410 lines
58 KiB
XML
410 lines
58 KiB
XML
<document id="53DB707E3383E2345FDD0B4738FFBABB" ID-DOI="10.11646/zootaxa.4803.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3907900" ID-ZooBank="2D391AAF-F105-4252-A090-34E725A05315" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593082538606" checkinUser="plazi" docAuthor="Chirivi-Joya, Daniel, Moreno-González, Jairo A. & Fagua, Giovanny" docDate="2020" docId="03B58796B645FFDEFF7EFF30FAA27492" docLanguage="en" docName="zootaxa.4803.1.1.pdf" docOrigin="Zootaxa 4803 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterophrynus awa Chirivi-Joya & Moreno-González & Fagua 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="27" masterDocId="FF8CFFEEB652FFC4FFE9FFA9FFE77661" masterDocTitle="Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species" masterLastPageNumber="41" masterPageNumber="1" pageNumber="24" updateTime="1700277040307" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BD9A49254835625B77D61F2187430D0C">
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<mods:title id="B9DE0411CC8CEEE3A83FF4C3B43F8A8A">Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species</mods:title>
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<mods:name id="F42B7321454CA138B6A1CDCA4C7BCC6B" type="personal">
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<mods:roleTerm id="E81CC1823527F92092A5797ED234E68F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="88D547E94F724BBAC5E4E356D64F2929">Chirivi-Joya, Daniel</mods:namePart>
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<mods:affiliation id="9E39615A736FA3DFF6052EAA30156405">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. & dhipnosodin @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1198 - 782 X</mods:affiliation>
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<mods:roleTerm id="A7EDCE9F138D786FA875F4E4C1B4B023">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="292D74435AE302DCCD84771A01A21191">Moreno-González, Jairo A.</mods:namePart>
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<mods:affiliation id="B41D9C1514FFF8E33E8920B05AF0DE7E">Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo. Rua do Matão, trav. 14, n ° 321, Cidade Universitária, São Paulo- SP, CEP: 05508 - 090. hansenochrus @ gmail. com; https: // orcid. org / 0000 - 0001 - 5980 - 1417</mods:affiliation>
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<mods:roleTerm id="7F647740E9AF2B205AA07A28E537053E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EADABDD88CCC38C9EE4C2BB7EDCF545F">Fagua, Giovanny</mods:namePart>
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<mods:affiliation id="125C1C8921F36127BE621A1FABE1FC37">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. & fagua @ javeriana. edu. co; https: // orcid. org / 0000 - 0003 - 1253 - 4416</mods:affiliation>
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<mods:typeOfResource id="19CD103F10FCDD8D8FB7482FDDA3FB8C">text</mods:typeOfResource>
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<mods:title id="029975FBFB2E0B9FE1C3EABAB9ABC123">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="79CE0B1522E575175A3E0868275F49AF">
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<mods:date id="392CEB2786CF899C4AD8521D02F3EF48">2020</mods:date>
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<mods:number id="F635592A423ABEBDF1F5B986DC6A35DB">2020-06-25</mods:number>
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<mods:detail id="AF4523B1E6AFBDF52A8260E420983604" type="volume">
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<mods:number id="C82F2ADED01A3E1F700FCAA6354BEE84">4803</mods:number>
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<mods:identifier id="82505D73E068A152C68C3FB5D5B2168C" type="DOI">10.11646/zootaxa.4803.1.1</mods:identifier>
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<treatment id="03B58796B645FFDEFF7EFF30FAA27492" LSID="urn:lsid:plazi:treatment:03B58796B645FFDEFF7EFF30FAA27492" httpUri="http://treatment.plazi.org/id/03B58796B645FFDEFF7EFF30FAA27492" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
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<subSubSection id="C306650BB645FFD3FF7EFF30FE0C76D5" box="[151,491,153,180]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BA33680B645FFD3FF7EFF30FE0C76D5" blockId="23.[151,556,153,216]" box="[151,491,153,180]" pageId="23" pageNumber="24">
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<heading id="D0EB81ECB645FFD3FF7EFF30FE0C76D5" bold="true" box="[151,491,153,180]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B968EA92B645FFD3FF7EFF30FE0C76D5" bold="true" box="[151,491,153,180]" pageId="23" pageNumber="24">
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<taxonomicName id="4C1C4D03B645FFD3FF7EFF30FE6176D5" ID-CoL="8VN4W" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[151,390,153,180]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
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<emphasis id="B968EA92B645FFD3FF7EFF30FE6176D5" bold="true" box="[151,390,153,180]" italics="true" pageId="23" pageNumber="24">Heterophrynus awa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57E9B645FFD3FE64FF33FE0C76D5" box="[397,491,154,180]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306650BB645FFD3FF7EFF14FDCB76B9" box="[151,556,189,216]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BA33680B645FFD3FF7EFF14FDCB76B9" blockId="23.[151,556,153,216]" box="[151,556,189,216]" pageId="23" pageNumber="24">
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<figureCitation id="13272A05B645FFD3FF7EFF14FEDC76B6" box="[151,315,189,215]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="23.[151,250,1934,1959]" captionStartId-1="24.[151,250,1928,1953]" captionStartId-2="26.[151,250,1910,1935]" captionTargetBox-0="[157,1426,315,1904]" captionTargetBox-1="[164,1423,534,1894]" captionTargetBox-2="[156,1432,784,1880]" captionTargetId-0="figure@23.[151,1436,309,1909]" captionTargetId-1="figure@24.[151,1436,525,1903]" captionTargetId-2="figure@26.[151,1437,777,1886]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="26" captionText-0="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." captionText-1="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." captionText-2="FIGURE 16. Male genital: Heterophrynus awa sp. nov. (ICN-Am-108): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (ICN-Am-75), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi-0="http://doi.org/10.5281/zenodo.3907930" figureDoi-1="http://doi.org/10.5281/zenodo.3907932" figureDoi-2="http://doi.org/10.5281/zenodo.3907934" httpUri-0="https://zenodo.org/record/3907930/files/figure.png" httpUri-1="https://zenodo.org/record/3907932/files/figure.png" httpUri-2="https://zenodo.org/record/3907934/files/figure.png" pageId="23" pageNumber="24">Figures 14–16</figureCitation>
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,
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<figureCitation id="13272A05B645FFD3FEAFFF14FE8576B6" box="[326,354,189,215]" captionStart="FIGURE 22" captionStartId="33.[151,250,1654,1679]" captionTargetBox="[185,1402,574,1621]" captionTargetId="figure@33.[177,1410,561,1629]" captionTargetPageId="33" captionText="FIGURE 22. Distribution map. Heterophrynus alces: purple circle. Heterophrynus armiger: green circle. Heterophrynus cheiracanthus: blue circle. Heterophrynus elaphus: orange circle. Heterophrynus awa sp. nov.: gray circle. Heterophrynus origamii sp. nov.: red circle." figureDoi="http://doi.org/10.5281/zenodo.3907947" httpUri="https://zenodo.org/record/3907947/files/figure.png" pageId="23" pageNumber="24">22</figureCitation>
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(in part).
