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<document id="B656F928D8985DCC89640BAEEF59FC35" ID-CLB-Dataset="21928" ID-DOI="10.11646/zootaxa.4779.4.1" ID-GBIF-Dataset="d48aecc5-2a44-491e-802f-4556f052a7a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3839329" ID-ZooBank="861A7289-DD34-4174-B6B2-783FBEC8DF36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590137887931" checkinUser="plazi" docAuthor="Kluge, Nikita J. &amp; Suttinun, Chanaporn" docDate="2020" docId="03B587A2F21BFF8AECABFA740D67FF42" docLanguage="en" docName="zootaxa.4779.4.1.pdf" docOrigin="Zootaxa 4779 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indocloeon (Hindocloeon) Kluge &amp; Suttinun 2020" docType="treatment" docVersion="6" lastPageNumber="481" masterDocId="FF8CFFDAF207FF95EC3CFFC90C6FFFB9" masterDocTitle="Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species" masterLastPageNumber="484" masterPageNumber="451" pageNumber="479" updateTime="1698840818980" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2FC1F7FDF5373B0DA4BC5A78981DE94A">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:namePart id="50719D705A084C8AEFEA1DE9A94F84A6">Kluge, Nikita J.</mods:namePart>
<mods:affiliation id="C40B7E7377C0639B417D31E044B9A5CC">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="D55042C915EBDFA49188F1FCE7F2C72C">Suttinun, Chanaporn</mods:namePart>
<mods:affiliation id="E4EC23A3165B62C1B47FAE9D407B33B3">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<mods:title id="8665E23A628037CC129A718ACC12AAB6">Zootaxa</mods:title>
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<mods:date id="15C49D84BE83F790F9FF0734DE888205">2020</mods:date>
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<mods:number id="6B5875D0AC511BF12590B5A2E78DC582">4779</mods:number>
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<mods:classification id="5F8E202CE99101349CA33B5839C26C40">journal article</mods:classification>
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<treatment id="03B587A2F21BFF8AECABFA740D67FF42" ID-DOI="http://doi.org/10.5281/zenodo.3853356" ID-GBIF-Taxon="164248613" ID-Zenodo-Dep="3853356" LSID="urn:lsid:plazi:treatment:03B587A2F21BFF8AECABFA740D67FF42" httpUri="http://treatment.plazi.org/id/03B587A2F21BFF8AECABFA740D67FF42" lastPageId="31" lastPageNumber="481" pageId="28" pageNumber="479">
<subSubSection id="C306653FF21BFF89ECABFA740E0EFA61" box="[151,609,1469,1496]" pageId="28" pageNumber="479" type="nomenclature">
<paragraph id="8BA336B4F21BFF89ECABFA740E0EFA61" blockId="28.[151,609,1469,1532]" box="[151,609,1469,1496]" pageId="28" pageNumber="479">
<heading id="D0EB81D8F21BFF89ECABFA740E0EFA61" bold="true" box="[151,609,1469,1496]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
<emphasis id="B968EAA6F21BFF89ECABFA740E0EFA61" bold="true" box="[151,609,1469,1496]" pageId="28" pageNumber="479">
<taxonomicName id="4C1C4D37F21BFF89ECABFA740DB7FA6E" authority="" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[151,472,1469,1496]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21BFF89ECABFA740D4FFA61" bold="true" box="[151,288,1469,1496]" italics="true" pageId="28" pageNumber="479">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F21BFF89ED0DFA740DA2FA61" bold="true" box="[305,461,1469,1496]" italics="true" pageId="28" pageNumber="479">Hindocloeon</emphasis>
)
</taxonomicName>
sp. «
<collectingRegion id="49D8F856F21BFF89EE2BFA770E3CFA61" box="[535,595,1470,1496]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="479">Java</collectingRegion>
»
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C306653FF21BFF89ECABFA280DC0FA45" box="[151,431,1505,1532]" pageId="28" pageNumber="479" type="description">
<paragraph id="8BA336B4F21BFF89ECABFA280DC0FA45" blockId="28.[151,609,1469,1532]" box="[151,431,1505,1532]" pageId="28" pageNumber="479">
(
<figureCitation id="13272A31F21BFF89ECA3FA280D6BFA42" box="[159,260,1505,1531]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="28" pageNumber="479">Figs 104</figureCitation>
