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<document id="6C0553B29D62567082FDCAB47B5B4C96" ID-CLB-Dataset="22833" ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee &amp; Nützel, Al- Exander" docDate="2020" docId="03B587AB4F2A1560FF517B17FACCFF7E" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trachydomia suwanneeae Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="30" updateTime="1698836971451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="579387B42239B5E60C06CA3E588D3BFD">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="0371C293C752D94226A3A17369B22EBE">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="B247107C5A75DA2F837DC30E590BEDDD">Karapunar, Baran</mods:namePart>
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<mods:namePart id="3D462E339831C37B20B916B7EEABB141">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="23A16222E382FC6308803579A5BCB3F6">Nützel, Al- Exander</mods:namePart>
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<heading id="D0EB81D14F2A1563FF517B17FDF3FC06" bold="true" box="[151,568,873,900]" fontSize="10" level="2" pageId="29" pageNumber="30" reason="2">
<emphasis id="B968EAAF4F2A1563FF517B17FDF3FC06" bold="true" box="[151,568,873,900]" pageId="29" pageNumber="30">
<taxonomicName id="4C1C4D3E4F2A1563FF517B17FE18FC06" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[151,467,873,900]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="suwanneeae" status="sp. nov.">
<emphasis id="B968EAAF4F2A1563FF517B17FE18FC06" bold="true" box="[151,467,873,900]" italics="true" pageId="29" pageNumber="30">Trachydomia suwanneeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F2A1563FE1C7B17FDF3FC06" box="[474,568,873,899]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C30665364F2A1563FF517BF3FEFBFC22" box="[151,304,909,935]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BA336BD4F2A1563FF517BF3FEFBFC22" blockId="29.[151,568,873,935]" box="[151,304,909,935]" pageId="29" pageNumber="30">
(
<figureCitation id="13272A384F2A1563FF667BF3FF25FC22" box="[160,238,909,935]" captionStart="FIGURE 21" captionStartId="30.[151,250,1173,1198]" captionTargetBox="[166,1421,273,1148]" captionTargetId="figure@30.[166,1421,273,1148]" captionTargetPageId="30" captionText="FIGURE 21. AD, Trachydomia suwanneeae sp. nov., ESKU-19-LP 26, holotype. EF, Trachydomia cf. nodusum (Meek &amp; Worthen, 1866), ESKU-19-LP 92. GK, Trachyspira eleganta sp. nov., ESKU-19-LP 28, holotype. Scale bars represent: 2 mm (AD, GK); 1 mm (EF)." figureDoi="http://doi.org/10.5281/zenodo.3763949" httpUri="https://zenodo.org/record/3763949/files/figure.png" pageId="29" pageNumber="30">Fig. 21</figureCitation>
AD)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F2A1563FF517BABFBB7FC6A" box="[151,1148,981,1007]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BA336BD4F2A1563FF517BABFBB7FC6A" blockId="29.[151,1437,981,2015]" box="[151,1148,981,1007]" pageId="29" pageNumber="30">
<emphasis id="B968EAAF4F2A1563FF517BABFED5FC6A" bold="true" box="[151,286,981,1007]" pageId="29" pageNumber="30">Etymology.</emphasis>
In honor of Suwannee Phomprasith for her work in biodiversity in
<collectingCountry id="F30B762D4F2A1563FBD77BABFBB3FC6A" box="[1041,1144,981,1007]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F2A1563FF017B87FCF5FBFA" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BA336BD4F2A1563FF017B87FDCFFB96" blockId="29.[151,1437,981,2015]" box="[199,516,1017,1043]" pageId="29" pageNumber="30">
<materialsCitation id="3B743CE04F2A1563FF017B87FDCFFB96" ID-GBIF-Occurrence="2608527132" box="[199,516,1017,1043]" collectionCode="LP" pageId="29" pageNumber="30" specimenCode="ESKU-19-LP 26" specimenCount="1" typeStatus="holotype">
<emphasis id="B968EAAF4F2A1563FF017B87FEF0FB96" bold="true" box="[199,315,1017,1043]" pageId="29" pageNumber="30">
<typeStatus id="54A7881F4F2A1563FF017B87FEFDFB96" box="[199,310,1017,1043]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBBA9EC64F2A1563FE847B87FE34FB96" box="[322,511,1017,1043]" pageId="29" pageNumber="30">ESKU-19-LP 26</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA336BD4F2A1563FF017C63FDF2FBB2" blockId="29.[151,1437,981,2015]" box="[199,569,1053,1079]" pageId="29" pageNumber="30">
<emphasis id="B968EAAF4F2A1563FF017C63FE8DFBB2" bold="true" box="[199,326,1053,1079]" pageId="29" pageNumber="30">
<typeStatus id="54A7881F4F2A1563FF017C63FE8AFBB2" box="[199,321,1053,1079]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
.
