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<document id="72C60B58B0A98AF7FE4EB1F69EA667F2" ID-DOI="10.11646/zootaxa.4743.2.1" ID-GBIF-Dataset="8eba416f-e2af-4cfe-96fb-fb6059a877ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3687704" ID-ZooBank="624E410F-A208-46D4-BF6B-5331B39582F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582709117521" checkinUser="plazi" docAuthor="Joharchi, Omid, Ermilov, Sergey G. &amp; Khaustov, Alexander A." docDate="2020" docId="03B58785DE0A1E345EA471B9FC0D82F2" docLanguage="en" docName="zootaxa.4743.2.1.pdf" docOrigin="Zootaxa 4743 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps trichiurus Joharchi &amp; Ermilov &amp; Khaustov 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="164" masterDocId="FF8CFFFDDE021E395E337475FFD28138" masterDocTitle="Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka" masterLastPageNumber="166" masterPageNumber="151" pageNumber="159" updateTime="1698721478006" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="41D4F7DFDBBBEB30DE4BB80500DB7EE9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="20263ED9653AA1E7BF16FFA6AE757B06">
<mods:title id="95B312503629B9E32BABA8551D666801">Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka</mods:title>
</mods:titleInfo>
<mods:name id="C8E73EA0037C608EE39C5B8235AA3EF5" type="personal">
<mods:role id="A230EF086DBB2ABCDB211E16077B70E1">
<mods:roleTerm id="2C24308EFDE7157B15973EAA19D5B317">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BB5EB6DF2021CC99266389CB16154B5">Joharchi, Omid</mods:namePart>
</mods:name>
<mods:name id="AF072D5AE10FCE7ABBA96A98991D746C" type="personal">
<mods:role id="7CE3C958BB6B13ABA5190D43075B4630">
<mods:roleTerm id="61ED83E991C98786F101ED1F32F838A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4043CC7DA4C1636605808D80DDE324A">Ermilov, Sergey G.</mods:namePart>
</mods:name>
<mods:name id="C776C98EA0654B4EEA2F5B8A632B4EDD" type="personal">
<mods:role id="DCF97EE9298CAB0EB615C8B248A27D4B">
<mods:roleTerm id="C289B8311DF9AE337C3155E06C2B1F38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C99079931DA7C4E9D179C5E478F5316">Khaustov, Alexander A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="504C1B4FA0CA4CCDBAB0A34DADC6E21D">text</mods:typeOfResource>
<mods:relatedItem id="7284CE308A776B2BBAB12104766AE3BD" type="host">
<mods:titleInfo id="F3154CED64BA36F7E27378AB452B0857">
<mods:title id="D669B2CB44951CA32B33A75C8C6B2553">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F34A035FB4D570580505BA38F33F0CE2">
<mods:date id="B5C4F0E22E4E482B7E0555563899F402">2020</mods:date>
<mods:detail id="DB3571020AF0EF7C2A8F49638646A98D" type="pubDate">
<mods:number id="470CBC55FA2B51633A81345DE803A96F">2020-02-25</mods:number>
</mods:detail>
<mods:detail id="D2CEF6CE925AC463DEE5AFCA43E4A753" type="volume">
<mods:number id="E75669D2983DF5CE82759D44C2C28453">4743</mods:number>
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<mods:detail id="C077EAD5FAA14F9C5ECEB851106EC382" type="issue">
<mods:number id="776BC3A04DD1178DA764B2CF3CDD7964">2</mods:number>
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<mods:start id="6D586BD658B0EE1EC775DBEAD7F7467A">151</mods:start>
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<mods:classification id="9F686CE82AFA2620C7176E7DF2D2E101">journal article</mods:classification>
<mods:identifier id="2441EE67B44E0CCE51BC9E22CA5ED984" type="DOI">10.11646/zootaxa.4743.2.1</mods:identifier>
<mods:identifier id="0D9A59A88C8FB91D3C5AFBE6C3A998A5" type="GBIF-Dataset">8eba416f-e2af-4cfe-96fb-fb6059a877ef</mods:identifier>
<mods:identifier id="7DB8EE99E9F31B8DD1B0D085F2917D3F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2E889D9E2F11865120F9F018DA5F8746" type="Zenodo-Dep">3687704</mods:identifier>
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<treatment id="03B58785DE0A1E345EA471B9FC0D82F2" ID-DOI="http://doi.org/10.5281/zenodo.3688016" ID-GBIF-Taxon="162267487" ID-Zenodo-Dep="3688016" LSID="urn:lsid:plazi:treatment:03B58785DE0A1E345EA471B9FC0D82F2" httpUri="http://treatment.plazi.org/id/03B58785DE0A1E345EA471B9FC0D82F2" lastPageId="13" lastPageNumber="164" pageId="8" pageNumber="159">
<subSubSection id="C3066518DE0A1E315EA471B9FDF684DF" box="[151,548,1484,1511]" pageId="8" pageNumber="159" type="nomenclature">
<paragraph id="8BA33693DE0A1E315EA471B9FDF684DF" blockId="8.[151,548,1484,1547]" box="[151,548,1484,1511]" pageId="8" pageNumber="159">
<heading id="D0EB81FFDE0A1E315EA471B9FDF684DF" bold="true" box="[151,548,1484,1511]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
<emphasis id="B968EA81DE0A1E315EA471B9FDF684DF" bold="true" box="[151,548,1484,1511]" pageId="8" pageNumber="159">
<taxonomicName id="4C1C4D10DE0A1E315EA471B9FE6D84DF" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[151,447,1484,1511]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="trichiurus" status="sp. nov.">
<emphasis id="B968EA81DE0A1E315EA471B9FE6D84DF" bold="true" box="[151,447,1484,1511]" italics="true" pageId="8" pageNumber="159">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57FADE0A1E315FF571B8FDF684DF" box="[454,548,1485,1511]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0A1E315EA47185FE998733" box="[151,331,1520,1547]" pageId="8" pageNumber="159" type="description">
<paragraph id="8BA33693DE0A1E315EA47185FE998733" blockId="8.[151,548,1484,1547]" box="[151,331,1520,1547]" pageId="8" pageNumber="159">
(
<figureCitation id="13272A16DE0A1E315EAC7185FE918733" box="[159,323,1520,1547]" captionStart-0="FIGURES 25–29" captionStart-1="FIGURES 30–36" captionStart-2="FIGURE 37–40" captionStartId-0="10.[151,264,1945,1970]" captionStartId-1="11.[151,264,1946,1971]" captionStartId-2="12.[151,250,1911,1936]" captionTargetBox-0="[190,1395,194,1908]" captionTargetBox-1="[182,1407,184,1918]" captionTargetBox-2="[166,1432,190,1874]" captionTargetId-0="figure@10.[178,1410,181,1919]" captionTargetId-1="figure@11.[180,1407,181,1920]" captionTargetId-2="figure@12.[151,1436,181,1885]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." captionText-1="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." captionText-2="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.3687716" figureDoi-1="http://doi.org/10.5281/zenodo.3687718" figureDoi-2="http://doi.org/10.5281/zenodo.3687720" httpUri-0="https://zenodo.org/record/3687716/files/figure.png" httpUri-1="https://zenodo.org/record/3687718/files/figure.png" httpUri-2="https://zenodo.org/record/3687720/files/figure.png" pageId="8" pageNumber="159">Figures 25–40</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0A1E315EA4724CFB8C87A3" pageId="8" pageNumber="159" type="materials_examined">