,
<figureCitation id="13272A31F21BFF89ED2CFA280D55FA42" box="[272,314,1505,1531]" captionStart="FIGURES 106–111" captionStartId="30.[151,264,1633,1658]" captionTargetBox="[151,1436,237,1608]" captionTargetId="figure@30.[151,1436,237,1608]" captionTargetPageId="30" captionText="FIGURES 106–111. Eggs of Hindocloeon. 106–108, Indocloeon (Hindocloeon) continentale sp. n.: 106–107, specimen from Erawan (Thailand); 108, specimen from Erumeli (India). 109–110, Indocloeon (Hindocloeon) indonesiae; 111, Indocloeon (Hindocloeon) sp. «Java»." figureDoi="http://doi.org/10.5281/zenodo.3839375" httpUri="https://zenodo.org/record/3839375/files/figure.png" pageId="28" pageNumber="479">111</figureCitation>
,
<figureCitation id="13272A31F21BFF89ED7AFA280DC8FA45" box="[326,423,1505,1532]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="28" pageNumber="479">115–118</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C306653FF21BFF8BECABF9E00FB6FF6E" lastPageId="30" lastPageNumber="481" pageId="28" pageNumber="479" type="materials_examined">
<paragraph id="8BA336B4F21BFF89ECABF9E00EEAF9DE" blockId="28.[151,1437,1577,2036]" pageId="28" pageNumber="479">
<emphasis id="B968EAA6F21BFF89ECABF9E00D17F9FD" bold="true" box="[151,376,1577,1604]" pageId="28" pageNumber="479">Material examined</emphasis>
(
<collectionCode id="ED0DAE71F21BFF89EDBAF9E00DD5F9FA" box="[390,442,1577,1603]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="28" pageNumber="479" type="Museum">ZIN</collectionCode>
).
<materialsCitation id="3B743CE9F21BFF89EDF7F9E00ECBF9FD" ID-GBIF-Occurrence="2622593306" box="[459,676,1577,1604]" country="Indonesia" location="Java" pageId="28" pageNumber="479" specimenCount="1" stateProvince="Jawa Timur">
<collectingCountry id="F30B7624F21BFF89EDF7F9E00E09F9FA" box="[459,614,1577,1603]" name="Indonesia" pageId="28" pageNumber="479">INDONESIA</collectingCountry>
,
<collectingRegion id="49D8F856F21BFF89EE4CF9E30ECBF9FD" box="[624,676,1578,1604]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="479">Java</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B743CE9F21BFF89EE93F9E00EEFF9DE" ID-GBIF-Occurrence="2622593303" collectingDate="2018-01-29" collectionCode="V" collectorName="S. Melnitsky &amp; Ivanov" country="Indonesia" elevation="99" latitude="-7.6383333" location="Parigaran" longLatPrecision="21" longitude="108.53001" pageId="28" pageNumber="479" specimenCount="101" specimenCount-female="1" specimenCount-male="1" stateProvince="Java">
<quantity id="4CE49B51F21BFF89EE93F9E00E9BF9FA" box="[687,756,1577,1604]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="479" unit="km" value="15.0">15 km</quantity>
NW
<location id="8EC3606FF21BFF89EF11F9E00FF4F9FA" LSID="urn:lsid:plazi:treatment:03B587A2F21BFF8AECABFA740D67FF42:8EC3606FF21BFF89EF11F9E00FF4F9FA" box="[813,923,1577,1603]" country="Indonesia" latitude="-7.6383333" longLatPrecision="21" longitude="108.53001" name="Parigaran" pageId="28" pageNumber="479" stateProvince="Java">Parigaran</location>
,
<geoCoordinate id="EE285073F21BFF89EF99F9E00871F9FA" box="[933,1054,1577,1603]" degrees="7" direction="south" minutes="38" orientation="latitude" pageId="28" pageNumber="479" precision="15" seconds="18" value="-7.6383333">S7°38’18”</geoCoordinate>
,
<geoCoordinate id="EE285073F21BFF89E814F9E008D1F9FA" box="[1064,1214,1577,1603]" degrees="108" direction="east" minutes="31" orientation="longitude" pageId="28" pageNumber="479" precision="15" seconds="48" value="108.53001">E108°31’48”</geoCoordinate>
, h=
<specimenCount id="9D1AFD3DF21BFF89E8DAF9E0094AF9FD" box="[1254,1317,1577,1604]" pageId="28" pageNumber="479" type="generic">
<quantity id="4CE49B51F21BFF89E8DAF9E0094EF9FD" box="[1254,1313,1577,1604]" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="28" pageNumber="479" unit="m" value="99.0">
<elevation id="0031D187F21BFF89E8DAF9E0094EF9FD" box="[1254,1313,1577,1604]" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="28" pageNumber="479" unit="m" value="99.0">99 m</elevation>
</quantity>
.