</emphasis>
ESKU-19-
<collectionCode id="ED0DAE784F2A1563FE047C63FE2FFBB2" box="[450,484,1053,1079]" country="Russia" name="Laboratory of Palaeontology" pageId="29" pageNumber="30">LP</collectionCode>
27, 94.
</paragraph>
<paragraph id="8BA336BD4F2A1563FF017C3FFCF5FBFA" blockId="29.[151,1437,981,2015]" pageId="29" pageNumber="30">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F2A1563FDD17C3FFDF2FBDE" box="[535,569,1089,1115]" country="Russia" name="Laboratory of Palaeontology" pageId="29" pageNumber="30">LP</collectionCode>
26: height = 12.1; width = 9.2; apical angle = 82
<superScript id="7C699BF54F2A1563FBA47C3EFBACFBCB" attach="none" box="[1122,1127,1088,1102]" fontSize="6" pageId="29" pageNumber="30">º</superScript>
. ESKU-19-
<collectionCode id="ED0DAE784F2A1563FB2D7C3FFAC6FBDE" box="[1259,1293,1089,1115]" country="Russia" name="Laboratory of Palaeontology" pageId="29" pageNumber="30">LP</collectionCode>
27: height = 6.4; width = 5.7. ESKU-19-
<collectionCode id="ED0DAE784F2A1563FE0B7C1BFE24FBFA" box="[461,495,1125,1151]" country="Russia" name="Laboratory of Palaeontology" pageId="29" pageNumber="30">LP</collectionCode>
94: height = 5.4; width = 4.2.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F2A1563FF017CF7FE74FB6E" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BA336BD4F2A1563FF017CF7FE74FB6E" blockId="29.[151,1437,981,2015]" pageId="29" pageNumber="30">
<emphasis id="B968EAAF4F2A1563FF017CF7FD6AFB26" bold="true" box="[199,673,1160,1187]" pageId="29" pageNumber="30">Type locality and stratigraphical range.</emphasis>
Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
<quantity id="4CE49B584F2A1563FEF17CD3FEB4FB42" box="[311,383,1197,1223]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="km" value="13.0">13 km</quantity>
east of
<collectingRegion id="49D8F85F4F2A1563FE1C7CD3FD69FB42" box="[474,674,1197,1223]" country="Thailand" name="Lop Buri" pageId="29" pageNumber="30">Lopburi Province</collectingRegion>
, Central
<collectingCountry id="F30B762D4F2A1563FCCF7CD3FCA6FB42" box="[777,877,1197,1223]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
(
<figureCitation id="13272A384F2A1563FCBB7CD3FC0BFB42" box="[893,960,1197,1223]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="29" pageNumber="30">Fig. 1</figureCitation>
), Khao Khad Formation,
<collectingRegion id="49D8F85F4F2A1563FB217CD3FA8DFB42" box="[1255,1350,1197,1223]" country="Thailand" name="Sara Buri" pageId="29" pageNumber="30">Saraburi</collectingRegion>
Group, Middle Permian, Roadian.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F2A1563FF017C8AFB67FA46" pageId="29" pageNumber="30" type="description">
<paragraph id="8BA336BD4F2A1563FF017C8AFB67FA46" blockId="29.[151,1437,981,2015]" pageId="29" pageNumber="30">
<emphasis id="B968EAAF4F2A1563FF017C8AFE92FA8A" bold="true" box="[199,345,1268,1295]" pageId="29" pageNumber="30">Description.</emphasis>
Shell naticiform to turbiniform; spire acute; whorls round, convex; body whorl distinctly higher than spire; whorls ornamented with distinct small nodes which are especially well-developed on the body whorl and weak or absent on the earliest preserved whorls; nodes roughly arranged in opisthocline arranged nodes; distance between nodes exceeds diameter of nodes; suture distinct; whorls convex with periphery at mid-whorl of body whorl but below mid-whorl of whorl face of spire whorls, with narrow subsutural ramp; base convex with gradually embracing of body whorl at transition below the periphery; aperture unknown; anomphalous.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F2A1560FF017DB3FACCFF7E" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BA336BD4F2A1560FF017DB3FACCFF7E" blockId="29.[151,1437,981,2015]" lastBlockId="30.[151,1436,152,251]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B968EAAF4F2A1563FF017DB3FEF0FA62" bold="true" box="[199,315,1485,1511]" pageId="29" pageNumber="30">Remarks.</emphasis>