<paragraph id="8BA33693DE0A1E315EA4724CFB8C87A3" blockId="8.[151,1437,1592,2043]" pageId="8" pageNumber="159">
<emphasis id="B968EA81DE0A1E315EA4724CFE9B876B" bold="true" box="[151,329,1592,1619]" pageId="8" pageNumber="159">Type material:</emphasis>
<materialsCitation id="3B743CCEDE0A1E315F62724DFE1587A3" ID-GBIF-Occurrence="2571176106" collectingDate="2019-01-24" collectionCode="TSUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="8" pageNumber="159" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<typeStatus id="54A78831DE0A1E315F62724DFE6E876A" box="[337,444,1592,1618]" pageId="8" pageNumber="159" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="F30B7603DE0A1E315C17724DFD44876A" box="[548,662,1592,1618]" name="Sri Lanka" pageId="8" pageNumber="159">Sri Lanka</collectingCountry>
,
<collectingRegion id="49D8F871DE0A1E315C91724DFC69876A" box="[674,955,1592,1618]" country="Sri Lanka" name="Sabaragamuwa" pageId="8" pageNumber="159">Sabaragamuwa Province</collectingRegion>
,
<location id="8EC36048DE0A1E315DFB724DFB83876A" LSID="urn:lsid:plazi:treatment:03B58785DE0A1E345EA471B9FC0D82F2:8EC36048DE0A1E315DFB724DFB83876A" box="[968,1105,1592,1618]" country="Sri Lanka" latitude="6.45" longLatPrecision="1304" longitude="80.2" name="Polgampola" pageId="8" pageNumber="159" stateProvince="Sabaragamuwa">Polgampola</location>
,
<geoCoordinate id="EE285054DE0A1E315A6E724DFB15876B" box="[1117,1223,1592,1619]" degrees="06" direction="north" minutes="27" orientation="latitude" pageId="8" pageNumber="159" precision="925" value="6.45">06°27’ N</geoCoordinate>
,
<geoCoordinate id="EE285054DE0A1E315AE7724CFA95876B" box="[1236,1351,1593,1619]" degrees="080" direction="east" minutes="12" orientation="longitude" pageId="8" pageNumber="159" precision="925" value="80.2">080°12’ E</geoCoordinate>
, alt.
<quantity id="4CE49B76DE0A1E315BB3724CFF7D874F" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="159" unit="m" value="42.0">
<elevation id="0031D1A0DE0A1E315BB3724CFF7D874F" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="159" unit="m" value="42.0">42 m</elevation>
</quantity>
,
<date id="FFA21053DE0A1E315E887228FEAA874F" box="[187,376,1629,1655]" pageId="8" pageNumber="159" value="2019-01-24">
<collectingDate id="EFE6E9BBDE0A1E315E887228FEAA874F" box="[187,376,1629,1655]" pageId="8" pageNumber="159" value="2019-01-24">24 January 2019</collectingDate>
</date>
,
<collectorName id="26E95345DE0A1E315FB77228FDDB874E" box="[388,521,1628,1655]" pageId="8" pageNumber="159">O. Joharchi</collectorName>
,
<collectorName id="26E95345DE0A1E315C277228FD7B874E" box="[532,681,1628,1655]" pageId="8" pageNumber="159">S.G. Ermilov</collectorName>
&amp;
<collectorName id="26E95345DE0A1E315CFF7228FCA7874E" box="[716,885,1628,1655]" pageId="8" pageNumber="159">A.A. Khaustov</collectorName>
coll., from forest soil-litter near the
<collectorName id="26E95345DE0A1E315B237229FF1687A2" pageId="8" pageNumber="159">Thambadola Ella</collectorName>
waterfall (in
<collectionCode id="ED0DAE56DE0A1E315F6872F4FE6D87A3" box="[347,447,1665,1691]" pageId="8" pageNumber="159">TSUMZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B743CCEDE0A1E315FE172F4FB8887A3" ID-GBIF-Occurrence="2571176104" box="[466,1114,1664,1691]" collectingDate="2019-01-24" collectionCode="TSUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="8" pageNumber="159" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="paratype">
<typeStatus id="54A78831DE0A1E315FE172F4FDE587A3" box="[466,567,1665,1691]" pageId="8" pageNumber="159" type="paratype">Paratype</typeStatus>
, one female, same data as holotype (in
<collectionCode id="ED0DAE56DE0A1E315DDD72F4FB8087A3" box="[1006,1106,1665,1691]" pageId="8" pageNumber="159">TSUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0A1E305EF472D1FEF68027" lastPageId="9" lastPageNumber="160" pageId="8" pageNumber="159" type="diagnosis">
<paragraph id="8BA33693DE0A1E315EF472D1FD2686C3" blockId="8.[151,1437,1592,2043]" pageId="8" pageNumber="159">
<emphasis id="B968EA81DE0A1E315EF472D1FEE88787" bold="true" box="[199,314,1700,1727]" pageId="8" pageNumber="159">Diagnosis</emphasis>
(adult female). Dorsal shield covering most of the idiosoma, with reticulate ornamentation over whole surface, narrowing from level of setae
<emphasis id="B968EA81DE0A1E315CAB72BCFD6087DB" box="[664,690,1737,1763]" italics="true" pageId="8" pageNumber="159">r3</emphasis>
, progressively tapering until rounded posteriorly, bearing 39 pairs setae. Most setae long enough to reach well past base of next posterior seta, all setae abruptly taper apically (
<figureCitation id="13272A16DE0A1E315B597299FF678613" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="8" pageNumber="159">Figs 25</figureCitation>
,
<figureCitation id="13272A16DE0A1E315EF17364FEDF8613" box="[194,269,1809,1835]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="8" pageNumber="159">30, 32</figureCitation>
), except
<emphasis id="B968EA81DE0A1E315F407364FE5B8613" box="[371,393,1809,1835]" italics="true" pageId="8" pageNumber="159">j1</emphasis>
,
<emphasis id="B968EA81DE0A1E315FAB7364FE638613" box="[408,433,1809,1835]" italics="true" pageId="8" pageNumber="159">z1</emphasis>
,
<emphasis id="B968EA81DE0A1E315FF27364FE0C8613" box="[449,478,1809,1835]" italics="true" pageId="8" pageNumber="159">Z5</emphasis>
very short (7–10) and
<emphasis id="B968EA81DE0A1E315CD77364FCD28613" box="[740,768,1809,1835]" italics="true" pageId="8" pageNumber="159">S5</emphasis>
longest (44–46). Sternal shield smooth almost throughout; anterior margin indistinct and posterior margin almost straight, bearing three pairs of setae and two pairs of large slit-shape pore-like structures (