</specimenCount>
<date id="FFA21074F21BFF89E916F9E009F7F9FA" box="[1322,1432,1577,1603]" pageId="28" pageNumber="479" value="2018-01-29">
<collectingDate id="EFE6E99CF21BFF89E916F9E009F7F9FA" box="[1322,1432,1577,1603]" pageId="28" pageNumber="479" value="2018-01-29">29.I.2018</collectingDate>
</date>
, coll.
<collectorName id="26E95362F21BFF89ECF3F9840D30F9DE" box="[207,351,1613,1639]" pageId="28" pageNumber="479">S. Melnitsky</collectorName>
&amp;
<collectionCode id="ED0DAE71F21BFF89EDBEF9870DF9F9D1" box="[386,406,1614,1640]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="479" type="Museum">V</collectionCode>
.
<collectorName id="26E95362F21BFF89ED9CF9870D9DF9D1" box="[416,498,1614,1640]" pageId="28" pageNumber="479">Ivanov</collectorName>
: 2 S
<specimenCount id="9D1AFD3DF21BFF89EE1EF9840E57F9DE" box="[546,568,1613,1639]" pageId="28" pageNumber="479" type="male">♂</specimenCount>
, 2 S
<specimenCount id="9D1AFD3DF21BFF89EE51F9840EEFF9DE" box="[621,640,1613,1639]" pageId="28" pageNumber="479" type="female">♀</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA336B4F21BFF89ECFBF9B80E8DF916" blockId="28.[151,1437,1577,2036]" pageId="28" pageNumber="479">
<emphasis id="B968EAA6F21BFF89ECFBF9B80E91F935" bold="true" box="[199,766,1649,1676]" pageId="28" pageNumber="479">
Descriptions.
<emphasis id="B968EAA6F21BFF89ED55F9B80E98F935" bold="true" box="[361,759,1649,1676]" italics="true" pageId="28" pageNumber="479">Subimaginal cuticular colouration</emphasis>
.
</emphasis>
Most part of cuticle colourless, lateral sclerite of parascutellum and posterior part of postsubalar sclerite brown.
</paragraph>
<paragraph id="8BA336B4F21BFF89ECFBF9700FC4F8DA" blockId="28.[151,1437,1577,2036]" pageId="28" pageNumber="479">
<emphasis id="B968EAA6F21BFF89ECFBF9700D16F96D" bold="true" box="[199,377,1721,1748]" italics="true" pageId="28" pageNumber="479">Male subimago</emphasis>
(
<figureCitation id="13272A31F21BFF89EDBBF9700E4BF96A" box="[391,548,1721,1747]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="28" pageNumber="479">Figs 117–118</figureCitation>
). Turbinate eyes red. Antenna: distal part of scapus with hypodermal pigmented spots on lateral and median sides; distal part of pedicellus with hypodermal pigmented spots on lateral and median sides (as in
<figureCitation id="13272A31F21BFF89ED26F8C80D05F8A2" box="[282,362,1793,1819]" captionStart="FIGURES 91–95" captionStartId="25.[151,264,1887,1912]" captionTargetBox="[151,1436,181,1861]" captionTargetId="figure@25.[151,1436,181,1861]" captionTargetPageId="25" captionText="FIGURES 91–95. Indocloeon (Hindocloeon) continentale sp. n. 91, fragment of subimaginal exuviae with postsubalar sclerite; 92, antennae of male imago; 93, male imago; 94, its metathorax and base of abdomen; 95, male subimago (arrow shows brown hypodermal spot just ventrad of metathoracic postnotum)." figureDoi="http://doi.org/10.5281/zenodo.3839367" httpUri="https://zenodo.org/record/3839367/files/figure.png" pageId="28" pageNumber="479">Fig. 92</figureCitation>
) [see
<taxonomicName id="4C1C4D37F21BFF89ED91F8C80E51F8A2" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[429,574,1793,1819]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="455" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21BFF89ED91F8C80E51F8A2" box="[429,574,1793,1819]" italics="true" pageId="28" pageNumber="479">Hindocloeon</emphasis>
</taxonomicName>
(9)]. First abdominal segment with pair of brown hypodermal spots just ventrad of metathoracic postnotum (