Based on the shell characters of the studied specimens such as having a naticiform shell and pustules on whorl face, the present material represents undoubtedly the genus
<taxonomicName id="4C1C4D3E4F2A1563FC6A7D8FFB8BF98E" authorityName="Meek &amp; Worthen" authorityYear="1866" box="[940,1088,1521,1547]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F2A1563FC6A7D8FFB8BF98E" box="[940,1088,1521,1547]" italics="true" pageId="29" pageNumber="30">Trachydomia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1C4D3E4F2A1563FB897D8FFABFF98E" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[1103,1396,1521,1547]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="suwanneeae" status="sp. nov.">
<emphasis id="B968EAAF4F2A1563FB897D8FFABFF98E" box="[1103,1396,1521,1547]" italics="true" pageId="29" pageNumber="30">Trachydomia suwanneeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F2A1563FABA7D8FFF0CF9AA" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
resembles most closely
<taxonomicName id="4C1C4D3E4F2A1563FE257E68FCEAF9AA" authority="Mansuy, 1913 a" authorityName="Mansuy" authorityYear="1913" box="[483,801,1557,1583]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="dussaulti">
<emphasis id="B968EAAF4F2A1563FE257E68FE38F9AA" box="[483,499,1558,1583]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FDC57E6BFDA3F9AA" box="[515,616,1557,1583]" italics="true" pageId="29" pageNumber="30">dussaulti</emphasis>
<bibRefCitation id="EF8D4B4C4F2A1563FDB47E6BFCEAF9AA" author="Mansuy" box="[626,801,1557,1583]" firstAuthor="Mansuy" pageId="29" pageNumber="30" pagination="1 - 133" refId="ref26022" refString="Mansuy, H. (1913 a) Faunes des calcaires a Productus de l'Indochine. Memoires du Service Geologique de l'Indochine, 2 (4), 1 - 133." type="journal article" year="1913" yearSuffix="a">
Mansuy, 1913
<emphasis id="B968EAAF4F2A1563FCD57E68FCEAF9AA" box="[787,801,1558,1583]" italics="true" pageId="29" pageNumber="30">a</emphasis>
</bibRefCitation>
</taxonomicName>
(p. 101, pl. 11, fig. 5ab) from the Permian Productus Limestone of
<collectingCountry id="F30B762D4F2A1563FEF47E47FEA3F9D6" box="[306,360,1593,1619]" name="Laos" pageId="29" pageNumber="30">Laos</collectingCountry>
in shape and ornaments that was also reported by
<bibRefCitation id="EF8D4B4C4F2A1563FC5D7E47FBE4F9D6" author="Batten" box="[923,1071,1593,1619]" firstAuthor="Batten" pageId="29" pageNumber="30" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
, p. 17, fig. 20) from the Permian of
<collectingRegion id="49D8F85F4F2A1563FF707E23FF32F9F2" box="[182,249,1629,1655]" country="Malaysia" name="Perak" pageId="29" pageNumber="30">Perak</collectingRegion>
,
<collectingCountry id="F30B762D4F2A1563FEC37E23FEA7F9F2" box="[261,364,1629,1655]" name="Malaysia" pageId="29" pageNumber="30">Malaysia</collectingCountry>
and by
<bibRefCitation id="EF8D4B4C4F2A1563FE017E23FDA8F9F2" author="Delpey" box="[455,611,1629,1655]" firstAuthor="Delpey" pageId="29" pageNumber="30" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey (1941</bibRefCitation>
, p. 268, fig. 10) from the Permian of
<collectingCountry id="F30B762D4F2A1563FBCD7E23FB49F9F2" box="[1035,1154,1629,1655]" name="Cambodia" pageId="29" pageNumber="30">Cambodia</collectingCountry>
. However,
<taxonomicName id="4C1C4D3E4F2A1563FAC77E20FA49F9F2" authorityName="Mansuy" authorityYear="1913" box="[1281,1410,1629,1655]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="dussaulti">
<emphasis id="B968EAAF4F2A1563FAC77E20FA49F9F2" box="[1281,1410,1629,1655]" italics="true" pageId="29" pageNumber="30">T. dussaulti</emphasis>
</taxonomicName>
is not as high-spired and hence its spire is not as acute.