<figureCitation id="13272A16DE0A1E315FCA732DFD9D864B" box="[505,591,1880,1907]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="8" pageNumber="159">Fig. 33</figureCitation>
), ratio of shield length/width (at broadest level) 1. Genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield≃ (
<figureCitation id="13272A16DE0A1E315ABD7309FB3486AE" box="[1166,1254,1916,1942]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="8" pageNumber="159">Figs 26</figureCitation>
&amp;
<figureCitation id="13272A16DE0A1E315B227308FAFD86AF" box="[1297,1327,1917,1943]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="8" pageNumber="159">31</figureCitation>
), ratio of length/width (at broadest level) 3.2, surface without any distinct reticulation, almost completely smooth (
<figureCitation id="13272A16DE0A1E315B0E73D5FA428683" box="[1341,1424,1952,1979]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="8" pageNumber="159">Fig. 34</figureCitation>
). Anal shield ratio of length/width≃(at broadest level) 1.8 and cribrum small, length of post-anal seta almost twice ≃
</paragraph>
<paragraph id="8BA33693DE0B1E305EA474ECFEF68027" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="160">para-anal setae. Peritreme long, extending to posterior of coxa I, post-stigmatal section of peritrematal shield oval, its tip extending to mid-level of coxa IV. Fixed digit of chelicera with six teeth in two rows, dorsal cheliceral setae thickened, arthrodial membrane reduced to a lobe and without filaments. Femur I and genu III with 12 and 8 setae, respectively.</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0B1E345EF4755CFD96818A" lastPageId="13" lastPageNumber="164" pageId="9" pageNumber="160" type="description">
<paragraph id="8BA33693DE0B1E305EF4755CFA40835B" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF4755CFDDA807B" bold="true" box="[199,520,296,323]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF4755CFE86807B" bold="true" box="[199,340,297,323]" italics="true" pageId="9" pageNumber="160">Description.</emphasis>
Female (n= 2).
</emphasis>
<emphasis id="B968EA81DE0B1E305C3C755CFD19807B" box="[527,715,297,323]" italics="true" pageId="9" pageNumber="160">Dorsal idiosoma</emphasis>
(
<figureCitation id="13272A16DE0B1E305CE9755CFCE3807B" box="[730,817,296,323]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figs 25</figureCitation>
,
<figureCitation id="13272A16DE0B1E305D0E755CFC4F807B" box="[829,925,297,323]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">30 &amp; 32</figureCitation>
). Dorsal shield 316–318 long, 160–162 wide, covering most of dorsal idiosoma, reticulated over whole surface, narrowing from level of setae
<emphasis id="B968EA81DE0B1E305AEE7538FB25805F" box="[1245,1271,333,359]" italics="true" pageId="9" pageNumber="160">r3</emphasis>
, progressively tapering until rounded posteriorly. Shield with 39 pairs of long setae, 22 pairs of podonotal setae on shield (
<emphasis id="B968EA81DE0B1E305B507504FA4680B3" box="[1379,1428,369,395]" italics="true" pageId="9" pageNumber="160">j1–6</emphasis>
;
<emphasis id="B968EA81DE0B1E305EA475E0FF1E8097" box="[151,204,405,431]" italics="true" pageId="9" pageNumber="160">z1–6</emphasis>
;
<emphasis id="B968EA81DE0B1E305EE975E0FEDD8097" box="[218,271,405,431]" italics="true" pageId="9" pageNumber="160">s1–6</emphasis>
;
<emphasis id="B968EA81DE0B1E305F2F75E0FE838097" box="[284,337,405,431]" italics="true" pageId="9" pageNumber="160">r2–5</emphasis>
), and 17 pairs of opisthonotal setae on shield (
<emphasis id="B968EA81DE0B1E305D6775E0FC588097" box="[852,906,405,431]" italics="true" pageId="9" pageNumber="160">J1–5</emphasis>
;
<emphasis id="B968EA81DE0B1E305DAB75E0FC008097" box="[920,978,405,431]" italics="true" pageId="9" pageNumber="160">Z1–5</emphasis>
;
<emphasis id="B968EA81DE0B1E305DEC75E0FBC58097" box="[991,1047,405,431]" italics="true" pageId="9" pageNumber="160">S1–5</emphasis>
; and two pairs of
<emphasis id="B968EA81DE0B1E305AEE75E3FB2B8097" box="[1245,1273,406,431]" italics="true" pageId="9" pageNumber="160">Zx</emphasis>
), supernumerary setae
<emphasis id="B968EA81DE0B1E305F3075CFFECE80EB" box="[259,284,442,467]" italics="true" pageId="9" pageNumber="160">Jx</emphasis>
between
<emphasis id="B968EA81DE0B1E305FBB75CFFE4680EB" box="[392,404,442,467]" italics="true" pageId="9" pageNumber="160">J</emphasis>
-series absent, most setae long enough to reach well past base of next posterior seta (28–38 long), all setae smooth and uniform in thickness and abruptly tapered apically (
<figureCitation id="13272A16DE0B1E305A3B75A8FB8980CF" box="[1032,1115,477,503]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">Fig. 32</figureCitation>
), except
<emphasis id="B968EA81DE0B1E305A8D75A8FB0680CF" box="[1214,1236,477,503]" italics="true" pageId="9" pageNumber="160">j1</emphasis>
,
<emphasis id="B968EA81DE0B1E305AD275A8FB2880CF" box="[1249,1274,477,503]" italics="true" pageId="9" pageNumber="160">z1</emphasis>
,
<emphasis id="B968EA81DE0B1E305B3B75A8FAF780CF" box="[1288,1317,477,503]" italics="true" pageId="9" pageNumber="160">Z5</emphasis>
very short (7–10) and
<emphasis id="B968EA81DE0B1E305F297674FEE48323" box="[282,310,513,539]" italics="true" pageId="9" pageNumber="160">S5</emphasis>
longest (44–46). Soft cuticle surrounding dorsal shield with six pairs setae (
<emphasis id="B968EA81DE0B1E305AA47674FB628323" box="[1175,1200,513,539]" italics="true" pageId="9" pageNumber="160">r6</emphasis>
,
<emphasis id="B968EA81DE0B1E305A8C7674FADE8323" box="[1215,1292,513,539]" italics="true" pageId="9" pageNumber="160">R2–R4</emphasis>
and
<emphasis id="B968EA81DE0B1E305B777674FA418323" box="[1348,1427,513,539]" italics="true" pageId="9" pageNumber="160">UR5–6</emphasis>
) setae. Shield with about 14 pairs of discernible pore-like structures, including ten poroids (
<emphasis id="B968EA81DE0B1E305AA67650FB6B8307" box="[1173,1209,549,575]" italics="true" pageId="9" pageNumber="160">id1</emphasis>
,
<emphasis id="B968EA81DE0B1E305AF47650FB388307" box="[1223,1258,549,575]" italics="true" pageId="9" pageNumber="160">id2</emphasis>
,
<emphasis id="B968EA81DE0B1E305ACA7650FACF8307" box="[1273,1309,549,575]" italics="true" pageId="9" pageNumber="160">id4</emphasis>