<figureCitation id="13272A31F21BFF89EE39F8EC0E0BF886" box="[517,612,1829,1855]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="28" pageNumber="479">Fig. 117</figureCitation>
) [see
<taxonomicName id="4C1C4D37F21BFF89EE9BF8EC0F57F886" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[679,824,1829,1855]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="28" pageNumber="455" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21BFF89EE9BF8EC0F57F886" box="[679,824,1829,1855]" italics="true" pageId="28" pageNumber="479">Hindocloeon</emphasis>
</taxonomicName>
(10)]. Other abdominal terga either without pigmentation, or terga VII–IX with brown hypodermal dot at middle (
<figureCitation id="13272A31F21BFF89EF02F8800FF0F8DA" box="[830,927,1865,1891]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="28" pageNumber="479">Fig. 118</figureCitation>
).
</paragraph>
<paragraph id="8BA336B4F21BFF88ECFBF8A40E44FF42" blockId="28.[151,1437,1577,2036]" lastBlockId="29.[151,1436,153,251]" lastPageId="29" lastPageNumber="480" pageId="28" pageNumber="479">
<emphasis id="B968EAA6F21BFF89ECFBF8A40DFBF831" bold="true" box="[199,404,1901,1928]" italics="true" pageId="28" pageNumber="479">Female subimago</emphasis>
(
<figureCitation id="13272A31F21BFF89ED9EF8A40E50F831" box="[418,575,1901,1928]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="28" pageNumber="479">Figs 115–116</figureCitation>
). Antennae as in male. Pronotum with three pairs of small, contrasting, brown hypodermal markings on lateral parts: zigzag anterior, elongate postero-median and triangular postero-lateral ones. Mesothorax with small, brown hypodermal spots on pleura. Abdomen with dark, lateral tracheal trunks; segment I with pair of brown hypodermal lateral spots just ventrad of metathoracic postnotum; each segment II–VII with fol- lowing contrasting brown hypodermal maculae: lateral side of tergum with longitudinal triangular macula (widest anteriorly) and darker brown dot near antero-median angle of this macula; lateral side of sternum with longitudinal macula of peculiar shape (
<figureCitation id="13272A31F21AFF88ED82FF280E70FF42" box="[446,543,224,251]" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="29" pageNumber="480">Fig. 115</figureCitation>
).
</paragraph>
<caption id="DF63663CF21AFF88ECABF8F40825F85F" ID-DOI="http://doi.org/10.5281/zenodo.3839373" ID-Zenodo-Dep="3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="29" pageNumber="480" startId="29.[151,264,1853,1878]" targetBox="[151,1427,281,1828]" targetPageId="29">
<paragraph id="8BA336B4F21AFF88ECABF8F40825F85F" blockId="29.[151,1437,1853,2022]" pageId="29" pageNumber="480">
<emphasis id="B968EAA6F21AFF88ECABF8F40D1FF8EF" bold="true" box="[151,368,1853,1878]" pageId="29" pageNumber="480">FIGURES 101–105.</emphasis>
Male genitals of
<taxonomicName id="4C1C4D37F21AFF88EE15F8F70EC3F8EF" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[553,684,1854,1878]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="455" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F21AFF88EE15F8F70EC3F8EF" box="[553,684,1854,1878]" italics="true" pageId="29" pageNumber="480">Hindocloeon</emphasis>
</taxonomicName>
. 101, imaginal genitals of
<taxonomicName id="4C1C4D37F21AFF88EFF9F8F70937F8EF" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[965,1368,1854,1878]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="continentale" status="sp. nov." subGenus="Hindocloeon">