<taxonomicName id="4C1C4D3E4F2A1563FD337EFCFCBDF91E" authorityName="Mansuy" authorityYear="1913" box="[757,886,1665,1691]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="dussaulti">
<emphasis id="B968EAAF4F2A1563FD337EFCFCBDF91E" box="[757,886,1665,1691]" italics="true" pageId="29" pageNumber="30">T. dussaulti</emphasis>
</taxonomicName>
has more nodes and the nodes are more densely spaced. The specimen illustrated as
<taxonomicName id="4C1C4D3E4F2A1563FDE47ED8FC98F93A" authority="Mansuy, 1913 a" authorityName="Mansuy" authorityYear="1913" box="[546,851,1701,1727]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="dussaulti">
<emphasis id="B968EAAF4F2A1563FDE47ED8FDF9F93A" box="[546,562,1702,1727]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FDF87EDBFD68F93A" box="[574,675,1701,1727]" italics="true" pageId="29" pageNumber="30">dussaulti</emphasis>
<bibRefCitation id="EF8D4B4C4F2A1563FD6E7EDBFC98F93A" author="Mansuy" box="[680,851,1701,1727]" firstAuthor="Mansuy" pageId="29" pageNumber="30" pagination="1 - 133" refId="ref26022" refString="Mansuy, H. (1913 a) Faunes des calcaires a Productus de l'Indochine. Memoires du Service Geologique de l'Indochine, 2 (4), 1 - 133." type="journal article" year="1913" yearSuffix="a">
Mansuy, 1913
<emphasis id="B968EAAF4F2A1563FC837ED8FC98F93A" box="[837,851,1702,1727]" italics="true" pageId="29" pageNumber="30">a</emphasis>
</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4C4F2A1563FCBC7EDBFBD1F93A" author="Delpey" box="[890,1050,1701,1727]" firstAuthor="Delpey" pageId="29" pageNumber="30" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey (1941)</bibRefCitation>
has a more distinctly inflated body whorl, a blunter spire and finer nodes, whereas the specimens assigned to
<taxonomicName id="4C1C4D3E4F2A1563FC127EB4FAB2F961" authority="(Meek &amp; Worthen, 1866)" baseAuthorityName="Meek &amp; Worthen" baseAuthorityYear="1866" box="[980,1401,1737,1764]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="nodusum">
<emphasis id="B968EAAF4F2A1563FC127EB4FC2FF966" box="[980,996,1738,1763]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FC347EB7FB9CF966" box="[1010,1111,1737,1763]" italics="true" pageId="29" pageNumber="30">nodusum</emphasis>
(
<bibRefCitation id="EF8D4B4C4F2A1563FBA37EB7FABAF961" author="Meek &amp; Worthen" box="[1125,1393,1737,1764]" firstAuthor="Meek" pageId="29" pageNumber="30" pagination="143 - 411" refId="ref26392" refString="Meek, F. B. &amp; Worthen, A. H. (1866) Descriptions of invertebrates from the Carboniferous system. Geological Survey of Illinois, 2, Palaeontology, 143 - 411." type="journal article" year="1866">Meek &amp; Worthen, 1866</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EF8D4B4C4F2A1563FF517E93FEF9F882" author="Delpey" box="[151,306,1773,1799]" firstAuthor="Delpey" pageId="29" pageNumber="30" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey (1941</bibRefCitation>
, p. 268, fig. 9) is more similar to the present specimens in having an acute spire and a rather elongated shape.