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<tableCitation id="C69E033BB645FFD3FE3BFF14FDC176B9" box="[466,550,189,216]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="23" pageNumber="24">Table 5</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306650BB645FFDCFF7EFEAFFBB77706" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BA33680B645FFD3FF7EFEAFFDF5777E" blockId="23.[151,530,261,288]" box="[151,530,261,288]" pageId="23" pageNumber="24">
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<materialsCitation id="3B743CDDB645FFD3FF7EFEAFFDE9777E" box="[151,526,261,288]" country="Colombia" location="Narino" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
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<emphasis id="B968EA92B645FFD3FF7EFEAFFEDE7741" bold="true" box="[151,313,261,288]" pageId="23" pageNumber="24">
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<typeStatus id="54A78822B645FFD3FF7EFEAFFF357741" box="[151,210,262,288]" pageId="23" pageNumber="24">Type</typeStatus>
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locality:
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</emphasis>
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<collectingCountry id="F30B7610B645FFD3FEA9FEACFE56777E" box="[320,433,261,287]" name="Colombia" pageId="23" pageNumber="24">Colombia</collectingCountry>
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,
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<collectingRegion id="49D8F862B645FFD3FE54FEACFDE9777E" box="[445,526,261,287]" country="Colombia" name="Narino" pageId="23" pageNumber="24">Nariño</collectingRegion>
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</materialsCitation>
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.
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</paragraph>
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<caption id="DF636608B645FFD3FF7EF827FD2A71AA" ID-DOI="http://doi.org/10.5281/zenodo.3907930" ID-Zenodo-Dep="3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1934,1959]" targetBox="[157,1426,315,1904]" targetPageId="23">
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<paragraph id="8BA33680B645FFD3FF7EF827FD2A71AA" blockId="23.[151,1437,1934,1995]" pageId="23" pageNumber="24">
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<emphasis id="B968EA92B645FFD3FF7EF827FEF871C6" bold="true" box="[151,287,1934,1959]" pageId="23" pageNumber="24">FIGURE 14.</emphasis>
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<taxonomicName id="4C1C4D03B645FFD3FECDF826FE0D71C6" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[292,490,1935,1959]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
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<emphasis id="B968EA92B645FFD3FECDF826FE0D71C6" box="[292,490,1935,1959]" italics="true" pageId="23" pageNumber="24">Heterophrynus awa</emphasis>
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</taxonomicName>
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<emphasis id="B968EA92B645FFD3FE06F826FDD971C6" bold="true" box="[495,574,1935,1959]" pageId="23" pageNumber="24">
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<taxonomicNameLabel id="A25B57E9B645FFD3FE06F826FDD971C6" box="[495,574,1935,1959]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
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</paragraph>
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</caption>
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<paragraph id="8BA33680B64AFFDCFF2EFF30FBB77706" blockId="24.[151,1437,152,503]" pageId="24" pageNumber="25">
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<emphasis id="B968EA92B64AFFDCFF2EFF30FE9176D2" bold="true" box="[199,374,152,179]" pageId="24" pageNumber="25">Type material:</emphasis>
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<materialsCitation id="3B743CDDB64AFFDCFE94FF30FDE4769A" collectingDate="2009-09-14" collectorName="Martinez, H" country="Colombia" county="Barbacoas" elevation="500" latitude="1.3833333" location="Reserva Natural Biotopo" longLatPrecision="21" longitude="-78.25" municipality="Tumaco-Pasto" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
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<typeStatus id="54A78822B64AFFDCFE94FF30FE0276D2" box="[381,485,153,179]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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(male):
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:
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<collectingRegion id="49D8F862B64AFFDCFD0FFF33FCD276D2" box="[742,821,154,179]" country="Colombia" name="Narino" pageId="24" pageNumber="25">Nariño</collectingRegion>
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</emphasis>
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:
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<collectingCounty id="62C24E0CB64AFFDCFCAAFF30FC5B76D2" box="[835,956,153,179]" pageId="24" pageNumber="25">Barbacoas</collectingCounty>
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: km 95 via
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<collectingMunicipality id="6BC7ACFAB64AFFDCFBD3FF30FB3976D2" box="[1082,1246,153,179]" pageId="24" pageNumber="25">Tumaco-Pasto</collectingMunicipality>
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,