<emphasis id="B968EAA6F21AFF88EFF9F8F70858F8EF" box="[965,1079,1854,1878]" italics="true" pageId="29" pageNumber="480">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F21AFF88E874F8F708A4F8EF" box="[1096,1227,1854,1878]" italics="true" pageId="29" pageNumber="480">Hindocloeon</emphasis>
)
<emphasis id="B968EAA6F21AFF88E8E7F8F70937F8EF" box="[1243,1368,1854,1878]" italics="true" pageId="29" pageNumber="480">continentale</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F21AFF88E95DF8F709F3F8EF" bold="true" box="[1377,1436,1854,1878]" pageId="29" pageNumber="480">
<taxonomicNameLabel id="A25B57DDF21AFF88E95DF8F709F3F8EF" box="[1377,1436,1854,1878]" pageId="29" pageNumber="480" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of
<emphasis id="B968EAA6F21AFF88E833F8630871F878" box="[1039,1054,1962,1985]" italics="true" pageId="29" pageNumber="480">I.</emphasis>
(
<emphasis id="B968EAA6F21AFF88E817F863082AF878" box="[1067,1093,1962,1985]" italics="true" pageId="29" pageNumber="480">H.</emphasis>
) sp. «Vietnam»; 104, subimaginal gonostyli of
<emphasis id="B968EAA6F21AFF88ED2BF8070D48F85C" box="[279,295,1998,2021]" italics="true" pageId="29" pageNumber="480">I.</emphasis>
(
<emphasis id="B968EAA6F21AFF88ED09F8070D21F85C" box="[309,334,1998,2021]" italics="true" pageId="29" pageNumber="480">H.</emphasis>
) sp. «Java»; 105, imaginal genitals of
<taxonomicName id="4C1C4D37F21AFF88EEE9F8070F37F85F" authorityName="Kluge &amp; Suttinun" authorityYear="2020" baseAuthorityName="Demoulin" baseAuthorityYear="1969" box="[725,856,1998,2022]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="longistylus">
<emphasis id="B968EAA6F21AFF88EEE9F8070F37F85F" box="[725,856,1998,2022]" italics="true" pageId="29" pageNumber="480">I. longistylus</emphasis>
</taxonomicName>
(after
<bibRefCitation id="EF8D4B45F21AFF88EFA0F8040851F85F" author="Demoulin, G." box="[924,1086,1997,2022]" pageId="29" pageNumber="480" pagination="225 - 241" refId="ref14512" refString="Demoulin, G. (1969) Les Ephemeropteres recoltes par la mission danoise du Noona Dan aux iles Philippines et Bismarck. Entomologiske Meddelelser, 37 (3), 225 - 241." type="journal article" year="1969">Demoulin 1969</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BA336B4F219FF8BECFBFF500845FF0A" blockId="30.[199,1066,152,216]" box="[199,1066,152,179]" pageId="30" pageNumber="481">
<emphasis id="B968EAA6F219FF8BECFBFF500C99FF0A" bold="true" box="[199,246,153,179]" italics="true" pageId="30" pageNumber="481">Egg</emphasis>
(
<figureCitation id="13272A31F219FF8BED39FF500D09FF0A" box="[261,358,153,179]" captionStart="FIGURES 106–111" captionStartId="30.[151,264,1633,1658]" captionTargetBox="[151,1436,237,1608]" captionTargetId="figure@30.[151,1436,237,1608]" captionTargetPageId="30" captionText="FIGURES 106–111. Eggs of Hindocloeon. 106–108, Indocloeon (Hindocloeon) continentale sp. n.: 106–107, specimen from Erawan (Thailand); 108, specimen from Erumeli (India). 109–110, Indocloeon (Hindocloeon) indonesiae; 111, Indocloeon (Hindocloeon) sp. «Java»." figureDoi="http://doi.org/10.5281/zenodo.3839375" httpUri="https://zenodo.org/record/3839375/files/figure.png" pageId="30" pageNumber="481">Fig. 111</figureCitation>
). Oval, about
<quantity id="4CE49B51F219FF8BEE3EFF510E09FF0A" box="[514,614,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="481" unit="mm" value="0.15">0.15 mm</quantity>
length. Chorion with very fine rugosity.