<bibRefCitation id="EF8D4B4C4F2A1563FF217F6FFE5EF8AE" author="Knight" box="[231,405,1809,1835]" firstAuthor="Knight" pageId="29" pageNumber="30" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
Knight (1933
<emphasis id="B968EAAF4F2A1563FEB87F6FFE47F8AE" box="[382,396,1809,1835]" italics="true" pageId="29" pageNumber="30">b</emphasis>
)
</bibRefCitation>
has reported several specimens of
<taxonomicName id="4C1C4D3E4F2A1563FCE17F6CFC60F8AE" baseAuthorityName="Meek &amp; Worthen" baseAuthorityYear="1866" box="[807,939,1809,1835]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="nodusum">
<emphasis id="B968EAAF4F2A1563FCE17F6CFCFCF8AE" box="[807,823,1810,1835]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FC807F6FFC60F8AE" box="[838,939,1809,1835]" italics="true" pageId="29" pageNumber="30">nodusum</emphasis>
</taxonomicName>
from the Pennsylvanian,
<collectingCountry id="F30B762D4F2A1563FB167F6FFACDF8AE" box="[1232,1286,1809,1835]" name="United States of America" pageId="29" pageNumber="30">USA</collectingCountry>
(from which this species was originally described) differs distinctly from the studied specimens in having a lower spire, a broader and more pronounced ramp, stronger and coarser nodes and more inflated body whorl.
<taxonomicName id="4C1C4D3E4F2A1563FBA37F24FAC7F8F6" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[1125,1292,1881,1907]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="suwanneeae" status="sp. nov.">
<emphasis id="B968EAAF4F2A1563FBA37F24FBBEF8F6" box="[1125,1141,1882,1907]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FB457F24FAC7F8F6" box="[1155,1292,1882,1907]" italics="true" pageId="29" pageNumber="30">suwanneeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F2A1563FAD47F27FAA2F8F6" box="[1298,1385,1881,1907]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
also resembles
<taxonomicName id="4C1C4D3E4F2A1563FEC97F00FDD5F812" authority="Knight, 1933 b" authorityName="Knight" authorityYear="1933" box="[271,542,1917,1943]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="whitei">
<emphasis id="B968EAAF4F2A1563FEC97F00FED4F812" box="[271,287,1918,1943]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FEE87F03FEB9F812" box="[302,370,1917,1943]" italics="true" pageId="29" pageNumber="30">whitei</emphasis>
<bibRefCitation id="EF8D4B4C4F2A1563FEBD7F03FDD5F812" author="Knight" box="[379,542,1917,1943]" firstAuthor="Knight" pageId="29" pageNumber="30" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
Knight, 1933
<emphasis id="B968EAAF4F2A1563FDD67F03FDD5F812" box="[528,542,1917,1943]" italics="true" pageId="29" pageNumber="30">b</emphasis>
</bibRefCitation>
</taxonomicName>
, both in having small and widely spaced nodes and a similar whorl profile but
<taxonomicName id="4C1C4D3E4F2A1563FF517FDCFF30F83E" authorityName="Knight" authorityYear="1933" box="[151,251,1953,1979]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="whitei">
<emphasis id="B968EAAF4F2A1563FF517FDCFF6CF83E" box="[151,167,1954,1979]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FF717FDFFF30F83E" box="[183,251,1953,1979]" italics="true" pageId="29" pageNumber="30">whitei</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C1C4D3E4F2A1563FE517FDCFD8BF83E" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[407,576,1953,1979]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="suwanneeae" status="sp. nov.">
<emphasis id="B968EAAF4F2A1563FE517FDCFE6CF83E" box="[407,423,1954,1979]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FE717FDCFD8BF83E" box="[439,576,1954,1979]" italics="true" pageId="29" pageNumber="30">suwanneeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F2A1563FD8F7FDFFD69F83E" box="[585,674,1953,1979]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
by its wider, more pronounced subsutural ramp, by being broader and by having stronger nodes.
<taxonomicName id="4C1C4D3E4F2A1563FE287FB8FD5EF85A" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[494,661,1989,2015]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="suwanneeae" status="sp. nov.">
<emphasis id="B968EAAF4F2A1563FE287FB8FE35F85A" box="[494,510,1990,2015]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FDCA7FB8FD5EF85A" box="[524,661,1990,2015]" italics="true" pageId="29" pageNumber="30">suwanneeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F2A1563FD5D7FBBFD39F85A" box="[667,754,1989,2015]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
is also similar to
<taxonomicName id="4C1C4D3E4F2A1563FC717FB8FA57F85A" authority="Nutzel &amp; Ketwetsuriya, 2016" authorityName="Nutzel &amp; Ketwetsuriya" authorityYear="2016" box="[951,1436,1989,2015]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="takhliensis">
<emphasis id="B968EAAF4F2A1563FC717FB8FC0CF85A" box="[951,967,1990,2015]" italics="true" pageId="29" pageNumber="30">T</emphasis>
.
<emphasis id="B968EAAF4F2A1563FC127FBBFB87F85A" box="[980,1100,1989,2015]" italics="true" pageId="29" pageNumber="30">takhliensis</emphasis>
Nützel &amp;
<bibRefCitation id="EF8D4B4C4F2A1563FB077FBBFA57F85A" author="Ketwetsuriya" box="[1217,1436,1989,2015]" firstAuthor="Ketwetsuriya" pageId="29" pageNumber="30" pagination="481 - 513" refId="ref24896" refString="Ketwetsuriya, C., Nu ¨ tzel, A. &amp; Kanjanapayont, P. (2016) Permian gastropods from the Tak Fa Limestone, Nakhon Sawan, Northern Thailand. Bulletin of Geosciences, 91, 481 - 513. https: // doi. org / 10.3140 / bull. geosci. 1617" type="journal article" year="2016">Ketwetsuriya, 2016</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF8D4B4C4F291560FF5978E7FE73FF36" author="Ketwetsuriya" box="[159,440,153,179]" etAl="et al." firstAuthor="Ketwetsuriya" pageId="30" pageNumber="31" pagination="481 - 513" refId="ref24896" refString="Ketwetsuriya, C., Nu ¨ tzel, A. &amp; Kanjanapayont, P. (2016) Permian gastropods from the Tak Fa Limestone, Nakhon Sawan, Northern Thailand. Bulletin of Geosciences, 91, 481 - 513. https: // doi. org / 10.3140 / bull. geosci. 1617" type="journal article" year="2016">
Ketwetsuriya
<emphasis id="B968EAAF4F291560FEFB78E4FEA4FF36" box="[317,367,153,179]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 2016
</bibRefCitation>
, p. 502, fig. 19JM) from the Middle Permian of the
<collectingRegion id="49D8F85F4F291560FBD478E7FBF7FF36" box="[1042,1084,153,179]" country="Thailand" name="Tak" pageId="30" pageNumber="31">Tak</collectingRegion>
Fa Limestone,
<collectingCountry id="F30B762D4F291560FB2C78E7FA9AFF36" box="[1258,1361,153,179]" name="Thailand" pageId="30" pageNumber="31">Thailand</collectingCountry>
. However, the shell of
<taxonomicName id="4C1C4D3E4F291560FE9C78C0FE3BFF52" authorityName="Nutzel &amp; Ketwetsuriya" authorityYear="2016" box="[346,496,189,215]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="takhliensis">
<emphasis id="B968EAAF4F291560FE9C78C0FEA1FF52" box="[346,362,190,215]" italics="true" pageId="30" pageNumber="31">T</emphasis>
.
<emphasis id="B968EAAF4F291560FEBE78C3FE3BFF52" box="[376,496,189,215]" italics="true" pageId="30" pageNumber="31">takhliensis</emphasis>
</taxonomicName>
is broader, its spire is less acute and it has more inflated body whorl.
<taxonomicName id="4C1C4D3E4F291560FAC078C0FA57FF52" authorityName="Nutzel &amp; Ketwetsuriya" authorityYear="2016" box="[1286,1436,189,215]" class="Gastropoda" family="Trachyspiridae" genus="Trachydomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="takhliensis">
<emphasis id="B968EAAF4F291560FAC078C0FADDFF52" box="[1286,1302,190,215]" italics="true" pageId="30" pageNumber="31">T</emphasis>
.
<emphasis id="B968EAAF4F291560FAE278C3FA57FF52" box="[1316,1436,189,215]" italics="true" pageId="30" pageNumber="31">takhliensis</emphasis>
</taxonomicName>
has more densely spaced, somewhat coarser and more protruded nodes and a more pronounced ramp.
</paragraph>
</subSubSection>
</treatment>
</document>