,
<emphasis id="B968EA81DE0B1E305B187650FA9D8307" box="[1323,1359,549,575]" italics="true" pageId="9" pageNumber="160">id5</emphasis>
,
<emphasis id="B968EA81DE0B1E305B6E7650FA478307" box="[1373,1429,549,575]" italics="true" pageId="9" pageNumber="160">idm1</emphasis>
,
<emphasis id="B968EA81DE0B1E305EA4763CFF1D835B" box="[151,207,585,611]" italics="true" pageId="9" pageNumber="160">idm2</emphasis>
,
<emphasis id="B968EA81DE0B1E305EEE763CFEC4835B" box="[221,278,585,611]" italics="true" pageId="9" pageNumber="160">idm3</emphasis>
,
<emphasis id="B968EA81DE0B1E305F10763CFE89835B" box="[291,347,585,611]" italics="true" pageId="9" pageNumber="160">idm5</emphasis>
,
<emphasis id="B968EA81DE0B1E305F5A763CFE73835B" box="[361,417,585,611]" italics="true" pageId="9" pageNumber="160">idm6</emphasis>
,
<emphasis id="B968EA81DE0B1E305F9C763CFE09835B" box="[431,475,585,611]" italics="true" pageId="9" pageNumber="160">idl1</emphasis>
) and four gland openings (
<emphasis id="B968EA81DE0B1E305D38763CFCE7835B" box="[779,821,585,611]" italics="true" pageId="9" pageNumber="160">gd1</emphasis>
,
<emphasis id="B968EA81DE0B1E305D70763CFCBF835B" box="[835,877,585,611]" italics="true" pageId="9" pageNumber="160">gd2</emphasis>
,
<emphasis id="B968EA81DE0B1E305D48763CFC77835B" box="[891,933,585,611]" italics="true" pageId="9" pageNumber="160">gd6</emphasis>
,
<emphasis id="B968EA81DE0B1E305D80763CFC0E835B" box="[947,988,585,611]" italics="true" pageId="9" pageNumber="160">gd9</emphasis>
), others indistinct (
<figureCitation id="13272A16DE0B1E305A85763CFAFD835B" box="[1206,1327,584,611]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figures 25</figureCitation>
and
<figureCitation id="13272A16DE0B1E305B55763CFA54835B" box="[1382,1414,585,611]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">30</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE0B1E305EF47619FDCD840C" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF47619FE5883BF" bold="true" box="[199,394,620,647]" italics="true" pageId="9" pageNumber="160">Ventral idiosoma</emphasis>
(
<figureCitation id="13272A16DE0B1E305FAB7618FE3D83BF" box="[408,495,621,647]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figs 26</figureCitation>
,
<figureCitation id="13272A16DE0B1E305FC97618FD5183BF" box="[506,643,621,647]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">31, 33 &amp; 34</figureCitation>
). Tritosternum with paired pilose laciniae (58–60), fused basally (2–4), columnar base 15–17 × 10–12 wide; presternal area poorly sclerotised, granulate-lineate (
<figureCitation id="13272A16DE0B1E305AAA76E4FB228393" box="[1177,1264,657,683]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figs 26</figureCitation>
&amp;
<figureCitation id="13272A16DE0B1E305B2B76E4FAE58393" box="[1304,1335,657,683]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">31</figureCitation>
). Sternal shield (length 99–102) narrowest between coxae II (73–75), widest (104–106), anterior margin completely indistinct and posterior margin almost straight; bears three pairs of smooth pointed setae (
<emphasis id="B968EA81DE0B1E305A6176ACFBA083CB" box="[1106,1138,729,755]" italics="true" pageId="9" pageNumber="160">st1</emphasis>
22–24,
<emphasis id="B968EA81DE0B1E305AFD76ACFB3C83CB" box="[1230,1262,729,755]" italics="true" pageId="9" pageNumber="160">st2</emphasis>
21–23,
<emphasis id="B968EA81DE0B1E305B7976ACFAB983CB" box="[1354,1387,729,755]" italics="true" pageId="9" pageNumber="160">st3</emphasis>
21– 23), not reaching base of next setae, one pair of lyrifissures adjacent to setae
<emphasis id="B968EA81DE0B1E305DCB7688FBCB822F" box="[1016,1049,765,791]" italics="true" pageId="9" pageNumber="160">st1</emphasis>
, and a pair of poroids between
<emphasis id="B968EA81DE0B1E305B4F7688FA4E822F" box="[1404,1436,765,791]" italics="true" pageId="9" pageNumber="160">st2</emphasis>
and
<emphasis id="B968EA81DE0B1E305EFB7754FF3B8203" box="[200,233,801,827]" italics="true" pageId="9" pageNumber="160">st3</emphasis>
; sternal shield smooth almost throughout (
<figureCitation id="13272A16DE0B1E305CFC7754FCF18203" box="[719,803,801,827]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">Fig. 33</figureCitation>
). Metasternal setae
<emphasis id="B968EA81DE0B1E305A367754FBF48203" box="[1029,1062,801,827]" italics="true" pageId="9" pageNumber="160">st4</emphasis>
(22–24) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates II/III completely fused to sternal shield, endopodal plates III/IV elongate, narrow and curved. Genital shield tongue-shaped, sides almost parallel, elongated, length 134–136, maximum width 40–42, surface without any distinct reticulation, almost completely smooth (
<figureCitation id="13272A16DE0B1E305B5C77F8FF6482F3" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">Fig. 34</figureCitation>
), posterior margin of genital shield almost reaches anterior margin of anal shield (
<figureCitation id="13272A16DE0B1E305A7E77C4FB7082F3" box="[1101,1186,945,971]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figs 26</figureCitation>
&amp;
<figureCitation id="13272A16DE0B1E305AF577C4FB3782F3" box="[1222,1253,945,971]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">31</figureCitation>
). Shield flanked by pair of minute, slightly elongate paragenital plaetelets; paragenital poroids
<emphasis id="B968EA81DE0B1E305A3F77A0FBFC82D7" box="[1036,1070,981,1007]" italics="true" pageId="9" pageNumber="160">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="B968EA81DE0B1E305F7E778CFEBC852B" box="[333,366,1017,1043]" italics="true" pageId="9" pageNumber="160">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 52–54, width 36–38, anterior half faintly lineate-reticulate, post-anal seta (20–22) twice as long as para-anal setae (10–12), cribrum small, with 3–4 irregular rows of spicules, limited to region posterior to post-anal seta; anal poroids
<emphasis id="B968EA81DE0B1E305A3C7034FBE58563" box="[1039,1079,1089,1115]" italics="true" pageId="9" pageNumber="160">gv3</emphasis>
on anterolateral margin of anal shield. Soft opisthogastric cuticle with pair of oval and slightly elongate metapodal plates (10–12 long × 2–4 wide), and 12 pairs of smooth setae (