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<location id="8EC3605BB64AFFDCFB01FF30FF1676B6" LSID="urn:lsid:plazi:treatment:03B58796B645FFDEFF7EFF30FAA27492:8EC3605BB64AFFDCFB01FF30FF1676B6" country="Colombia" county="Barbacoas" latitude="1.3833333" longLatPrecision="21" longitude="-78.25" municipality="Tumaco-Pasto" name="Reserva Natural Biotopo" pageId="24" pageNumber="25" stateProvince="Narino">Reserva Natural Biotopo</location>
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(
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;
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):
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<location id="8EC3605BB64AFFDCFDADFF14FD2F76B6" LSID="urn:lsid:plazi:treatment:03B58796B645FFDEFF7EFF30FAA27492:8EC3605BB64AFFDCFDADFF14FD2F76B6" box="[580,712,189,215]" country="Colombia" county="Barbacoas" latitude="1.3833333" longLatPrecision="21" longitude="-78.25" municipality="Tumaco-Pasto" name="Rain Forest" pageId="24" pageNumber="25" stateProvince="Narino">Rain Forest</location>
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,
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<quantity id="4CE49B65B64AFFDCFD3AFF15FCDC76B6" box="[723,827,188,215]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">
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<elevation id="0031D1B3B64AFFDCFD3AFF15FCDC76B6" box="[723,827,188,215]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">500 masl</elevation>
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</quantity>
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,
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<date id="FFA21040B64AFFDCFCACFF14FBC676B6" box="[837,1057,189,215]" pageId="24" pageNumber="25" value="2009-09-14">
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<collectingDate id="EFE6E9A8B64AFFDCFCACFF14FBC676B6" box="[837,1057,189,215]" pageId="24" pageNumber="25" value="2009-09-14">14 September 2009</collectingDate>
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</date>
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, D,
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<collectorName id="26E95356B64AFFDCFBA4FF14FB3176B6" box="[1101,1238,189,215]" pageId="24" pageNumber="25">Martinez, H</collectorName>
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||
. Herreño, manual collection, male (ICN-Am-108)
|
||
</materialsCitation>
|
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.
|
||
<materialsCitation id="3B743CDDB64AFFDCFDE6FF48FBAB7706" collectingDate="2009-09-14" collectorName="Martinez, H & Vereda Altaquer & April" country="Colombia" county="Barbacoas" elevation="500" latitude="1.246" location="Reserva Natural Biotopo" longLatPrecision="1" longitude="-78.11531" municipality="Tumaco-Pasto" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Narino" typeStatus="paratype">
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<typeStatus id="54A78822B64AFFDCFDE6FF48FD9B769A" box="[527,636,225,251]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
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(
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<specimenCount id="9D1AFD09B64AFFDCFD78FF48FD12769A" box="[657,757,225,251]" pageId="24" pageNumber="25" type="female">1 female</specimenCount>
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,
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<specimenCount id="9D1AFD09B64AFFDCFCEAFF48FCB1769D" box="[771,854,225,252]" pageId="24" pageNumber="25" type="male">1 male</specimenCount>
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):
|
||
<collectingCountry id="F30B7610B64AFFDCFC82FF48FBE2769A" box="[875,1029,225,251]" name="Colombia" pageId="24" pageNumber="25">COLOMBIA</collectingCountry>
|
||
:
|
||
<emphasis id="B968EA92B64AFFDCFBF8FF4BFB87769A" box="[1041,1120,226,251]" italics="true" pageId="24" pageNumber="25">
|
||
<collectingRegion id="49D8F862B64AFFDCFBF8FF4BFB87769A" box="[1041,1120,226,251]" country="Colombia" name="Narino" pageId="24" pageNumber="25">Nariño</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty id="62C24E0CB64AFFDCFB9BFF48FB0C769A" box="[1138,1259,225,251]" pageId="24" pageNumber="25">Barbacoas</collectingCounty>
|
||
: km 95 via
|
||
<collectingMunicipality id="6BC7ACFAB64AFFDCFA9CFF48FEFA777E" pageId="24" pageNumber="25">Tumaco-Pasto</collectingMunicipality>
|
||
,
|
||
<location id="8EC3605BB64AFFDCFEC1FEACFDD9777E" LSID="urn:lsid:plazi:treatment:03B58796B645FFDEFF7EFF30FAA27492:8EC3605BB64AFFDCFEC1FEACFDD9777E" box="[296,574,261,287]" country="Colombia" county="Barbacoas" latitude="1.246" longLatPrecision="1" longitude="-78.11531" municipality="Tumaco-Pasto" name="Reserva Natural Biotopo" pageId="24" pageNumber="25" stateProvince="Narino">Reserva Natural Biotopo</location>
|
||
(
|
||
<geoCoordinate id="EE285047B64AFFDCFDA6FEACFD3D777E" box="[591,730,261,287]" degrees="1" direction="north" minutes="23" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="00" value="1.3833333">1°23´00´´ N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE285047B64AFFDCFD0EFEADFC66777E" box="[743,897,260,287]" degrees="78" direction="west" minutes="15" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="00" value="-78.25">78°15´00´´W</geoCoordinate>
|
||
):
|
||