</paragraph>
<paragraph id="8BA336B4F219FF8BECFBFF750FB6FF6E" blockId="30.[199,1066,152,216]" box="[199,985,188,216]" pageId="30" pageNumber="481">
<emphasis id="B968EAA6F219FF8BECFBFF750D21FF6E" bold="true" box="[199,334,188,215]" pageId="30" pageNumber="481">Dimension.</emphasis>
Fore wing length (and approximate body length)
<quantity id="4CE49B51F219FF8BEF40FF750FBAFF6E" box="[892,981,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="30" pageNumber="481" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF63663CF219FF8BECABF9A80DCDF978" ID-DOI="http://doi.org/10.5281/zenodo.3839375" ID-Zenodo-Dep="3839375" httpUri="https://zenodo.org/record/3839375/files/figure.png" pageId="30" pageNumber="481" startId="30.[151,264,1633,1658]" targetBox="[151,1436,237,1608]" targetPageId="30">
<paragraph id="8BA336B4F219FF8BECABF9A80DCDF978" blockId="30.[151,1436,1633,1730]" pageId="30" pageNumber="481">
<emphasis id="B968EAA6F219FF8BECABF9A80D03F9C3" bold="true" box="[151,364,1633,1658]" pageId="30" pageNumber="481">FIGURES 106–111.</emphasis>
Eggs of
<taxonomicName id="4C1C4D37F219FF8BEDF4F9AB0E24F9C3" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[456,587,1634,1658]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="455" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BEDF4F9AB0E24F9C3" box="[456,587,1634,1658]" italics="true" pageId="30" pageNumber="481">Hindocloeon</emphasis>
</taxonomicName>
. 106–108,
<taxonomicName id="4C1C4D37F219FF8BEE82F9AB0820F9C3" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[702,1103,1634,1658]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="continentale" status="sp. nov." subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BEE82F9AB0F5FF9C3" box="[702,816,1634,1658]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F219FF8BEF7CF9AB0FACF9C3" box="[832,963,1634,1658]" italics="true" pageId="30" pageNumber="481">Hindocloeon</emphasis>
)
<emphasis id="B968EAA6F219FF8BEFEEF9AB0820F9C3" box="[978,1103,1634,1658]" italics="true" pageId="30" pageNumber="481">continentale</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F219FF8BE86AF9AB08FFF9C3" bold="true" box="[1110,1168,1634,1658]" pageId="30" pageNumber="481">
<taxonomicNameLabel id="A25B57DDF219FF8BE86AF9AB08FFF9C3" box="[1110,1168,1634,1658]" pageId="30" pageNumber="481" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 106–107, specimen from Erawan (Thailand); 108, specimen from Erumeli (India). 109–110,
<taxonomicName id="4C1C4D37F219FF8BEF62F94F088AF927" authorityName="Kluge" authorityYear="2012" box="[862,1253,1670,1694]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="indonesiae" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BEF62F94F0FBFF927" box="[862,976,1670,1694]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F219FF8BEFDEF94F080AF927" box="[994,1125,1670,1694]" italics="true" pageId="30" pageNumber="481">Hindocloeon</emphasis>
)
<emphasis id="B968EAA6F219FF8BE84BF94F088AF927" box="[1143,1253,1670,1694]" italics="true" pageId="30" pageNumber="481">indonesiae</emphasis>
</taxonomicName>
; 111,
<taxonomicName id="4C1C4D37F219FF8BE916F94F0D45F97B" authority="" authorityName="Kluge &amp; Suttinun" authorityYear="2020" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="subGenus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BE916F94F09F3F927" box="[1322,1436,1670,1694]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F219FF8BEC9CF9630D4CF97B" box="[160,291,1706,1730]" italics="true" pageId="30" pageNumber="481">Hindocloeon</emphasis>
)
</taxonomicName>
sp. «Java».
</paragraph>
</caption>
<subSubSection id="C306653FF219FF8AECFBF9260D67FF42" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481" type="discussion">
<paragraph id="8BA336B4F219FF8AECFBF9260D67FF42" blockId="30.[151,1437,1775,2019]" lastBlockId="31.[151,1436,153,251]" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
<emphasis id="B968EAA6F219FF8BECFBF9260D20F8B3" bold="true" box="[199,335,1775,1802]" pageId="30" pageNumber="481">Comments.</emphasis>
Male subimagines are determined as belonging to
<taxonomicName id="4C1C4D37F219FF8BEFB3F9390862F8B3" authorityName="Muller-Liebenau" authorityYear="1982" box="[911,1037,1776,1802]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F219FF8BEFB3F9390862F8B3" box="[911,1037,1776,1802]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF219FF8BE828F9380852F8B3" box="[1044,1085,1776,1803]" pageId="30" pageNumber="481" sensu="lato">s. l.</taxonomicNameLabel>
based on the peculiar structure of gonostyli (
<figureCitation id="13272A31F219FF8BED12F8DD0DFFF897" box="[302,400,1812,1838]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="30" pageNumber="481">Fig. 104</figureCitation>
). Female subimagines are determined as belonging to the same species based on equally colourless cuticle and the same structure of middle and hind legs as in males. Belonging of the female subimagines to the genus
<taxonomicName id="4C1C4D37F219FF8BED34F8950DE9F8CF" authorityName="Muller-Liebenau" authorityYear="1982" box="[264,390,1884,1910]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F219FF8BED34F8950DE9F8CF" box="[264,390,1884,1910]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF219FF8BEDB0F8940DDBF8CF" box="[396,436,1884,1911]" pageId="30" pageNumber="481" sensu="lato">s. l.</taxonomicNameLabel>
is proven by the absence of the patella-tibial suture on fore legs. This species differs from
<taxonomicName id="4C1C4D37F219FF8BECABF8480D31F823" authorityName="Kluge &amp; Suttinun" authorityYear="2020" baseAuthorityName="Demoulin" baseAuthorityYear="1969" box="[151,350,1920,1947]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="longistylus" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BECABF8480CC7F823" box="[151,168,1921,1946]" italics="true" pageId="30" pageNumber="481">I.</emphasis>
(
<emphasis id="B968EAA6F219FF8BEC85F8480CBBF823" box="[185,212,1921,1946]" italics="true" pageId="30" pageNumber="481">H.</emphasis>
)
<emphasis id="B968EAA6F219FF8BECD9F8490D31F823" box="[229,350,1920,1946]" italics="true" pageId="30" pageNumber="481">longistylus</emphasis>
</taxonomicName>
by having a relatively longer proximal segment of the gonostylus even in the subimaginal stage (
<figureCitation id="13272A31F219FF8BEC9CF86D0D6DF807" box="[160,258,1956,1982]" captionStart="FIGURES 101–105" captionStartId="29.[151,264,1853,1878]" captionTargetBox="[151,1427,281,1828]" captionTargetId="figure@29.[151,1436,273,1828]" captionTargetPageId="29" captionText="FIGURES 101–105. Male genitals of Hindocloeon. 101, imaginal genitals of Indocloeon (Hindocloeon) continentale sp. n. (penis dotted; 1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; gv-d, distal portion of gonovectis; gv-p, proximal portion of gonovectis; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 102, their subimaginal exuviae; 103, subimaginal gonostyli of I. (H.) sp. «Vietnam»; 104, subimaginal gonostyli of I. (H.) sp. «Java»; 105, imaginal genitals of I. longistylus (after Demoulin 1969)." figureDoi="http://doi.org/10.5281/zenodo.3839373" httpUri="https://zenodo.org/record/3839373/files/figure.png" pageId="30" pageNumber="481">Fig. 104</figureCitation>
). Male and female subimagines differ from the subgenus
<taxonomicName id="4C1C4D37F219FF8BEFBBF86D086AF807" authorityName="Muller-Liebenau" authorityYear="1982" box="[903,1029,1956,1982]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAA6F219FF8BEFBBF86D086AF807" box="[903,1029,1956,1982]" italics="true" pageId="30" pageNumber="481">Indocloeon</emphasis>
</taxonomicName>
by the absence of reddish stripes on posterior margins of abdominal terga. The male differs from