<emphasis id="B968EA81DE0B1E305FD870FCFD95859B" box="[491,583,1161,1187]" italics="true" pageId="9" pageNumber="160">Jv1–Jv5</emphasis>
,
<emphasis id="B968EA81DE0B1E305C6570FCFD78859B" box="[598,682,1161,1187]" italics="true" pageId="9" pageNumber="160">Zv1–Zv</emphasis>
3) and four pairs of poroids including
<emphasis id="B968EA81DE0B1E305A6970FCFBAE859B" box="[1114,1148,1161,1187]" italics="true" pageId="9" pageNumber="160">iv5</emphasis>
. Exopodal and parapodal platelets not fused, strip-like extending narrowly behind coxae IV. Peritreme extending anteriorly to posterior level of coxa I, peritrematal shield narrow, expanded anteriorly, free from exopodal shields, each shield bearing five pore-like structures, a lyrifissure
<emphasis id="B968EA81DE0B1E305C387083FDF38437" box="[523,545,1270,1295]" italics="true" pageId="9" pageNumber="160">ip</emphasis>
and a gland pore
<emphasis id="B968EA81DE0B1E305CD97083FCD48437" box="[746,774,1270,1295]" italics="true" pageId="9" pageNumber="160">gp</emphasis>
at level of coxa III, two lyrifissures
<emphasis id="B968EA81DE0B1E305A927083FB658437" box="[1185,1207,1270,1295]" italics="true" pageId="9" pageNumber="160">ip</emphasis>
and a gland pore
<emphasis id="B968EA81DE0B1E305BB37083FA4E8437" box="[1408,1436,1270,1295]" italics="true" pageId="9" pageNumber="160">gp</emphasis>
on post-stigmatic section (
<figureCitation id="13272A16DE0B1E305F8D716CFDC1840C" box="[446,531,1305,1332]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE0B1E305EF47149FCB8874F" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="160">
<taxonomicName id="4C1C4D10DE0B1E305EF47149FE89846F" authorityName="Krefft" authorityYear="1871" box="[199,347,1340,1367]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="160" phylum="Nematoda" rank="genus">
<emphasis id="B968EA81DE0B1E305EF47149FE89846F" bold="true" box="[199,347,1340,1367]" italics="true" pageId="9" pageNumber="160">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13272A16DE0B1E305F5F7148FE3D846F" box="[364,495,1341,1367]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Figs 27–29</figureCitation>
,
<figureCitation id="13272A16DE0B1E305FCF7149FD95846F" box="[508,583,1340,1367]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">35–36</figureCitation>
). Epistome subtriangular and irregularly denticulate (
<figureCitation id="13272A16DE0B1E305A9B7148FB2C846F" box="[1192,1278,1341,1367]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Fig. 28</figureCitation>
). Hypostomal groove with seven transverse rows of denticles, each row with 8–15 small denticles, with smooth anterior transverse lines. Hypostome with four pairs of setae, internal posterior hypostomal setae
<emphasis id="B968EA81DE0B1E305A3671F0FBF384A7" box="[1029,1057,1413,1439]" italics="true" pageId="9" pageNumber="160">h3</emphasis>
(30–32),
<emphasis id="B968EA81DE0B1E305ABC71F0FB7984A7" box="[1167,1195,1413,1439]" italics="true" pageId="9" pageNumber="160">h1</emphasis>
(24–26),
<emphasis id="B968EA81DE0B1E305B2A71F0FAE484A7" box="[1305,1334,1413,1439]" italics="true" pageId="9" pageNumber="160">h2</emphasis>
(13–15), palpcoxal
<emphasis id="B968EA81DE0B1E305F3F71DFFEF484FB" box="[268,294,1450,1475]" italics="true" pageId="9" pageNumber="160">pc</emphasis>
(22–24) (
<figureCitation id="13272A16DE0B1E305FAB71DCFE3F84FB" box="[408,493,1449,1475]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Fig. 27</figureCitation>
). Corniculi robust and horn-like, extending slightly beyond palptrochanter. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except except
<emphasis id="B968EA81DE0B1E305B6471B8FAA984DF" box="[1367,1403,1485,1511]" italics="true" pageId="9" pageNumber="160">al1</emphasis>
on palp femur and
<emphasis id="B968EA81DE0B1E305F747184FEBE8733" box="[327,364,1521,1547]" italics="true" pageId="9" pageNumber="160">al1</emphasis>
,
<emphasis id="B968EA81DE0B1E305F4A7184FE448733" box="[377,406,1521,1547]" italics="true" pageId="9" pageNumber="160">d1</emphasis>
on palp genu thickened and apically spatulate, palp tarsal apotele two-tined (
<figureCitation id="13272A16DE0B1E305AC67184FA9A8733" box="[1269,1352,1520,1547]" captionStart="FIGURES 30–36" captionStartId="11.[151,264,1946,1971]" captionTargetBox="[182,1407,184,1918]" captionTargetId="figure@11.[180,1407,181,1920]" captionTargetPageId="11" captionText="FIGURES 30–36. DIC micrographs of Cosmolaelaps trichiurus sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="9" pageNumber="160">Fig. 35</figureCitation>
). Fixed digit of chelicera with an offset and most distal large tooth (gabelzhan), followed five variously sized teeth in two rows, a setaceous pilus dentilis, dorsal cheliceral setae thickened prostrate, arthrodial membrane reduced to a lobe and without filaments; cheliceral lyrifissures indistinct (
<figureCitation id="13272A16DE0B1E305D3A7228FC8F874F" box="[777,861,1629,1655]" captionStart="FIGURES 25–29" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[190,1395,194,1908]" captionTargetId="figure@10.[178,1410,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–29. Cosmolaelaps trichiurus sp. nov., female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="9" pageNumber="160">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE0B1E305EF472F4FDE78683" blockId="9.[151,1437,153,2015]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF472F4FF2F87A3" bold="true" box="[199,253,1665,1691]" italics="true" pageId="9" pageNumber="160">Legs</emphasis>
(
<figureCitation id="13272A16DE0B1E305F3E72F4FE4287A3" box="[269,400,1665,1691]" captionStart="FIGURE 37–40" captionStartId="12.[151,250,1911,1936]" captionTargetBox="[166,1432,190,1874]" captionTargetId="figure@12.[151,1436,181,1885]" captionTargetPageId="12" captionText="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687720" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="9" pageNumber="160">Figs 37–40</figureCitation>
). Legs II (232–237) and III, (206–195) short, I (306–309) and IV (278–286) longer. Chaetotaxy normal for free-living
<taxonomicName id="4C1C4D10DE0B1E305FF672D0FD968787" box="[453,580,1701,1727]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="13272A16DE0B1E305CA472D0FD388787" box="[663,746,1701,1727]" captionStart="FIGURE 37–40" captionStartId="12.[151,250,1911,1936]" captionTargetBox="[166,1432,190,1874]" captionTargetId="figure@12.[151,1436,181,1885]" captionTargetPageId="12" captionText="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687720" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="9" pageNumber="160">Fig. 37</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 1/2-1, femur 2-2/1, 3/2-2 (
<emphasis id="B968EA81DE0B1E305E9272BCFF1B87DB" box="[161,201,1737,1763]" italics="true" pageId="9" pageNumber="160">pv3</emphasis>
absent), genu 2-3/2, 3/1-2 (
<emphasis id="B968EA81DE0B1E305C3472BFFDF387DB" box="[519,545,1738,1763]" italics="true" pageId="9" pageNumber="160">pv</emphasis>
slightly thickened), tibia 2-3/2, 3/1-2 (ventral setae slightly thickened). Leg II (
<figureCitation id="13272A16DE0B1E305E937298FF21863F" box="[160,243,1773,1799]" captionStart="FIGURE 37–40" captionStartId="12.[151,250,1911,1936]" captionTargetBox="[166,1432,190,1874]" captionTargetId="figure@12.[151,1436,181,1885]" captionTargetPageId="12" captionText="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687720" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="9" pageNumber="160">Fig. 38</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1, femur 2-3/1, 2/2-1, genu 2-3/1, 2/1-2, tibia 2-2/1, 2/1-2 (ventral setae slightly thickened). Leg III (
<figureCitation id="13272A16DE0B1E305C297364FDA28613" box="[538,624,1809,1835]" captionStart="FIGURE 37–40" captionStartId="12.[151,250,1911,1936]" captionTargetBox="[166,1432,190,1874]" captionTargetId="figure@12.[151,1436,181,1885]" captionTargetPageId="12" captionText="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687720" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="9" pageNumber="160">Fig. 39</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 2-1/1, 0/1-0, femur 1-2/1, 1/0-1, genu 2- 2/1, 2/0-1 (
<emphasis id="B968EA81DE0B1E305F247343FEE38677" box="[279,305,1846,1871]" italics="true" pageId="9" pageNumber="160">pv</emphasis>
absent), tibia: 2-1/1, 2/1-1 (ventral setae thickened). Leg IV (
<figureCitation id="13272A16DE0B1E305DDD7340FB978677" box="[1006,1093,1845,1871]" captionStart="FIGURE 37–40" captionStartId="12.[151,250,1911,1936]" captionTargetBox="[166,1432,190,1874]" captionTargetId="figure@12.[151,1436,181,1885]" captionTargetPageId="12" captionText="FIGURE 37–40. Cosmolaelaps trichiurus sp. nov., female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687720" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="9" pageNumber="160">Fig. 40</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 2-1/1, 0/1-1, femur 1-2/1, 1/0-1, genu 2-2/1, 3/0-1, tibia 2-1/1, 3/1-2. Tarsi II-IV with 18 setae (3- 3/2, 3/2-3 +
<emphasis id="B968EA81DE0B1E305B46732FFA44864B" box="[1397,1430,1882,1907]" italics="true" pageId="9" pageNumber="160">mv</emphasis>
,
<emphasis id="B968EA81DE0B1E305EA47308FF6B86AF" box="[151,185,1917,1943]" italics="true" pageId="9" pageNumber="160">md</emphasis>
); some ventral and lateral setae on tarsi II-III thickened. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="8BA33693DE0B1E305EF473B0FCA786E7" blockId="9.[151,1437,153,2015]" box="[199,885,1989,2015]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF473B0FE3686E7" bold="true" box="[199,484,1989,2015]" pageId="9" pageNumber="160">
<emphasis id="B968EA81DE0B1E305EF473B0FE0986E7" bold="true" box="[199,475,1989,2015]" italics="true" pageId="9" pageNumber="160">Insemination structures</emphasis>
:
</emphasis>
Not seen, apparently unsclerotised.
</paragraph>
<caption id="DF63661BDE081E335EA473ECFE5B86EE" ID-DOI="http://doi.org/10.5281/zenodo.3687716" ID-Zenodo-Dep="3687716" httpUri="https://zenodo.org/record/3687716/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,264,1945,1970]" targetBox="[190,1395,194,1908]" targetPageId="10">
<paragraph id="8BA33693DE081E335EA473ECFE5B86EE" blockId="10.[151,1436,1945,2006]" pageId="10" pageNumber="161">
<emphasis id="B968EA81DE081E335EA473ECFE87868A" bold="true" box="[151,341,1945,1970]" pageId="10" pageNumber="161">FIGURES 25–29.</emphasis>
<taxonomicName id="4C1C4D10DE081E335F6873EFFD87868A" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[347,597,1946,1970]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="trichiurus" status="sp. nov.">
<emphasis id="B968EA81DE081E335F6873EFFD87868A" box="[347,597,1946,1970]" italics="true" pageId="10" pageNumber="161">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE081E335C6F73EFFD7F868A" bold="true" box="[604,685,1946,1970]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="A25B57FADE081E335C6F73EFFD7F868A" box="[604,685,1946,1970]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 25, dorsal idiosoma; 26, ventral idiosoma; 27, subcapitulum; 28, epistome; 29, chelicera.
</paragraph>
</caption>
<caption id="DF63661BDE091E325EA473EFFAE486EF" ID-DOI="http://doi.org/10.5281/zenodo.3687718" ID-Zenodo-Dep="3687718" httpUri="https://zenodo.org/record/3687718/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,264,1946,1971]" targetBox="[182,1407,184,1918]" targetPageId="11">
<paragraph id="8BA33693DE091E325EA473EFFAE486EF" blockId="11.[151,1437,1946,2007]" pageId="11" pageNumber="162">
<emphasis id="B968EA81DE091E325EA473EFFE86868B" bold="true" box="[151,340,1946,1971]" pageId="11" pageNumber="162">FIGURES 30–36.</emphasis>
DIC micrographs of
<taxonomicName id="4C1C4D10DE091E325C1F73EFFCF7868A" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[556,805,1946,1970]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="trichiurus" status="sp. nov.">
<emphasis id="B968EA81DE091E325C1F73EFFCF7868A" box="[556,805,1946,1970]" italics="true" pageId="11" pageNumber="162">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE091E325D1F73EFFCAE868A" bold="true" box="[812,892,1946,1970]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="A25B57FADE091E325D1F73EFFCAE868A" box="[812,892,1946,1970]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some dorsal setae; 33, sternal shield; 34, genital shield; 35, palp; 36, subcapitulum.