<location id="8EC3605BB64AFFDCFC7DFEACFBFE777E" LSID="urn:lsid:plazi:treatment:03B58796B645FFDEFF7EFF30FAA27492:8EC3605BB64AFFDCFC7DFEACFBFE777E" box="[916,1049,261,287]" country="Colombia" county="Barbacoas" latitude="1.246" longLatPrecision="1" longitude="-78.11531" municipality="Tumaco-Pasto" name="Rain Forest" pageId="24" pageNumber="25" stateProvince="Narino">Rain Forest</location>
|
||
,
|
||
<quantity id="4CE49B65B64AFFDCFBCCFEADFB69777E" box="[1061,1166,260,287]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">
|
||
<elevation id="0031D1B3B64AFFDCFBCCFEADFB69777E" box="[1061,1166,260,287]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="500.0">500 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA21040B64AFFDCFB70FEACFA91777E" box="[1177,1398,261,287]" pageId="24" pageNumber="25" value="2009-09-14">
|
||
<collectingDate id="EFE6E9A8B64AFFDCFB70FEACFA91777E" box="[1177,1398,261,287]" pageId="24" pageNumber="25" value="2009-09-14">14 September 2009</collectingDate>
|
||
</date>
|
||
, D,
|
||
<collectorName id="26E95356B64AFFDCFF7EFE80FEC77722" box="[151,288,297,323]" pageId="24" pageNumber="25">Martinez, H</collectorName>
|
||
. Herreño, manual collection, male (ICN-Am-108);
|
||
<collectorName id="26E95356B64AFFDCFCB0FE80FBEB7722" box="[857,1036,297,323]" pageId="24" pageNumber="25">Vereda Altaquer</collectorName>
|
||
(
|
||
<geoCoordinate id="EE285047B64AFFDCFBF2FE81FB2F7722" box="[1051,1224,296,323]" degrees="1" direction="north" minutes="14" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="45.60" value="1.246">1°14´45.60´´ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE285047B64AFFDCFB3BFE81FA767722" box="[1234,1425,296,323]" degrees="78" direction="west" minutes="06" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="55.12" value="-78.11531">78°06´55.12´´W</geoCoordinate>
|
||
): on the ground,
|
||
<collectorName id="26E95356B64AFFDCFED6FEE4FE9D7706" box="[319,378,333,359]" pageId="24" pageNumber="25">April</collectorName>
|
||
15 of 2010, 1270masl, manual collection, female (ICN-Am-75)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB64AFFDCFF2EFED9FDDB77EA" box="[199,572,368,395]" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="8BA33680B64AFFDCFF2EFED9FDDB77EA" blockId="24.[151,1437,152,503]" box="[199,572,368,395]" pageId="24" pageNumber="25">
|
||
<emphasis id="B968EA92B64AFFDCFF2EFED9FE8477EA" bold="true" box="[199,355,368,395]" pageId="24" pageNumber="25">Distribution:</emphasis>
|
||
<collectingCountry id="F30B7610B64AFFDCFE83FED8FE3C77EA" box="[362,475,369,395]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49D8F862B64AFFDCFE0FFED8FDD077EA" box="[486,567,369,395]" country="Colombia" name="Narino" pageId="24" pageNumber="25">Nariño</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB64AFFDCFF2EFE3CFAD87796" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BA33680B64AFFDCFF2EFE3CFAD87796" blockId="24.[151,1437,152,503]" pageId="24" pageNumber="25">
|
||
<emphasis id="B968EA92B64AFFDCFF2EFE3CFEB577CE" bold="true" box="[199,338,405,431]" pageId="24" pageNumber="25">Etymology:</emphasis>
|
||
This species is dedicated to the Colombian native community Awá (Kwaiker or Awa-Kwaiker), which inhabit between the boundaries of
|
||
<collectingCountry id="F30B7610B64AFFDCFD8DFE10FD3577B2" box="[612,722,441,467]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30B7610B64AFFDCFCE0FE10FC8F77B2" box="[777,872,441,467]" name="Ecuador" pageId="24" pageNumber="25">Ecuador</collectingCountry>
|
||
, and some they settle in the
|
||
<typeStatus id="54A78822B64AFFDCFB4AFE10FB3477B2" box="[1187,1235,441,467]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality (Nariño). We consider that this species deserves to be considered as part of his natural richness. Name in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF636608B64AFFDCFF7EF821FCBA71A4" ID-DOI="http://doi.org/10.5281/zenodo.3907932" ID-Zenodo-Dep="3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1928,1953]" targetBox="[164,1423,534,1894]" targetPageId="24">
|
||
<paragraph id="8BA33680B64AFFDCFF7EF821FCBA71A4" blockId="24.[151,1437,1928,1989]" pageId="24" pageNumber="25">
|
||
<emphasis id="B968EA92B64AFFDCFF7EF821FEF871C0" bold="true" box="[151,287,1928,1953]" pageId="24" pageNumber="25">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C1C4D03B64AFFDCFECCF820FE0B71C1" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[293,492,1929,1953]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
|
||
<emphasis id="B968EA92B64AFFDCFECCF820FE0B71C1" box="[293,492,1929,1953]" italics="true" pageId="24" pageNumber="25">Heterophrynus awa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EA92B64AFFDCFE1BF820FDA571C0" bold="true" box="[498,578,1929,1953]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A25B57E9B64AFFDCFE1BF820FDA571C0" box="[498,578,1929,1953]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Male (ICN-Am-108): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral
|
||
<taxonomicName id="4C1C4D03B64AFFDCFAB0F821FF2171A4" authority="(Pv)" baseAuthorityName="Pv" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="24" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pv)</taxonomicName>
|
||
. D. Dorsal
|
||
<taxonomicName id="4C1C4D03B64AFFDCFED1F805FE5671A4" authority="(Pd)" baseAuthorityName="Pd" box="[312,433,1964,1989]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="24" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pd)</taxonomicName>
|
||
. E. Ventral tibia (Tv). F. Dorsal tibia (Td).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C306650BB64BFFDDFF2EFF31FCE47402" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="8BA33680B64BFFDDFF2EFF31FBB2769A" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFF31FEA576D2" bold="true" box="[199,322,152,179]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C1C4D03B64BFFDDFEA1FF30FDC276D2" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[328,549,153,179]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
|
||
<emphasis id="B968EA92B64BFFDDFEA1FF30FDC276D2" box="[328,549,153,179]" italics="true" pageId="25" pageNumber="26">Heterophrynus awa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EA92B64BFFDDFDC2FF30FD6276D2" bold="true" box="[555,645,153,179]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A25B57E9B64BFFDDFDC2FF30FD6276D2" box="[555,645,153,179]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from other species by present the following characters together: four teeth in the ectal portion of the basal segment of the chelicerae. Five long spines in the ventral femur, spine FIII is smaller than FII and FV, FIV is small, presents a small spine PVI.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFEACFE907709" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
Female genitalia with gonopod sclerites elongated and with laminar shape, a medial portion wider than the basal and distal portion, distal portion thick and with a rounded apex. Seminal receptacles oval with a sclerotized area at the base (
|
||
<figureCitation id="13272A05B64BFFDDFEE8FEE4FE8D7709" box="[257,362,333,360]" captionStart="FIGURE 16" captionStartId="26.[151,250,1910,1935]" captionTargetBox="[156,1432,784,1880]" captionTargetId="figure@26.[151,1437,777,1886]" captionTargetPageId="26" captionText="FIGURE 16. Male genital: Heterophrynus awa sp. nov. (ICN-Am-108): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (ICN-Am-75), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907934" httpUri="https://zenodo.org/record/3907934/files/figure.png" pageId="25" pageNumber="26">Fig. 16D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFED8FC607799" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
Similar species: This species is similar to
|
||
<taxonomicName id="4C1C4D03B64BFFDDFD41FEDBFCE177EA" authorityName="Pocock" authorityYear="1902" box="[680,774,369,395]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alces">
|
||
<emphasis id="B968EA92B64BFFDDFD41FEDBFCE177EA" box="[680,774,369,395]" italics="true" pageId="25" pageNumber="26">H. alces</emphasis>
|
||
</taxonomicName>
|
||
but it presents a small spine TIII and T3, which is absent in
|
||
<taxonomicName id="4C1C4D03B64BFFDDFF5DFE3FFEE177CE" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[180,262,406,431]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
|
||
<emphasis id="B968EA92B64BFFDDFF5DFE3FFEE177CE" box="[180,262,406,431]" italics="true" pageId="25" pageNumber="26">H. awa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EA92B64BFFDDFEE4FE3CFE8077CE" bold="true" box="[269,359,405,431]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A25B57E9B64BFFDDFEE4FE3CFE8077CE" box="[269,359,405,431]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the female genitalia is completely different, presenting the sclerites of the gonopods with a rounded shape. The species
|
||
<taxonomicName id="4C1C4D03B64BFFDDFE38FE13FDA977B2" authorityName="Pocock" authorityYear="1902" box="[465,590,442,467]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="armiger">
|
||
<emphasis id="B968EA92B64BFFDDFE38FE13FDA977B2" box="[465,590,442,467]" italics="true" pageId="25" pageNumber="26">H. armiger</emphasis>
|
||
</taxonomicName>
|
||
is very similar, however, it presents female genitalia with sclerites not widened at the medial portion, and with an acuminated apex (
|
||
<figureCitation id="13272A05B64BFFDDFCF6FE74FC9D7799" box="[799,890,477,504]" captionStart="FIGURE 6" captionStartId="11.[151,250,1755,1780]" captionTargetBox="[155,1433,529,1726]" captionTargetId="figure@11.[151,1436,525,1730]" captionTargetPageId="11" captionText="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="25" pageNumber="26">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFDA8FCE47402" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
The male genital of
|
||
<taxonomicName id="4C1C4D03B64BFFDDFE41FDABFDC3747A" authorityName="Pocock" authorityYear="1902" box="[424,548,514,539]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="armiger">
|
||
<emphasis id="B968EA92B64BFFDDFE41FDABFDC3747A" box="[424,548,514,539]" italics="true" pageId="25" pageNumber="26">H. armiger</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D03B64BFFDDFDB3FDABFD4B747A" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[602,684,514,539]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
|
||
<emphasis id="B968EA92B64BFFDDFDB3FDABFD4B747A" box="[602,684,514,539]" italics="true" pageId="25" pageNumber="26">H. awa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EA92B64BFFDDFD5BFDA8FCE1747A" bold="true" box="[690,774,513,539]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A25B57E9B64BFFDDFD5BFDA8FCE1747A" box="[690,774,513,539]" pageId="25" pageNumber="26" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