<taxonomicName id="4C1C4D37F219FF8BEF74F8000860F85B" authorityName="Kluge" authorityYear="2012" box="[840,1039,1992,2019]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="indonesiae" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BEF74F8000F37F85B" box="[840,856,1993,2018]" italics="true" pageId="30" pageNumber="481">I.</emphasis>
(
<emphasis id="B968EAA6F219FF8BEF55F8000FEAF85B" box="[873,901,1993,2018]" italics="true" pageId="30" pageNumber="481">H.</emphasis>
)
<emphasis id="B968EAA6F219FF8BEFAAF8010860F85B" box="[918,1039,1992,2018]" italics="true" pageId="30" pageNumber="481">indonesiae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D37F219FF8BE822F8000899F85B" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[1054,1270,1992,2019]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="continentale" subGenus="Hindocloeon">
<emphasis id="B968EAA6F219FF8BE822F8000840F85B" box="[1054,1071,1993,2018]" italics="true" pageId="30" pageNumber="481">I.</emphasis>
(
<emphasis id="B968EAA6F219FF8BE803F8000834F85B" box="[1087,1115,1993,2018]" italics="true" pageId="30" pageNumber="481">H.</emphasis>
)
<emphasis id="B968EAA6F219FF8BE850F8010899F85B" box="[1132,1270,1992,2018]" italics="true" pageId="30" pageNumber="481">continentale</emphasis>
</taxonomicName>
and
<emphasis id="B968EAA6F219FF8BE912F8000950F85B" box="[1326,1343,1993,2018]" italics="true" pageId="30" pageNumber="481">I.</emphasis>
(
<emphasis id="B968EAA6F219FF8BE96CF8000903F85B" box="[1360,1388,1993,2018]" italics="true" pageId="30" pageNumber="481">H.</emphasis>
) sp. «
<collectingCountry id="F30B7624F218FF8AEC9FFF500D69FF0A" box="[163,262,153,179]" name="Vietnam" pageId="31" pageNumber="482">Vietnam</collectingCountry>
» by the shape of the gonostylus. The female subimago differs from
<taxonomicName id="4C1C4D37F218FF8AEFCAFF5308D5FF0A" authorityName="Kluge" authorityYear="2012" box="[1014,1210,153,179]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="indonesiae" subGenus="Hindocloeon">
<emphasis id="B968EAA6F218FF8AEFCAFF530868FF0A" box="[1014,1031,154,179]" italics="true" pageId="31" pageNumber="482">I.</emphasis>
(
<emphasis id="B968EAA6F218FF8AE82AFF53085DFF0A" box="[1046,1074,154,179]" italics="true" pageId="31" pageNumber="482">H.</emphasis>
)
<emphasis id="B968EAA6F218FF8AE87DFF5008D5FF0A" box="[1089,1210,153,179]" italics="true" pageId="31" pageNumber="482">indonesiae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D37F218FF8AE8FBFF5309F3FF0A" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[1223,1436,153,179]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="continentale" subGenus="Hindocloeon">
<emphasis id="B968EAA6F218FF8AE8FBFF5308B7FF0A" box="[1223,1240,154,179]" italics="true" pageId="31" pageNumber="482">I.</emphasis>
(
<emphasis id="B968EAA6F218FF8AE8DBFF53096CFF0A" box="[1255,1283,154,179]" italics="true" pageId="31" pageNumber="482">H.</emphasis>
)
<emphasis id="B968EAA6F218FF8AE92EFF5009F3FF0A" box="[1298,1436,153,179]" italics="true" pageId="31" pageNumber="482">continentale</emphasis>
</taxonomicName>
and
<emphasis id="B968EAA6F218FF8AECF4FF770CB7FF6E" box="[200,216,190,215]" italics="true" pageId="31" pageNumber="482">I.</emphasis>
(
<emphasis id="B968EAA6F218FF8AECD5FF770D6AFF6E" box="[233,261,190,215]" italics="true" pageId="31" pageNumber="482">H.</emphasis>
) sp. «
<collectingCountry id="F30B7624F218FF8AED76FF740DC2FF6E" box="[330,429,189,215]" name="Vietnam" pageId="31" pageNumber="482">Vietnam</collectingCountry>
» by the peculiar shape of brown maculae on abdominal terga and on the pronotum (
<figureCitation id="13272A31F218FF8AE956FF740C93FF42" captionStart="FIGURES 115–118" captionStartId="32.[151,264,1464,1489]" captionTargetBox="[152,1436,381,1439]" captionTargetId="figure@32.[152,1436,381,1439]" captionTargetPageId="32" captionText="FIGURES 115–118. Indocloeon (Hindocloeon) sp. «Java». 115–116, female subimago; 117–118, male subimago." figureDoi="http://doi.org/10.5281/zenodo.3839379" httpUri="https://zenodo.org/record/3839379/files/figure.png" pageId="31" pageNumber="482">Figs 115–116</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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