</paragraph>
</caption>
<caption id="DF63661BDE0E1E355EA47302FA8786B7" ID-DOI="http://doi.org/10.5281/zenodo.3687720" ID-Zenodo-Dep="3687720" box="[151,1365,1911,1936]" httpUri="https://zenodo.org/record/3687720/files/figure.png" pageId="12" pageNumber="163" startId="12.[151,250,1911,1936]" targetBox="[166,1432,190,1874]" targetPageId="12">
<paragraph id="8BA33693DE0E1E355EA47302FA8786B7" blockId="12.[151,1365,1911,1936]" box="[151,1365,1911,1936]" pageId="12" pageNumber="163">
<emphasis id="B968EA81DE0E1E355EA47302FE9486A8" bold="true" box="[151,326,1911,1936]" pageId="12" pageNumber="163">FIGURE 37–40.</emphasis>
<taxonomicName id="4C1C4D10DE0E1E355F7F7302FD9486B7" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[332,582,1911,1935]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="trichiurus" status="sp. nov.">
<emphasis id="B968EA81DE0E1E355F7F7302FD9486B7" box="[332,582,1911,1935]" italics="true" pageId="12" pageNumber="163">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE0E1E355C7F730DFD4F86A8" bold="true" box="[588,669,1912,1936]" pageId="12" pageNumber="163">
<taxonomicNameLabel id="A25B57FADE0E1E355C7F730DFD4F86A8" box="[588,669,1912,1936]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 31, leg I (trochanter-tibia); 32, leg II; 33, leg III; 34, leg IV.
</paragraph>
</caption>
<paragraph id="8BA33693DE0F1E345EF474ECFD96818A" blockId="13.[151,1437,152,971]" box="[199,580,152,179]" pageId="13" pageNumber="164">
<emphasis id="B968EA81DE0F1E345EF474ECFE15818B" bold="true" box="[199,455,152,179]" pageId="13" pageNumber="164">Male and immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0F1E345EF474C8FF2B81C3" pageId="13" pageNumber="164" type="etymology">
<paragraph id="8BA33693DE0F1E345EF474C8FF2B81C3" blockId="13.[151,1437,152,971]" pageId="13" pageNumber="164">
<emphasis id="B968EA81DE0F1E345EF474C8FE9C81EF" bold="true" box="[199,334,189,215]" pageId="13" pageNumber="164">Etymology.</emphasis>
The species name
<taxonomicName id="4C1C4D10DE0F1E345C1174C9FD4281EE" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[546,656,188,214]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="trichiurus">
<emphasis id="B968EA81DE0F1E345C1174C9FD4281EE" box="[546,656,188,214]" italics="true" pageId="13" pageNumber="164">trichiurus</emphasis>
</taxonomicName>
(Greek
<emphasis id="B968EA81DE0F1E345CD874C9FCF281EE" box="[747,800,188,214]" italics="true" pageId="13" pageNumber="164">thrix</emphasis>
, hair +
<emphasis id="B968EA81DE0F1E345D4474C8FC7E81EE" box="[887,940,189,214]" italics="true" pageId="13" pageNumber="164">oura</emphasis>
, tail) refers to the tail-like appearance of the setae
<emphasis id="B968EA81DE0F1E345EE57494FF2081C3" box="[214,242,225,251]" italics="true" pageId="13" pageNumber="164">S5</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0F1E345EF47571FC0D82F2" pageId="13" pageNumber="164" type="diagnosis">
<paragraph id="8BA33693DE0F1E345EF47571FC0D82F2" blockId="13.[151,1437,152,971]" pageId="13" pageNumber="164">
<emphasis id="B968EA81DE0F1E345EF47571FE1D8027" bold="true" box="[199,463,260,287]" pageId="13" pageNumber="164">Differential diagnosis.</emphasis>
<taxonomicName id="4C1C4D10DE0F1E345FEB7571FD3F8026" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[472,749,260,286]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="trichiurus">
<emphasis id="B968EA81DE0F1E345FEB7571FD3F8026" box="[472,749,260,286]" italics="true" pageId="13" pageNumber="164">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
is similar in general appearance and the shape of the dorsal setae to
<taxonomicName id="4C1C4D10DE0F1E345EC7755CFE3C807B" authority="(Karg, 1993)" baseAuthorityName="Karg" baseAuthorityYear="1993" box="[244,494,296,323]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="gladii">
<emphasis id="B968EA81DE0F1E345EC7755CFED5807B" box="[244,263,297,323]" italics="true" pageId="13" pageNumber="164">C</emphasis>
.
<emphasis id="B968EA81DE0F1E345F26755DFE84807A" box="[277,342,296,322]" italics="true" pageId="13" pageNumber="164">gladii</emphasis>
(
<bibRefCitation id="EF8D4B62DE0F1E345F56755CFE34807B" author="Karg, W." box="[357,486,297,323]" pageId="13" pageNumber="164" pagination="1 - 523" refId="ref8189" refString="Karg, W. (1993) Acari (Acarina), Milben Parasitiformes (Anactinochaeta) Cohors Gamasina Leach Raubmilben. 2. Uberarbeitete Auflage. Die Tierwelt Deutschlands, 59, 1 - 523." type="journal article" year="1993">Karg, 1993</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1C4D10DE0F1E345C16755DFB68807B" authority="Joharchi, Fan &amp; Arjomandi, 2017" authorityName="Joharchi, Fan &amp; Arjomandi" authorityYear="2017" box="[549,1210,296,323]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="dioscorea">
<emphasis id="B968EA81DE0F1E345C16755DFCEA807A" box="[549,824,296,322]" italics="true" pageId="13" pageNumber="164">Cosmolaelaps dioscorea</emphasis>
<bibRefCitation id="EF8D4B62DE0F1E345D73755DFB68807B" author="Joharchi, O. &amp; Fan, Q. - H. &amp; Arjomandi, E." box="[832,1210,296,323]" pageId="13" pageNumber="164" pagination="789 - 796" refId="ref7701" refString="Joharchi, O., Fan, Q. - H. &amp; Arjomandi, E. (2017) A new species of Cosmolaelaps Berlese (Acari: Mesostigmata: Laelapidae) from the Pacific Islands. Systematic &amp; Applied Acarology, 22 (6), 789 - 796. https: // doi. org / 10.11158 / saa. 22.6.5" type="journal article" year="2017">Joharchi, Fan &amp; Arjomandi, 2017</bibRefCitation>
</taxonomicName>
described from Galápagos Islands and
<collectingCountry id="F30B7603DE0F1E345F4F7539FE74805E" box="[380,422,332,358]" name="Fiji" pageId="13" pageNumber="164">Fiji</collectingCountry>
, respectively.
<taxonomicName id="4C1C4D10DE0F1E345C757539FC88805E" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[582,858,332,358]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="trichiurus">
<emphasis id="B968EA81DE0F1E345C757539FC88805E" box="[582,858,332,358]" italics="true" pageId="13" pageNumber="164">Cosmolaelaps trichiurus</emphasis>
</taxonomicName>
can be easily distinguished from these two species by
<emphasis id="B968EA81DE0F1E345E887504FF0580B3" box="[187,215,369,395]" italics="true" pageId="13" pageNumber="164">S5</emphasis>
much longer than other dorsal setae (vs.
<emphasis id="B968EA81DE0F1E345C997504FD1480B3" box="[682,710,369,395]" italics="true" pageId="13" pageNumber="164">
S
<quantity id="4CE49B76DE0F1E345C8B7504FD1480B3" box="[696,710,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="164" unit="in" value="5.0">5</quantity>
</emphasis>
in the same size with other dorsal setae in both species), genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield (genital shield not elongated in both species), length of post-anal seta almost twice that of para-anal setae (vs. length of post-anal seta shorter or at most subequal of para-anal setae but never longer in both species), fixed digit of chelicera with six teeth, dorsal cheliceral setae thickened, arthrodial membrane reduced to a lobe and without filaments (fixed digit of chelicera with 11 and eight teeth in
<taxonomicName id="4C1C4D10DE0F1E345CB67650FD358306" baseAuthorityName="Karg" baseAuthorityYear="1993" box="[645,743,548,575]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="gladii">
<emphasis id="B968EA81DE0F1E345CB67650FD4A8307" box="[645,664,549,575]" italics="true" pageId="13" pageNumber="164">C</emphasis>
.
<emphasis id="B968EA81DE0F1E345C957651FD358306" box="[678,743,548,574]" italics="true" pageId="13" pageNumber="164">gladii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D10DE0F1E345D2C7650FC7F8306" authorityName="Joharchi, Fan &amp; Arjomandi" authorityYear="2017" box="[799,941,548,575]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="dioscorea">
<emphasis id="B968EA81DE0F1E345D2C7650FCE08307" box="[799,818,549,575]" italics="true" pageId="13" pageNumber="164">C</emphasis>
.
<emphasis id="B968EA81DE0F1E345D737651FC7F8306" box="[832,941,548,574]" italics="true" pageId="13" pageNumber="164">dioscorea</emphasis>
</taxonomicName>
, respectively, cheliceral setae not thickened and arthrodial membrane with a rounded flap and normal filaments in both species). According to shape of dorsal setae,
<taxonomicName id="4C1C4D10DE0F1E345EEE7618FEB983BE" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[221,363,620,647]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="trichiurus">
<emphasis id="B968EA81DE0F1E345EEE7618FF2283BF" box="[221,240,621,647]" italics="true" pageId="13" pageNumber="164">C</emphasis>
.
<emphasis id="B968EA81DE0F1E345ECE7619FEB983BE" box="[253,363,620,646]" italics="true" pageId="13" pageNumber="164">trichiurus</emphasis>
</taxonomicName>
in the classification of
<bibRefCitation id="EF8D4B62DE0F1E345C437618FD2E83BF" author="Karg, W." box="[624,764,621,647]" pageId="13" pageNumber="164" pagination="213 - 232" refId="ref8030" refString="Karg, W. (1981) Die Untergattung Cosmolaelaps Berlese, 1903 der Raubmilbengattung Hypoaspis Canestrini, 1884. Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 213 - 232." type="journal article" year="1981">Karg (1981)</bibRefCitation>
, falls into the
<emphasis id="B968EA81DE0F1E345DAA7618FC2983BE" box="[921,1019,621,646]" italics="true" pageId="13" pageNumber="164">nasoseta</emphasis>
species group of
<taxonomicName id="4C1C4D10DE0F1E345A8C7618FF218393" authority="" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="subGenus" subGenus="Cosmolaelaps">
<emphasis id="B968EA81DE0F1E345A8C7618FAE783BE" box="[1215,1333,621,646]" italics="true" pageId="13" pageNumber="164">Hypoaspis</emphasis>
(
<emphasis id="B968EA81DE0F1E345B767618FF3A8392" italics="true" pageId="13" pageNumber="164">Cosmolaelaps</emphasis>
)
</taxonomicName>
, but
<taxonomicName id="4C1C4D10DE0F1E345F1976E4FE6B8392" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[298,441,656,683]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="trichiurus">
<emphasis id="B968EA81DE0F1E345F1976E4FEEF8393" box="[298,317,657,683]" italics="true" pageId="13" pageNumber="164">C</emphasis>
.
<emphasis id="B968EA81DE0F1E345F7876E5FE6B8392" box="[331,441,656,682]" italics="true" pageId="13" pageNumber="164">trichiurus</emphasis>
</taxonomicName>
is distinguished easily from all species in
<emphasis id="B968EA81DE0F1E345DA076E4FC278392" box="[915,1013,657,682]" italics="true" pageId="13" pageNumber="164">nasoseta</emphasis>
species group by following combination of unique characters: (1) dorsal shield narrowing from level of setae
<emphasis id="B968EA81DE0F1E345DE976C0FC2183F7" box="[986,1011,693,719]" italics="true" pageId="13" pageNumber="164">r3</emphasis>
, progressively tapering until rounded posteriorly and most setae long enough to reach well past base of next posterior seta, all setae abruptly taper apically (except
<emphasis id="B968EA81DE0F1E345EC07688FEDA822F" box="[243,264,765,791]" italics="true" pageId="13" pageNumber="164">j1</emphasis>
,
<emphasis id="B968EA81DE0F1E345F2B7688FEE3822F" box="[280,305,765,791]" italics="true" pageId="13" pageNumber="164">z1</emphasis>
,
<emphasis id="B968EA81DE0F1E345F727688FE8D822F" box="[321,351,765,791]" italics="true" pageId="13" pageNumber="164">Z5</emphasis>
very short) and setae
<emphasis id="B968EA81DE0F1E345C6C7688FDA9822F" box="[607,635,765,791]" italics="true" pageId="13" pageNumber="164">S5</emphasis>
much longer than other dorsal setae; (2) sternal shield smooth almost throughout; anterior margin indistinct and posterior margin almost straight; (3) genital shield elongated, posterior margin of genital shield almost reaches anterior margin of anal shield and surface almost completely smooth; (4) length of post-anal seta almost twice of para-anal setae; (5) fixed digit of chelicera with six teeth, dorsal cheliceral setae thickened and arthrodial membrane reduced to a lobe (filaments absent); (6) Femur I (2-2/1, 3/2-2) and genu III (2-2/1, 2/0-1) with 12 (
<emphasis id="B968EA81DE0F1E345F8E77C4FE3482F3" box="[445,486,945,971]" italics="true" pageId="13" pageNumber="164">pv3</emphasis>
absent) and 8 (
<emphasis id="B968EA81DE0F1E345CA177C4FD7E82F2" box="[658,684,945,970]" italics="true" pageId="13" pageNumber="164">pv</emphasis>
absent) setae, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>
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