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<document id="124200534E6F4C86EB85C902854B3B37" ID-CLB-Dataset="22833" ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee &amp; Nützel, Al- Exander" docDate="2020" docId="03B587AB4F16155FFF5178E7FBD7FCB9" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterosubulites longusapertura Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="34" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="34" updateTime="1698836971451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="088E33606E8E589052AF5152AC6684C9">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="C3A9DEF74856AED0B8E5260AA265BB47">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="1C3C654C2693F0981624B20D1360DB78">Karapunar, Baran</mods:namePart>
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<mods:namePart id="168FA3824F7926015DDFA74D8C12DFEC">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="AB200487CB00CE570D050B7F48DC4958">Nützel, Al- Exander</mods:namePart>
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<treatment id="03B587AB4F16155FFF5178E7FBD7FCB9" ID-DOI="http://doi.org/10.5281/zenodo.3803831" ID-GBIF-Taxon="163737165" ID-Zenodo-Dep="3803831" LSID="urn:lsid:plazi:treatment:03B587AB4F16155FFF5178E7FBD7FCB9" httpUri="http://treatment.plazi.org/id/03B587AB4F16155FFF5178E7FBD7FCB9" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="C30665364F16155FFF5178E7FDB7FF31" box="[151,636,153,180]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BA336BD4F16155FFF5178E7FDB7FF31" blockId="33.[151,636,153,215]" box="[151,636,153,180]" pageId="33" pageNumber="34">
<heading id="D0EB81D14F16155FFF5178E7FDB7FF31" bold="true" box="[151,636,153,180]" fontSize="10" level="2" pageId="33" pageNumber="34" reason="2">
<emphasis id="B968EAAF4F16155FFF5178E7FDB7FF31" bold="true" box="[151,636,153,180]" pageId="33" pageNumber="34">
<taxonomicName id="4C1C4D3E4F16155FFF5178E7FDDCFF31" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[151,535,153,180]" class="Gastropoda" family="Heterosubulitidae" genus="Heterosubulites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="longusapertura" status="sp. nov.">
<emphasis id="B968EAAF4F16155FFF5178E7FDDCFF31" bold="true" box="[151,535,153,180]" italics="true" pageId="33" pageNumber="34">Heterosubulites longusapertura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F16155FFDD878E4FDB7FF31" box="[542,636,154,180]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF5178C3FEC5FF52" box="[151,270,189,215]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BA336BD4F16155FFF5178C3FEC5FF52" blockId="33.[151,636,153,215]" box="[151,270,189,215]" pageId="33" pageNumber="34">
(
<figureCitation id="13272A384F16155FFF6678C3FECDFF52" box="[160,262,189,215]" captionStart="FIGURE 22" captionStartId="33.[151,250,1781,1806]" captionTargetBox="[166,1421,849,1755]" captionTargetId="figure@33.[166,1421,849,1755]" captionTargetPageId="33" captionText="FIGURE 22. AC, Strobeus? sp. 1, AB, ESKU-19-LP 49. C, ESKU-19-LP 90. DE, Strobeus? sp. 2, D, ESKU-19-LP 217. E, ESKU-19-LP 220. F, Girtyspira? sp., ESKU-19-LP 222. G, Heterosubulites longusapertura sp. nov. ESKU-19-LP 215, holotype. Scale bars represent: 2 mm (AC); 200 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.3763951" httpUri="https://zenodo.org/record/3763951/files/figure.png" pageId="33" pageNumber="34">Fig. 22G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF517978FD31FEA5" box="[151,762,261,288]" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BA336BD4F16155FFF517978FD31FEA5" blockId="33.[151,1437,261,828]" box="[151,762,261,288]" pageId="33" pageNumber="34">
<emphasis id="B968EAAF4F16155FFF517978FED5FEA5" bold="true" box="[151,286,262,288]" pageId="33" pageNumber="34">Etymology.</emphasis>
Latin, because of the long-shape aperture.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF017954FBA2FEED" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BA336BD4F16155FFF017954FD32FEC1" blockId="33.[151,1437,261,828]" box="[199,761,297,324]" pageId="33" pageNumber="34">
<materialsCitation id="3B743CE04F16155FFF017954FD32FEC1" ID-GBIF-Occurrence="2608527137" box="[199,761,297,324]" collectionCode="LP" pageId="33" pageNumber="34" specimenCode="ESKU-19-LP 215" specimenCount="1" typeStatus="holotype">
<emphasis id="B968EAAF4F16155FFF017954FEF0FEC1" bold="true" box="[199,315,298,324]" pageId="33" pageNumber="34">
<typeStatus id="54A7881F4F16155FFF017954FEFDFEC1" box="[199,310,298,324]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
.
</emphasis>
Only
<specimenCount id="9D1AFD344F16155FFE477954FDD5FEC6" box="[385,542,297,324]" pageId="33" pageNumber="34">one specimen</specimenCount>
:
<specimenCode id="DBBA9EC64F16155FFDEF7957FD3FFEC1" box="[553,756,297,324]" pageId="33" pageNumber="34">ESKU-19-LP 215</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA336BD4F16155FFF017933FBA2FEED" blockId="33.[151,1437,261,828]" box="[199,1129,332,360]" pageId="33" pageNumber="34">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F16155FFDD07933FDF3FEE2" box="[534,568,333,359]" country="Russia" name="Laboratory of Palaeontology" pageId="33" pageNumber="34">LP</collectionCode>
215: height = 1.6; width = 0.9; apical angle = 82
<superScript id="7C699BF54F16155FFB9B7932FBA9FEDF" attach="none" box="[1117,1122,332,346]" fontSize="6" pageId="33" pageNumber="34">º</superScript>
.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF01790CFE74FE56" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BA336BD4F16155FFF01790CFE74FE56" blockId="33.[151,1437,261,828]" pageId="33" pageNumber="34">
<emphasis id="B968EAAF4F16155FFF01790CFD6AFE09" bold="true" box="[199,673,369,396]" pageId="33" pageNumber="34">Type locality and stratigraphical range.</emphasis>
Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
<quantity id="4CE49B584F16155FFEF179EBFEB4FE2A" box="[311,383,405,431]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="34" unit="km" value="13.0">13 km</quantity>
east of
<collectingRegion id="49D8F85F4F16155FFE1C79EBFD69FE2A" box="[474,674,405,431]" country="Thailand" name="Lop Buri" pageId="33" pageNumber="34">Lopburi Province</collectingRegion>
, Central
<collectingCountry id="F30B762D4F16155FFCCF79EBFCA6FE2A" box="[777,877,405,431]" name="Thailand" pageId="33" pageNumber="34">Thailand</collectingCountry>
(
<figureCitation id="13272A384F16155FFCBB79EBFC0BFE2A" box="[893,960,405,431]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="33" pageNumber="34">Fig. 1</figureCitation>
), Khao Khad Formation,
<collectingRegion id="49D8F85F4F16155FFB2179EBFA8DFE2A" box="[1255,1350,405,431]" country="Thailand" name="Sara Buri" pageId="33" pageNumber="34">Saraburi</collectingRegion>
Group, Middle Permian, Roadian.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF0179A3FB89FDE6" pageId="33" pageNumber="34" type="description">
<paragraph id="8BA336BD4F16155FFF0179A3FB89FDE6" blockId="33.[151,1437,261,828]" pageId="33" pageNumber="34">
<emphasis id="B968EAAF4F16155FFF0179A3FE92FE7D" bold="true" box="[199,345,477,504]" pageId="33" pageNumber="34">Description.</emphasis>
Shell very small, fusiform; whorls smooth; whorls embracing high on the whorls; whorls evenly convex with periphery above mid-whorl; spire whorls only slightly convex; spire small, blunt, rounded; body-whorl inflated, much higher than spire with height about 90% of total height; sutures indistinct; base flat; aperture elongated, narrow, with wide and short anterior siphonal canal; outer lip almost straight.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F16155FFF017A10FBD7FCB9" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BA336BD4F16155FFF017A10FBD7FCB9" blockId="33.[151,1437,261,828]" pageId="33" pageNumber="34">
<emphasis id="B968EAAF4F16155FFF017A10FEF0FD0D" bold="true" box="[199,315,622,648]" pageId="33" pageNumber="34">Remarks.</emphasis>
<taxonomicName id="4C1C4D3E4F16155FFE857A13FD63FD02" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[323,680,621,647]" class="Gastropoda" family="Heterosubulitidae" genus="Heterosubulites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="longusapertura" status="sp. nov.">
<emphasis id="B968EAAF4F16155FFE857A13FD63FD02" box="[323,680,621,647]" italics="true" pageId="33" pageNumber="34">Heterosubulites longusapertura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F16155FFD767A10FCCCFD0D" box="[688,775,622,648]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
is very close to the
<typeStatus id="54A7881F4F16155FFC2A7A10FBD7FD0D" box="[1004,1052,622,648]" pageId="33" pageNumber="34">type</typeStatus>
species
<taxonomicName id="4C1C4D3E4F16155FFBBB7A10FB2AFD02" authorityName="Knight" authorityYear="1931" box="[1149,1249,621,648]" class="Gastropoda" family="Acteoninidae" genus="Ceraunocochlis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="blatta">
<emphasis id="B968EAAF4F16155FFBBB7A10FB5AFD02" box="[1149,1169,622,647]" italics="true" pageId="33" pageNumber="34">H</emphasis>
.
<emphasis id="B968EAAF4F16155FFB667A13FB2AFD02" box="[1184,1249,621,647]" italics="true" pageId="33" pageNumber="34">blatta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8D4B4C4F16155FFB377A13FA5EFD02" author="Knight" box="[1265,1429,621,647]" firstAuthor="Knight" pageId="33" pageNumber="34" pagination="177 - 229" refId="ref25046" refString="Knight, J. B. (1931 a) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: the Subulitidae. Journal of Paleontology, 5 (3), 177 - 229." type="journal article" year="1931" yearSuffix="a">
Knight, 1931
<emphasis id="B968EAAF4F16155FFA417A10FA5EFD02" box="[1415,1429,622,647]" italics="true" pageId="33" pageNumber="34">a</emphasis>
</bibRefCitation>
, p. 203204, pl. 21, fig. 2ad) (see also
<bibRefCitation id="EF8D4B4C4F16155FFD8B7AEFFD22FD2E" author="Bandel" box="[589,745,657,683]" firstAuthor="Bandel" pageId="33" pageNumber="34" pagination="45 - 80" refId="ref22773" refString="Bandel, K. (2002 a) About the Heterostropha (Gastropoda) from the Carboniferous and Permian. Mitteilungen des Geologisch- Palaontologischen Instituts der Universitat Hamburg, 86, 45 - 80." type="journal article" year="2002" yearSuffix="a">
Bandel 2002
<emphasis id="B968EAAF4F16155FFD1D7AECFD22FD2E" box="[731,745,658,683]" italics="true" pageId="33" pageNumber="34">a</emphasis>
</bibRefCitation>
) from the Pennsylvanian of
<collectingRegion id="49D8F85F4F16155FFBEE7AEFFB45FD2E" box="[1064,1166,657,683]" country="United States of America" name="Missouri" pageId="33" pageNumber="34">Missouri</collectingRegion>
,
<collectingCountry id="F30B762D4F16155FFB5C7AEFFB1BFD2E" box="[1178,1232,657,683]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
in shape, size and shell proportions, but
<taxonomicName id="4C1C4D3E4F16155FFE577AC8FDAAFD4A" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[401,609,693,719]" class="Gastropoda" family="Heterosubulitidae" genus="Heterosubulites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="longusapertura" status="sp. nov.">
<emphasis id="B968EAAF4F16155FFE577AC8FDAAFD4A" box="[401,609,693,719]" italics="true" pageId="33" pageNumber="34">H. longusapertura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F16155FFDAF7AC8FD6DFD55" box="[617,678,694,720]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C1C4D3E4F16155FFCF87AC8FC69FD4A" authorityName="Knight" authorityYear="1931" box="[830,930,693,719]" class="Gastropoda" family="Acteoninidae" genus="Ceraunocochlis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="blatta">
<emphasis id="B968EAAF4F16155FFCF87AC8FC69FD4A" box="[830,930,693,719]" italics="true" pageId="33" pageNumber="34">H. blatta</emphasis>
</taxonomicName>
having a narrower aperture, a much broader anterior siphonal canal and a much more arched outer lip.
<taxonomicName id="4C1C4D3E4F16155FFCFB7AA4FBA4FD76" authority="Nutzel, 2012" authorityName="Nutzel" authorityYear="2012" box="[829,1135,729,756]" class="Gastropoda" family="Heterosubulitidae" genus="Heterosubulites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="fusiformis">
<emphasis id="B968EAAF4F16155FFCFB7AA4FC9AFD76" box="[829,849,730,755]" italics="true" pageId="33" pageNumber="34">H</emphasis>
.
<emphasis id="B968EAAF4F16155FFCA47AA4FC19FD71" box="[866,978,730,756]" italics="true" pageId="33" pageNumber="34">fusiformis</emphasis>
Nützel, 2012
</taxonomicName>
from the Middle Permian Akasaka Limestone,
<collectingCountry id="F30B762D4F16155FFE427A80FE0FFC9D" box="[388,452,766,792]" name="Japan" pageId="33" pageNumber="34">Japan</collectingCountry>
is also similar but its shell is larger, the shell is more slender, whorls embrace lower on preceding whorl, its aperture is wider and its anterior siphonal canal is narrower.
</paragraph>
</subSubSection>
</treatment>
</document>