, presents Pi rounded and so smaller than the LaM, the LoL 1 has a similar length with the LaM. Both species differ because
|
||
<taxonomicName id="4C1C4D03B64BFFDDFC69FD8FFC19745E" authorityName="Pocock" authorityYear="1902" box="[896,1022,550,575]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="armiger">
|
||
<emphasis id="B968EA92B64BFFDDFC69FD8FFC19745E" box="[896,1022,550,575]" italics="true" pageId="25" pageNumber="26">H. armiger</emphasis>
|
||
</taxonomicName>
|
||
presents a spine on LOL 1 which is absent in
|
||
<taxonomicName id="4C1C4D03B64BFFDDFEEBFDE3FEB37402" authority="Chirivi-Joya & Moreno-González & Fagua, 2020" authorityName="Chirivi-Joya & Moreno-González & Fagua" authorityYear="2020" box="[258,340,586,611]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="awa" status="sp. nov.">
|
||
<emphasis id="B968EA92B64BFFDDFEEBFDE3FEB37402" box="[258,340,586,611]" italics="true" pageId="25" pageNumber="26">H. awa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EA92B64BFFDDFEB3FDE0FE487402" bold="true" box="[346,431,585,611]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A25B57E9B64BFFDDFEB3FDE0FE487402" box="[346,431,585,611]" pageId="25" pageNumber="26" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13272A05B64BFFDDFE57FDE0FDBE7402" box="[446,601,585,611]" captionStart="FIGURE 6" captionStartId="11.[151,250,1755,1780]" captionTargetBox="[155,1433,529,1726]" captionTargetId="figure@11.[151,1436,525,1730]" captionTargetPageId="11" captionText="FIGURE 6. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MUSENUV 24303), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907913" httpUri="https://zenodo.org/record/3907913/files/figure.png" pageId="25" pageNumber="26">Fig. 6A, B, C</figureCitation>
|
||
;
|
||
<figureCitation id="13272A05B64BFFDDFD8DFDE0FD937402" box="[612,628,585,611]" captionStart="FIGURE 7" captionStartId="13.[151,250,670,695]" captionTargetBox="[152,1435,181,644]" captionTargetId="figure@13.[152,1436,181,644]" captionTargetPageId="13" captionText="FIGURE 7. Male genital: Heterophrynus armiger. (IAvH-I 2924): A. Spine over LoL1, dorsal view. B. Detail of spine, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3907916" httpUri="https://zenodo.org/record/3907916/files/figure.png" pageId="25" pageNumber="26">7</figureCitation>
|
||
;
|
||
<figureCitation id="13272A05B64BFFDDFD68FDE0FD137402" box="[641,756,585,611]" captionStart="FIGURE 16" captionStartId="26.[151,250,1910,1935]" captionTargetBox="[156,1432,784,1880]" captionTargetId="figure@26.[151,1437,777,1886]" captionTargetPageId="26" captionText="FIGURE 16. Male genital: Heterophrynus awa sp. nov. (ICN-Am-108): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (ICN-Am-75), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907934" httpUri="https://zenodo.org/record/3907934/files/figure.png" pageId="25" pageNumber="26">16A, B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB64BFFDEFF2EFDC5FAA27492" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8BA33680B64BFFDDFF2EFDC5FB7174CA" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFDC5FDBD74E6" bold="true" box="[199,602,620,647]" pageId="25" pageNumber="26">Description. Male (ICN-Am-108):</emphasis>
|
||
Total length
|
||
<quantity id="4CE49B65B64BFFDDFD07FDC4FCB274E6" box="[750,853,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="25" pageNumber="26" unit="mm" value="20.9">20.9 mm</quantity>
|
||
. Carapace, opisthosoma, and legs have a brown coloration, with red tonalities, pedipalps brown. Measures of the male are provided in
|
||
<tableCitation id="C69E033BB64BFFDDFBD5FD38FB7674CA" box="[1084,1169,656,683]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="25" pageNumber="26">Table 5</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFD1CFD8C7576" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFD1CFEDB74AE" bold="true" box="[199,316,693,719]" italics="true" pageId="25" pageNumber="26">Carapace.</emphasis>
|
||
Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin with a concave area in the medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (
|
||
<figureCitation id="13272A05B64BFFDDFE1FFD54FDB97576" box="[502,606,765,791]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFC88FD4E75E2" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFC88FED6755A" bold="true" box="[199,305,801,827]" italics="true" pageId="25" pageNumber="26">Sternum.</emphasis>
|
||
Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with more than 10 setae in the basal region, five in the medial region three in distal region and two more in the apex.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFC24FB80758E" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
The second segment (tetrasternum) rounded and with four setae; two on the medial region and two on the apical region. The third segment (pentasternum) rounded, smaller than the second segment, has two setae on medial region. Metasternum longitudinally divided with two setae in each segment (
|
||
<figureCitation id="13272A05B64BFFDDFC1AFC7CFBBD758E" box="[1011,1114,981,1007]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFC51FD7A7256" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFC51FEDB7272" bold="true" box="[199,316,1016,1043]" italics="true" pageId="25" pageNumber="26">Abdomen.</emphasis>
|
||
Oblong, color brown, with soft punctuations, and darker areas in the medial area of the sclerites. Carapace wider than the Abdomen (
|
||
<figureCitation id="13272A05B64BFFDDFDC3FBB4FD767256" box="[554,657,1053,1079]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFBE9FA71728A" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFBE9FED9723A" bold="true" box="[199,318,1088,1115]" italics="true" pageId="25" pageNumber="26">Chelicera.</emphasis>
|
||
Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (
|
||
<figureCitation id="13272A05B64BFFDDFE25FB20FDD272C2" box="[460,565,1161,1187]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14D</figureCitation>
|
||
). The ectal row of the basal segment of the chelicerae have four conspicuous teeth, two with acuminated apex placed in the proximal and medial portions (tooth 1 and 2), and two with acuminated apex placed on the medial portion (tooth 3 and 4) (
|
||
<figureCitation id="13272A05B64BFFDDFCFAFB78FC90728A" box="[787,887,1233,1259]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14F</figureCitation>
|
||
). Mobile finger of the chelicerae with five teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EFB5DFE0E7112" blockId="25.[151,1437,152,2015]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EFB5DFECB736E" bold="true" box="[199,300,1268,1295]" italics="true" pageId="25" pageNumber="26">Pedipalp</emphasis>
|
||
. Trochanter: Prolateral face with four spines; spines Tr1 and Tr3 placed in the medial region, Tr2 placed near to ventral margin. Tr3 and Tr4 with subequal lengths. Spines lengths: Tr2>Tr1>Tr4≥Tr3. Dorsal oblique series of six setal tubercles. The dorsomedial area without spines but with one tubercle (
|
||
<figureCitation id="13272A05B64BFFDDFBFCFA94FB9E7336" box="[1045,1145,1341,1367]" captionStart="FIGURE 14" captionStartId="23.[151,250,1934,1959]" captionTargetBox="[157,1426,315,1904]" captionTargetId="figure@23.[151,1436,309,1909]" captionTargetPageId="23" captionText="FIGURE 14. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907930" httpUri="https://zenodo.org/record/3907930/files/figure.png" pageId="25" pageNumber="26">Fig. 14E</figureCitation>
|
||
). Femur: Ventral face with five major spines, there are three tubercles between FII–FIII, one small spine FIV with two tubercles at each side between FIII–FV, a small spine with three tubercles at each side between FV–FVI, ten tubercles more distally than FVI, FIV has almost the size of a tubercle. Spine lengths: FI>FII>FV>FVI>FIII>FIV (
|
||
<figureCitation id="13272A05B64BFFDDFB8FFA00FB2873A2" box="[1126,1231,1448,1475]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15A</figureCitation>
|
||
). Dorsal face with five major spines, there are three tubercles between F2–F3, a very small spine and four tubercles between F3–F4, two very small spines and three tubercles between F4–F5, and eight tubercles more distally than F5, F1 and F2 are joined at the base, F3 clearly separated from F2. Spine lengths: F3>F4>F1=F2>F5 (
|
||
<figureCitation id="13272A05B64BFFDDFBA8F9BCFB4F704E" box="[1089,1192,1556,1583]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1C4D03B64BFFDDFB52F9BCFA80704E" authority=": Ventral" authorityName="Ventral" box="[1211,1383,1557,1583]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">Patella: Ventral</taxonomicName>
|
||
face with six major spines, presents a small PVI, there are three tubercles more proximally than PVI, one small spine and three tubercles between PVI–PV, four tubercles between PV–PIV, and PIV–PIII, three tubercles between PIII–PII and five between PII–PI, PI wider than the other spines. Spine lengths: PIV>PII>PV>PI>PIII>PVI (
|
||
<figureCitation id="13272A05B64BFFDDFB06F928FAB270FA" box="[1263,1365,1664,1691]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15C</figureCitation>
|
||
). Dorsal face with six major spines, seven tubercles between P6–P5, P7 is absent. Spine lengths: P3=P4≥P5>P6>P1>P2 (
|
||
<figureCitation id="13272A05B64BFFDDFF49F960FEE07082" box="[160,263,1736,1763]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15D</figureCitation>
|
||
).
|
||
<emphasis id="B968EA92B64BFFDDFEF3F960FEB37082" box="[282,340,1737,1763]" italics="true" pageId="25" pageNumber="26">Tibia</emphasis>
|
||
: Ventral face with two major spines. Spine lengths: TII>TI (
|
||
<figureCitation id="13272A05B64BFFDDFC1DF960FBBE7082" box="[1012,1113,1736,1763]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15E</figureCitation>
|
||
). Dorsal face with two major spines. Spine lengths: T2>T1 (
|
||
<figureCitation id="13272A05B64BFFDDFE1AF944FDB07166" box="[499,599,1772,1799]" captionStart="FIGURE 15" captionStartId="24.[151,250,1928,1953]" captionTargetBox="[164,1423,534,1894]" captionTargetId="figure@24.[151,1436,525,1903]" captionTargetPageId="24" captionText="FIGURE 15. Heterophrynus awa sp. nov. Male (ICN-Am-108):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907932" httpUri="https://zenodo.org/record/3907932/files/figure.png" pageId="25" pageNumber="26">Fig. 15F</figureCitation>
|
||
).
|
||
<emphasis id="B968EA92B64BFFDDFD84F947FCA37166" box="[621,836,1774,1799]" italics="true" pageId="25" pageNumber="26">Tarsus-metatarsus:</emphasis>
|
||
Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible.
|
||
<emphasis id="B968EA92B64BFFDDFC1AF8B8FBCE714A" bold="true" box="[1011,1065,1809,1835]" italics="true" pageId="25" pageNumber="26">Legs</emphasis>
|
||
. Femora brown. Femora lengths: I>III>II>IV (
|
||
<tableCitation id="C69E033BB64BFFDDFEC3F89CFE65712E" box="[298,386,1844,1871]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="25" pageNumber="26">Table 5</tableCitation>
|
||
). Leg I: Tibia with 31 segments and tarsus with 70 segments in the right and left leg. Leg IV: Basitibia with three segments.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B64BFFDDFF2EF8D4FA8571F6" blockId="25.[151,1437,152,2015]" box="[199,1378,1917,1943]" pageId="25" pageNumber="26">Basitibia-distitibia lengths: BT1>DT>BT3>BT2. Basitarsus as long as the telotarsus. Tarsus tetramerous.</paragraph>
|
||
<paragraph id="8BA33680B64BFFDEFF2EF809FCFE7741" blockId="25.[151,1437,152,2015]" lastBlockId="26.[151,1437,153,755]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EF809FDDA71DA" bold="true" box="[199,573,1952,1979]" pageId="25" pageNumber="26">
|
||
<emphasis id="B968EA92B64BFFDDFF2EF809FE6C71DA" bold="true" box="[199,395,1952,1979]" italics="true" pageId="25" pageNumber="26">Female genitalia</emphasis>
|
||
(ICN-Am-75):
|
||
</emphasis>
|
||
Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with an oval shape, completely separated at the medial portion, dorsal surface sclerotized at the proximal region, with brown coloration, medial and distal portion white. Sclerites with laminar shape, the medial region is wider than the basal and distal region, distal portion a little curved and thinner than the basal portion, with rounded apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites longer than the total length of the seminal receptacle (
|
||
<figureCitation id="13272A05B648FFDEFD4DFEACFCEB7741" box="[676,780,261,288]" captionStart="FIGURE 16" captionStartId="26.[151,250,1910,1935]" captionTargetBox="[156,1432,784,1880]" captionTargetId="figure@26.[151,1437,777,1886]" captionTargetPageId="26" captionText="FIGURE 16. Male genital: Heterophrynus awa sp. nov. (ICN-Am-108): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (ICN-Am-75), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907934" httpUri="https://zenodo.org/record/3907934/files/figure.png" pageId="26" pageNumber="27">Fig. 16D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B648FFDEFF2EFE81FA7A7722" blockId="26.[151,1437,153,755]" box="[199,1437,296,323]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF2EFE81FED87722" bold="true" box="[199,319,296,323]" pageId="26" pageNumber="27">Variation.</emphasis>
|
||
Variation in measures and segments of the basitibia IV, Tibia I and tarsus I, are provided in
|
||
<tableCitation id="C69E033BB648FFDEFAABFE80FA707722" box="[1346,1431,296,323]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="26" pageNumber="27">Table 5</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B648FFDEFF7EFEE4FD4E745E" blockId="26.[151,1437,153,755]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF7EFEE4FEE57706" bold="true" box="[151,258,333,359]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF7EFEE4FF1C7706" bold="true" box="[151,251,333,359]" italics="true" pageId="26" pageNumber="27">Sternum</emphasis>
|
||
.
|
||
</emphasis>
|
||
The number of setae over all segments of the sternum was variable.
|
||
<emphasis id="B968EA92B648FFDEFBFFFEE5FB9C7706" bold="true" box="[1046,1147,332,359]" italics="true" pageId="26" pageNumber="27">Pedipalp</emphasis>
|
||
.
|
||
<emphasis id="B968EA92B648FFDEFB62FEE4FAE17706" box="[1163,1286,333,359]" italics="true" pageId="26" pageNumber="27">Trochanter</emphasis>
|
||
: line of setal tubercles vary between five and six.
|
||
<emphasis id="B968EA92B648FFDEFDDCFEDBFD6777EA" box="[565,640,370,395]" italics="true" pageId="26" pageNumber="27">Femur</emphasis>
|
||
: Ventral face: FIV can be present as a spine or tubercle, number of tubercles between spines is variable, and can be more conspicuous. Dorsal face: number of tubercles between spines is variable, tubercles can be more conspicuous.
|
||
<taxonomicName id="4C1C4D03B648FFDEFD5AFE10FCE377B2" box="[691,772,441,467]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EA92B648FFDEFD5AFE10FCE377B2" box="[691,772,441,467]" italics="true" pageId="26" pageNumber="27">Patella</emphasis>
|
||
</taxonomicName>
|
||
. Ventral face: relative size between PI and PIII is variable, number and size of small spines and tubercles between major spines are variable. Dorsal face: number of tubercles are variable.
|
||
<emphasis id="B968EA92B648FFDEFECEFDA8FE82747A" bold="true" box="[295,357,513,539]" italics="true" pageId="26" pageNumber="27">Legs.</emphasis>
|
||
Leg I: Number of segments of tibia vary between 31 and 33 and in the tarsus between 66 and 76, the relative size of femora is variable (
|
||
<tableCitation id="C69E033BB648FFDEFDACFD8CFD7C745E" box="[581,667,548,575]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="26" pageNumber="27">Table 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B648FFDEFF2EFDE0FBAC74E6" blockId="26.[151,1437,153,755]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF2EFDE0FEEC7402" bold="true" box="[199,267,585,611]" pageId="26" pageNumber="27">Male.</emphasis>
|
||
Observed Males were bigger than females (
|
||
<tableCitation id="C69E033BB648FFDEFD1EFDE0FCA97402" box="[759,846,584,611]" captionStart="TABLE 5" captionStartId="27.[151,239,154,178]" captionText="TABLE 5. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus awa sp. nov. Empty cells correspond to missing segment or data." pageId="26" pageNumber="27">Table 5</tableCitation>
|
||
). Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B648FFDEFF2EFD39FAA27492" blockId="26.[151,1437,153,755]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF2EFD39FE9274CA" bold="true" box="[199,373,656,683]" pageId="26" pageNumber="27">
|
||
<emphasis id="B968EA92B648FFDEFF2EFD39FE8974CA" bold="true" box="[199,366,656,683]" italics="true" pageId="26" pageNumber="27">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
(ICN-Am-108): LaM longer than Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In dorsal view, the LoD are, long, rounded and do not cover LoL1 and LoL 2 (
|
||
<figureCitation id="13272A05B648FFDEFB66FD70FADF7492" box="[1167,1336,729,755]" captionStart="FIGURE 16" captionStartId="26.[151,250,1910,1935]" captionTargetBox="[156,1432,784,1880]" captionTargetId="figure@26.[151,1437,777,1886]" captionTargetPageId="26" captionText="FIGURE 16. Male genital: Heterophrynus awa sp. nov. (ICN-Am-108): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (ICN-Am-75), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907934" httpUri="https://zenodo.org/record/3907934/files/figure.png" pageId="26" pageNumber="27">Fig. 16A, B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |