil, and postoculars 2–4. Lateral tongue papillae absent; left lung absent, tracheal, cardiac and right lungs multicameral (with 8–29 + 2–9 + 4–10 chambers); testes unsegmented; and rectal caecum usually absent (but 0.5–8.2% SVL in
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<emphasis id="B968EAD9C10DB17BCCF2CB9AFBE5FE0F" box="[946,1116,404,429]" italics="true" pageId="55" pageNumber="56">
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Mal.
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<taxonomicName id="4C1C4D48C10DB17BCCAECB9AFBE5FE0F" box="[1006,1116,404,429]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="koekkoeki">koekkoeki</taxonomicName>
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</emphasis>
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and
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<emphasis id="B968EAD9C10DB17BCBD3CB9AFAB5FE0F" box="[1171,1292,404,429]" italics="true" pageId="55" pageNumber="56">
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Mal.
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<taxonomicName id="4C1C4D48C10DB17BCB8ECB9AFAB5FE0F" box="[1230,1292,404,429]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="ruber">ruber</taxonomicName>
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</emphasis>
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). Coloration of dorsum brown, reddish-brown, or blackish-brown, venter yellow, gold or orangish-brown; a light nuchal collar may be present as well as light supralabials.
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</paragraph>
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<paragraph id="8BA336CBC10DB17BCF87CBF1FE6FFDC2" blockId="55.[151,1436,151,717]" pageId="55" pageNumber="56">
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<emphasis id="B968EAD9C10DB17BCF87CBF1FE52FDBA" bold="true" box="[199,491,511,536]" pageId="55" pageNumber="56">Phylogenetic definition.</emphasis>
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Includes the MRCA of
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<taxonomicName id="4C1C4D48C10DB17BCC6CCBF1FB89FDBA" box="[812,1072,511,536]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="ruber">
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<emphasis id="B968EAD9C10DB17BCC6CCBF1FB89FDBA" box="[812,1072,511,536]" italics="true" pageId="55" pageNumber="56">Malayotyphlops ruber</emphasis>
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</taxonomicName>
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and
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<emphasis id="B968EAD9C10DB17BCB3BCBF1FA8CFDBA" box="[1147,1333,511,536]" italics="true" pageId="55" pageNumber="56">
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Mal.
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<taxonomicName id="4C1C4D48C10DB17BCB80CBF1FA8CFDBA" box="[1216,1333,511,536]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="luzonensis">luzonensis</taxonomicName>
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</emphasis>
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and all descendants thereof, and all species more closely related to
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<emphasis id="B968EAD9C10DB17BCC0EC82AFC73FD9F" box="[846,970,548,573]" italics="true" pageId="55" pageNumber="56">
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Mal.
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<taxonomicName id="4C1C4D48C10DB17BCCCCC82AFC73FD9F" box="[908,970,548,573]" class="Reptilia" family="Typhlopidae" genus="Malayotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="ruber">ruber</taxonomicName>
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</emphasis>
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than to the
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<typeStatus id="54A78869C10DB17BCB1AC82BFB33FD9F" box="[1114,1162,549,573]" pageId="55" pageNumber="56">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10DB17BCF87C862FBF1FD27" box="[199,1096,620,645]" pageId="55" pageNumber="56" type="etymology">
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<paragraph id="8BA336CBC10DB17BCF87C862FBF1FD27" blockId="55.[151,1436,151,717]" box="[199,1096,620,645]" pageId="55" pageNumber="56">
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<emphasis id="B968EAD9C10DB17BCF87C862FEF7FD27" bold="true" box="[199,334,620,645]" pageId="55" pageNumber="56">Etymology.</emphasis>
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Name refers to the distribution of species in the Malay archipelago.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10DB17BCF87C881FEE0FD6E" pageId="55" pageNumber="56" type="distribution">
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<paragraph id="8BA336CBC10DB17BCF87C881FEE0FD6E" blockId="55.[151,1436,151,717]" pageId="55" pageNumber="56">
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<emphasis id="B968EAD9C10DB17BCF87C881FED9FD0A" bold="true" box="[199,352,655,680]" pageId="55" pageNumber="56">Distribution.</emphasis>
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Species are primarily found in the
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<collectingCountry id="F30B765BC10DB17BCDB8C881FCC0FD0A" box="[760,889,655,680]" name="Philippines" pageId="55" pageNumber="56">Philippines</collectingCountry>
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, though some occur on Borneo and the
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<collectingCountry id="F30B765BC10DB17BCA05C881FF5FFD6F" name="Indonesia" pageId="55" pageNumber="56">Maluku Islands</collectingCountry>
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(Table 1).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |