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<document id="5BEF424E9C10A4A0143B99B049A06B32" ID-DOI="10.11646/zootaxa.4743.2.1" ID-GBIF-Dataset="8eba416f-e2af-4cfe-96fb-fb6059a877ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3687704" ID-ZooBank="624E410F-A208-46D4-BF6B-5331B39582F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582709117521" checkinUser="plazi" docAuthor="Joharchi, Omid, Ermilov, Sergey G. &amp; Khaustov, Alexander A." docDate="2020" docId="03B58785DE031E315EA470B3FDAE8442" docLanguage="en" docName="zootaxa.4743.2.1.pdf" docOrigin="Zootaxa 4743 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps ceylonensis Joharchi &amp; Ermilov &amp; Khaustov 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="159" masterDocId="FF8CFFFDDE021E395E337475FFD28138" masterDocTitle="Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka" masterLastPageNumber="166" masterPageNumber="151" pageNumber="152" updateTime="1698721478006" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="12FE63677389716FEA15F8E39EFD9408">
<mods:title id="BC670AB6B955D0B41EF04D86B5B54E52">Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Sri Lanka</mods:title>
</mods:titleInfo>
<mods:name id="6F93FBB2076428B9CF9CE79883810C04" type="personal">
<mods:role id="18807AF91883A6FDB341AC0DEB3B6BEF">
<mods:roleTerm id="331FAC597798C91DC9F5FC90E9BE9C2E">Author</mods:roleTerm>
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<mods:namePart id="7A1EFE36EFBCAB2A1EF2CD8B4690E9A4">Joharchi, Omid</mods:namePart>
</mods:name>
<mods:name id="84BF86B7932E0DFF4AE35E7D1BADC9B2" type="personal">
<mods:role id="D9A4934F69FFEC0E70DEFD521B0EE9A3">
<mods:roleTerm id="0FBD5D923EC40F5A77C5D3DCF1C0EFCB">Author</mods:roleTerm>
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<mods:namePart id="D1C89271CCBCFF83F123DC8889A68235">Ermilov, Sergey G.</mods:namePart>
</mods:name>
<mods:name id="FBFD1B3BC829CB2575CDC1F63449D765" type="personal">
<mods:role id="96E2620DD6D7A3EE3FD09B1DFA468D9F">
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<mods:namePart id="97A78A23284200BEC04C915F13288558">Khaustov, Alexander A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3732D8E0D2B00A844EF70376681A3994">text</mods:typeOfResource>
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<mods:title id="F991FC6DD7A5F2BC3A4B56057261B540">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1588C5AD79C16E8DF46D4908F71C1F88">
<mods:date id="6B0F29FFBB2D4EE027AFE222D391027B">2020</mods:date>
<mods:detail id="56D0BEAB3ECBC16510AF97A061D233F4" type="pubDate">
<mods:number id="EC98C2344C5FDB76EE598905C20D7B4E">2020-02-25</mods:number>
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<mods:detail id="AC5CC5B11AF0773CC12F7D17481D6DF9" type="volume">
<mods:number id="84CF849BDC8D486E47E28913BA766CEF">4743</mods:number>
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<mods:detail id="30C6420F95B29F5CAEED0419963DB2EC" type="issue">
<mods:number id="B7B83C8B17C6B52B985608036DB8CB61">2</mods:number>
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<mods:classification id="1B7AF40ED3C5A196D911233AAC14182C">journal article</mods:classification>
<mods:identifier id="684F8289BDD511D579B34DBAEA0FF8AA" type="DOI">10.11646/zootaxa.4743.2.1</mods:identifier>
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<treatment id="03B58785DE031E315EA470B3FDAE8442" ID-DOI="http://doi.org/10.5281/zenodo.3688002" ID-GBIF-Taxon="162267490" ID-Zenodo-Dep="3688002" LSID="urn:lsid:plazi:treatment:03B58785DE031E315EA470B3FDAE8442" httpUri="http://treatment.plazi.org/id/03B58785DE031E315EA470B3FDAE8442" lastPageId="8" lastPageNumber="159" pageId="1" pageNumber="152">
<subSubSection id="C3066518DE031E385EA470B3FDE485D9" box="[151,566,1222,1249]" pageId="1" pageNumber="152" type="nomenclature">
<paragraph id="8BA33693DE031E385EA470B3FDE485D9" blockId="1.[151,566,1222,1284]" box="[151,566,1222,1249]" pageId="1" pageNumber="152">
<heading id="D0EB81FFDE031E385EA470B3FDE485D9" bold="true" box="[151,566,1222,1249]" fontSize="11" level="1" pageId="1" pageNumber="152" reason="1">
<emphasis id="B968EA81DE031E385EA470B3FDE485D9" bold="true" box="[151,566,1222,1249]" pageId="1" pageNumber="152">
<taxonomicName id="4C1C4D10DE031E385EA470B3FE0385D9" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[151,465,1222,1249]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE031E385EA470B3FE0385D9" bold="true" box="[151,465,1222,1249]" italics="true" pageId="1" pageNumber="152">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57FADE031E385FEB70B2FDE485D9" box="[472,566,1223,1249]" pageId="1" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE031E385EA4709FFEEE843C" box="[151,316,1258,1284]" pageId="1" pageNumber="152" type="description">
<paragraph id="8BA33693DE031E385EA4709FFEEE843C" blockId="1.[151,566,1222,1284]" box="[151,316,1258,1284]" pageId="1" pageNumber="152">
(
<figureCitation id="13272A16DE031E385EAC709FFEE6843C" box="[159,308,1258,1284]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 714" captionStart-2="FIGURE 1518" captionStart-3="FIGURES 1920" captionStart-4="FIGURES 2124" captionStartId-0="3.[151,264,1945,1970]" captionStartId-1="4.[151,264,1945,1970]" captionStartId-2="5.[151,250,1910,1935]" captionStartId-3="6.[151,264,1987,2012]" captionStartId-4="7.[151,264,1945,1970]" captionTargetBox-0="[182,1399,191,1905]" captionTargetBox-1="[180,1407,182,1918]" captionTargetBox-2="[162,1425,192,1867]" captionTargetBox-3="[164,1421,650,1948]" captionTargetBox-4="[175,1409,187,1920]" captionTargetId-0="figure@3.[173,1415,179,1917]" captionTargetId-1="figure@4.[179,1409,181,1920]" captionTargetId-2="figure@5.[151,1436,181,1885]" captionTargetId-3="figure@6.[151,1435,633,1962]" captionTargetId-4="figure@7.[175,1413,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." captionText-1="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." captionText-2="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." captionText-3="FIGURES 1920. Cosmolaelaps ceylonensis sp. nov., male. 19, ventral idiosoma; 20, chelicera." captionText-4="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi-0="http://doi.org/10.5281/zenodo.3687706" figureDoi-1="http://doi.org/10.5281/zenodo.3687708" figureDoi-2="http://doi.org/10.5281/zenodo.3687710" figureDoi-3="http://doi.org/10.5281/zenodo.3687712" figureDoi-4="http://doi.org/10.5281/zenodo.3687714" httpUri-0="https://zenodo.org/record/3687706/files/figure.png" httpUri-1="https://zenodo.org/record/3687708/files/figure.png" httpUri-2="https://zenodo.org/record/3687710/files/figure.png" httpUri-3="https://zenodo.org/record/3687712/files/figure.png" httpUri-4="https://zenodo.org/record/3687714/files/figure.png" pageId="1" pageNumber="152">Figures 124</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE031E385EA47147FCDB8480" pageId="1" pageNumber="152" type="materials_examined">
<paragraph id="8BA33693DE031E385EA47147FCDB8480" blockId="1.[151,1437,1329,2041]" pageId="1" pageNumber="152">
<emphasis id="B968EA81DE031E385EA47147FE9A8475" bold="true" box="[151,328,1329,1357]" pageId="1" pageNumber="152">Type material:</emphasis>
<materialsCitation id="3B743CCEDE031E385F627147FE1A84AC" ID-GBIF-Occurrence="2571176103" collectingDate="2019-01-24" collectionCode="TSUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="1" pageNumber="152" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<typeStatus id="54A78831DE031E385F627147FE6E8474" box="[337,444,1330,1356]" pageId="1" pageNumber="152" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="F30B7603DE031E385C177147FD448474" box="[548,662,1330,1356]" name="Sri Lanka" pageId="1" pageNumber="152">Sri Lanka</collectingCountry>
,
<collectingRegion id="49D8F871DE031E385C917147FC698474" box="[674,955,1330,1356]" country="Sri Lanka" name="Sabaragamuwa" pageId="1" pageNumber="152">Sabaragamuwa Province</collectingRegion>
,
<location id="8EC36048DE031E385DFB7147FB838474" LSID="urn:lsid:plazi:treatment:03B58785DE031E315EA470B3FDAE8442:8EC36048DE031E385DFB7147FB838474" box="[968,1105,1330,1356]" country="Sri Lanka" latitude="6.45" longLatPrecision="1304" longitude="80.2" name="Polgampola" pageId="1" pageNumber="152" stateProvince="Sabaragamuwa">Polgampola</location>
,
<geoCoordinate id="EE285054DE031E385A6E7147FB158475" box="[1117,1223,1330,1357]" degrees="06" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="152" precision="925" value="6.45">06°27 N</geoCoordinate>
,
<geoCoordinate id="EE285054DE031E385AE77147FA958475" box="[1236,1351,1330,1357]" degrees="080" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="152" precision="925" value="80.2">080°12 E</geoCoordinate>
, alt.
<quantity id="4CE49B76DE031E385BB37147FF7D8449" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="152" unit="m" value="42.0">
<elevation id="0031D1A0DE031E385BB37147FF7D8449" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="152" unit="m" value="42.0">42 m</elevation>
</quantity>
,
<date id="FFA21053DE031E385E887123FEAA8448" box="[187,376,1366,1393]" pageId="1" pageNumber="152" value="2019-01-24">
<collectingDate id="EFE6E9BBDE031E385E887123FEAA8448" box="[187,376,1366,1393]" pageId="1" pageNumber="152" value="2019-01-24">24 January 2019</collectingDate>
</date>
,
<collectorName id="26E95345DE031E385FB77123FDDB8448" box="[388,521,1366,1392]" pageId="1" pageNumber="152">O. Joharchi</collectorName>
,
<collectorName id="26E95345DE031E385C277123FD7B8448" box="[532,681,1366,1392]" pageId="1" pageNumber="152">S.G. Ermilov</collectorName>
&amp;
<collectorName id="26E95345DE031E385CFF7123FCA78448" box="[716,885,1366,1392]" pageId="1" pageNumber="152">A.A. Khaustov</collectorName>
coll., from forest soil-litter near the
<collectorName id="26E95345DE031E385B237123FF1684AC" pageId="1" pageNumber="152">Thambadola Ella</collectorName>
waterfall (in
<collectionCode id="ED0DAE56DE031E385F68710FFE6D84AC" box="[347,447,1402,1428]" pageId="1" pageNumber="152">TSUMZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B743CCEDE031E385FE1710EFCD78480" ID-GBIF-Occurrence="2571176105" collectingDate="2019-01-24" collectionCode="TSUMZ, ZISP" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="1" pageNumber="152" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="paratype">
<typeStatus id="54A78831DE031E385FE1710EFD9084AD" box="[466,578,1403,1429]" pageId="1" pageNumber="152" type="paratype">Paratypes</typeStatus>
, three females and seven males same data as holotype (two females and four males in
<collectionCode id="ED0DAE56DE031E385ECF71EBFE8C8480" box="[252,350,1438,1464]" pageId="1" pageNumber="152">TSUMZ</collectionCode>
, one female and three males in
<collectionCode id="ED0DAE56DE031E385C8C71EBFD2F8480" box="[703,765,1438,1464]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="152">ZISP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE031E385EF471B4FA9A8688" pageId="1" pageNumber="152" type="diagnosis">
<paragraph id="8BA33693DE031E385EF471B4FA9A8688" blockId="1.[151,1437,1329,2041]" pageId="1" pageNumber="152">
<emphasis id="B968EA81DE031E385EF471B4FEE884E4" bold="true" box="[199,314,1473,1500]" pageId="1" pageNumber="152">Diagnosis</emphasis>
(adult female and male). Dorsal shield with well reticulate ornamentation over whole surface, covering most of the idiosoma, narrowing from level of setae
<emphasis id="B968EA81DE031E385D377193FCCF8738" box="[772,797,1510,1536]" italics="true" pageId="1" pageNumber="152">r3</emphasis>
, progressively tapering until rounded posteriorly, bearing 39 pairs of subequal oblong-mucronate setae (22 pairs of podonotal setae and 17 pairs of opisthonotal setae, including two pairs of
<emphasis id="B968EA81DE031E385F78725AFEB58770" box="[331,359,1583,1608]" italics="true" pageId="1" pageNumber="152">Zx</emphasis>
setae) except
<emphasis id="B968EA81DE031E385C37725BFDC88770" box="[516,538,1582,1608]" italics="true" pageId="1" pageNumber="152">j1</emphasis>
,
<emphasis id="B968EA81DE031E385C14725BFD928770" box="[551,576,1582,1608]" italics="true" pageId="1" pageNumber="152">z1</emphasis>
and
<emphasis id="B968EA81DE031E385C46725AFD408771" box="[629,658,1583,1609]" italics="true" pageId="1" pageNumber="152">Z5</emphasis>
foliate-dentate and shorter. Adult female sternal shield smooth almost throughout; anterior margin hardly conspicuous and posterior margin concave, bearing three pairs of setae and two pairs of large slit shape pore-like structures,
<emphasis id="B968EA81DE031E385CF27203FD3087A8" box="[705,738,1654,1680]" italics="true" pageId="1" pageNumber="152">st1</emphasis>
long enough to reach base of
<emphasis id="B968EA81DE031E385A097203FB8887A8" box="[1082,1114,1654,1680]" italics="true" pageId="1" pageNumber="152">st2</emphasis>
, ratio of shield length/width (at broadest level) 1. Genital shield ratio of length/width (at broadest level) 2.4, surface without any distinct reticulation, almost≃completely smooth, except two diagonal lines in posterior part ≃ which confluence with another horizontal line and forming semi-triangular in posterior part of surface.Anal shield ratio of length/width (at broadest level) 1.4, cribrum well developed. Peritreme long, reaching mid-level of coxa I (near
<emphasis id="B968EA81DE031E385A5F7373FB578618" box="[1132,1157,1798,1824]" italics="true" pageId="1" pageNumber="152">s1</emphasis>
). Fixed digit of chelicera with 10 ≃11 teeth of various sizes. Some dorsal setae on tibia and genu of legs IIV and
<emphasis id="B968EA81DE031E385AB8735FFB67867C" box="[1163,1205,1834,1860]" italics="true" pageId="1" pageNumber="152">pd2</emphasis>
,
<emphasis id="B968EA81DE031E385AF6735FFB3C867C" box="[1221,1262,1834,1860]" italics="true" pageId="1" pageNumber="152">pd3</emphasis>
on tarsi IIIIV feather-like. Male with holoventral shield; shield with a strongly sclerotised median circular shape projection; spermatodactyl short and apically curved dorsally;
<emphasis id="B968EA81DE031E385C917307FD1886B4" box="[674,714,1906,1932]" italics="true" pageId="1" pageNumber="152">av1</emphasis>
on femur of leg II lanceolate,
<emphasis id="B968EA81DE031E385A287306FBE786B4" box="[1051,1077,1907,1932]" italics="true" pageId="1" pageNumber="152">pv</emphasis>
on tibia IV and
<emphasis id="B968EA81DE031E385ADA7307FAC086B4" box="[1257,1298,1906,1932]" italics="true" pageId="1" pageNumber="152">av2</emphasis>
on tarsus II, IV,
<emphasis id="B968EA81DE031E385E8C73E3FF358688" box="[191,231,1942,1968]" italics="true" pageId="1" pageNumber="152">pv2</emphasis>
on tarsus III, IV spur-like (apically blunt) and trochanter bearing a large bump shape spur (
<figureCitation id="13272A16DE031E385AD473E3FAE98688" box="[1255,1339,1942,1968]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="1" pageNumber="152">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE031E315EF473CCFED280B3" lastPageId="8" lastPageNumber="159" pageId="1" pageNumber="152" type="description">
<paragraph id="8BA33693DE031E3B5EF473CCFD188097" blockId="1.[151,1437,1329,2041]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="153" pageId="1" pageNumber="152">
<emphasis id="B968EA81DE031E385EF473CCFDD186EC" bold="true" box="[199,515,1977,2005]" pageId="1" pageNumber="152">Description. Female (n=4).</emphasis>
<emphasis id="B968EA81DE031E385C3973CFFD1786EC" box="[522,709,1978,2004]" italics="true" pageId="1" pageNumber="152">Dorsal idiosoma</emphasis>
(
<figureCitation id="13272A16DE031E385CE073CFFCCB86EC" box="[723,793,1978,2004]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="1" pageNumber="152">Figs 1</figureCitation>
,
<figureCitation id="13272A16DE031E385D1473CEFCB586EC" box="[807,871,1978,2005]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="1" pageNumber="152">7 &amp; 9</figureCitation>
). Dorsal shield 295305 long, 180190 wide, covering most of dorsal idiosoma, well reticulated over whole surface, narrowing from level of setae
<emphasis id="B968EA81DE031E385AED73ABFB2A86C0" box="[1246,1272,2014,2040]" italics="true" pageId="1" pageNumber="152">r3</emphasis>
, progressively tapering until rounded posteriorly. Shield with 39 pairs of oblong-mucronate setae, 22 pairs of podonotal setae (
<emphasis id="B968EA81DE001E3B5E9274C8FF0081EF" box="[161,210,189,215]" italics="true" pageId="2" pageNumber="153">j16</emphasis>
;
<emphasis id="B968EA81DE001E3B5ED274C8FEC481EF" box="[225,278,189,215]" italics="true" pageId="2" pageNumber="153">z16</emphasis>
;
<emphasis id="B968EA81DE001E3B5F1674C8FE8881EF" box="[293,346,189,215]" italics="true" pageId="2" pageNumber="153">s16</emphasis>
;
<emphasis id="B968EA81DE001E3B5F5B74C8FE4F81EF" box="[360,413,189,215]" italics="true" pageId="2" pageNumber="153">r25</emphasis>
), and 17 pairs of opisthonotal setae (
<emphasis id="B968EA81DE001E3B5D0F74C8FCA081EF" box="[828,882,189,215]" italics="true" pageId="2" pageNumber="153">J15</emphasis>
;
<emphasis id="B968EA81DE001E3B5DB274C8FC6981EF" box="[897,955,189,215]" italics="true" pageId="2" pageNumber="153">Z15</emphasis>
;
<emphasis id="B968EA81DE001E3B5DFA74C8FBD381EF" box="[969,1025,189,215]" italics="true" pageId="2" pageNumber="153">S15</emphasis>
and two pairs of
<emphasis id="B968EA81DE001E3B5AF774CBFB3281EF" box="[1220,1248,190,215]" italics="true" pageId="2" pageNumber="153">Zx</emphasis>
setae) except
<emphasis id="B968EA81DE001E3B5B4C74C8FA4781EF" box="[1407,1429,189,215]" italics="true" pageId="2" pageNumber="153">j1</emphasis>
,
<emphasis id="B968EA81DE001E3B5EA47494FF6281C3" box="[151,176,225,251]" italics="true" pageId="2" pageNumber="153">z1</emphasis>
and
<emphasis id="B968EA81DE001E3B5ED57494FED681C3" box="[230,260,225,251]" italics="true" pageId="2" pageNumber="153">Z5</emphasis>
foliate-dentate; two unpaired supernumerary setae (
<emphasis id="B968EA81DE001E3B5D7F7497FCB781C3" box="[844,869,226,251]" italics="true" pageId="2" pageNumber="153">Jx</emphasis>
) present (
<figureCitation id="13272A16DE001E3B5DE67494FBC981C3" box="[981,1051,225,251]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 1</figureCitation>
,
<figureCitation id="13272A16DE001E3B5A1A7494FBB981C3" box="[1065,1131,225,251]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">7 &amp; 9</figureCitation>
). Almost all setae with 24 minute barbs (
<figureCitation id="13272A16DE001E3B5F087570FE538027" box="[315,385,261,287]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 9</figureCitation>
) and uniform in length (2126) and thickness, except
<emphasis id="B968EA81DE001E3B5DC37570FBDF8027" box="[1008,1037,261,287]" italics="true" pageId="2" pageNumber="153">Z5</emphasis>
(1315),
<emphasis id="B968EA81DE001E3B5A4C7570FB458027" box="[1151,1175,261,287]" italics="true" pageId="2" pageNumber="153">z1</emphasis>
(810) and
<emphasis id="B968EA81DE001E3B5B167570FAE88027" box="[1317,1338,261,287]" italics="true" pageId="2" pageNumber="153">j1</emphasis>
(1517) length of setae slightly increasing from central to lateral (
<figureCitation id="13272A16DE001E3B5D16755CFCBF807B" box="[805,877,297,323]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 1</figureCitation>
,
<figureCitation id="13272A16DE001E3B5D4E755CFC13807B" box="[893,961,297,323]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">7 &amp; 9</figureCitation>
). Narrow strip of soft cuticle surrounding dorsal shield with two pairs of
<emphasis id="B968EA81DE001E3B5FCB753BFDDB805F" box="[504,521,334,359]" italics="true" pageId="2" pageNumber="153">R</emphasis>
setae (
<emphasis id="B968EA81DE001E3B5C697538FDAB805F" box="[602,633,333,359]" italics="true" pageId="2" pageNumber="153">R3</emphasis>
,
<emphasis id="B968EA81DE001E3B5CBA7538FD7A805F" box="[649,680,333,359]" italics="true" pageId="2" pageNumber="153">R4</emphasis>
). Shield with about 16 pairs of discernible pore-like structures, including 15 poroids (
<emphasis id="B968EA81DE001E3B5F4F7504FE7280B3" box="[380,416,369,395]" italics="true" pageId="2" pageNumber="153">id1</emphasis>
,
<emphasis id="B968EA81DE001E3B5F9D7504FE0080B3" box="[430,466,369,395]" italics="true" pageId="2" pageNumber="153">id2</emphasis>
,
<emphasis id="B968EA81DE001E3B5FD27504FDD780B3" box="[481,517,369,395]" italics="true" pageId="2" pageNumber="153">id4</emphasis>
,
<emphasis id="B968EA81DE001E3B5C277504FDE580B3" box="[532,567,369,395]" italics="true" pageId="2" pageNumber="153">id5</emphasis>
,
<emphasis id="B968EA81DE001E3B5C757504FDB880B3" box="[582,618,369,395]" italics="true" pageId="2" pageNumber="153">id6</emphasis>
,
<emphasis id="B968EA81DE001E3B5C4A7504FD6280B3" box="[633,688,369,395]" italics="true" pageId="2" pageNumber="153">idm1</emphasis>
,
<emphasis id="B968EA81DE001E3B5CF37504FD2580B3" box="[704,759,369,395]" italics="true" pageId="2" pageNumber="153">idm2</emphasis>
,
<emphasis id="B968EA81DE001E3B5D357504FCED80B3" box="[774,831,369,395]" italics="true" pageId="2" pageNumber="153">idm3</emphasis>
,
<emphasis id="B968EA81DE001E3B5D7E7504FC5780B3" box="[845,901,369,395]" italics="true" pageId="2" pageNumber="153">idm4</emphasis>
,
<emphasis id="B968EA81DE001E3B5DA77504FC1E80B3" box="[916,972,369,395]" italics="true" pageId="2" pageNumber="153">idm5</emphasis>
,
<emphasis id="B968EA81DE001E3B5DE87504FC2F80B3" box="[987,1021,369,395]" italics="true" pageId="2" pageNumber="153">idx</emphasis>
,
<emphasis id="B968EA81DE001E3B5A3F7504FBE580B3" box="[1036,1079,369,395]" italics="true" pageId="2" pageNumber="153">idl3</emphasis>
) and four gland openings (
<emphasis id="B968EA81DE001E3B5B587504FA4780B3" box="[1387,1429,369,395]" italics="true" pageId="2" pageNumber="153">gd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5EA475E0FF138097" box="[151,193,405,431]" italics="true" pageId="2" pageNumber="153">gd2</emphasis>
,
<emphasis id="B968EA81DE001E3B5EFC75E0FF288097" box="[207,250,405,431]" italics="true" pageId="2" pageNumber="153">gd5</emphasis>
,
<emphasis id="B968EA81DE001E3B5F3475E0FEE38097" box="[263,305,405,431]" italics="true" pageId="2" pageNumber="153">gd6</emphasis>
), others indistinct (
<figureCitation id="13272A16DE001E3B5C3875E0FDA48097" box="[523,630,405,431]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figures 1</figureCitation>
and
<figureCitation id="13272A16DE001E3B5C9F75E0FD6E8097" box="[684,700,405,431]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">7</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE001E3B5EF475CDFDBB859B" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="153">
<emphasis id="B968EA81DE001E3B5EF475CDFE5880EB" bold="true" box="[199,394,440,467]" italics="true" pageId="2" pageNumber="153">Ventral idiosoma</emphasis>
(
<figureCitation id="13272A16DE001E3B5FAB75CCFE0D80EB" box="[408,479,441,467]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 2</figureCitation>
&amp;
<figureCitation id="13272A16DE001E3B5C3275CCFDC380EB" box="[513,529,441,467]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">8</figureCitation>
). Tritosternum with paired pilose laciniae (5358), fused basally (24), columnar base 2024 × 811 wide; presternal area poorly sclerotised. Sternal shield (length 8690) narrowest between coxae II (5860), widest (8890), anterior margin completely indistinct and posterior margin concave; shield with three pairs of smooth pointed setae (
<emphasis id="B968EA81DE001E3B5FC47650FDCA8307" box="[503,536,549,575]" italics="true" pageId="2" pageNumber="153">st1</emphasis>
3234,
<emphasis id="B968EA81DE001E3B5C457650FD448307" box="[630,662,549,575]" italics="true" pageId="2" pageNumber="153">st2</emphasis>
2224,
<emphasis id="B968EA81DE001E3B5CC67650FCC48307" box="[757,790,549,575]" italics="true" pageId="2" pageNumber="153">st3</emphasis>
2224,
<emphasis id="B968EA81DE001E3B5D477650FC468307" box="[884,916,549,575]" italics="true" pageId="2" pageNumber="153">st1</emphasis>
reaching base of next setae) and two pairs of large slit-shaped pore-like structures (
<emphasis id="B968EA81DE001E3B5C77763CFDB4835B" box="[580,614,585,611]" italics="true" pageId="2" pageNumber="153">iv1</emphasis>
adjacent to setae
<emphasis id="B968EA81DE001E3B5D01763CFC81835B" box="[818,851,585,611]" italics="true" pageId="2" pageNumber="153">st1</emphasis>
;
<emphasis id="B968EA81DE001E3B5D50763CFC57835B" box="[867,901,585,611]" italics="true" pageId="2" pageNumber="153">iv2</emphasis>
between
<emphasis id="B968EA81DE001E3B5DC0763CFBC1835B" box="[1011,1043,585,611]" italics="true" pageId="2" pageNumber="153">st2</emphasis>
and
<emphasis id="B968EA81DE001E3B5A7F763CFBBF835B" box="[1100,1133,585,611]" italics="true" pageId="2" pageNumber="153">st3</emphasis>
); sternal shield smooth almost throughout and without distinct reticulation. Metasternal setae
<emphasis id="B968EA81DE001E3B5DA27618FC6383BF" box="[913,945,621,647]" italics="true" pageId="2" pageNumber="153">st4</emphasis>
(1921) and metasternal poroids located on soft integument; metasternal platelets absent. Endopodal plates II/III completely fused to sternal shield, endopodal plates III/IV elongate, narrow and curved. Genital shield broad, flask-shaped, not adjacent to anal shield, expanded laterally behind setae
<emphasis id="B968EA81DE001E3B5FB876ACFE7983CB" box="[395,427,729,755]" italics="true" pageId="2" pageNumber="153">st5</emphasis>
, length 137139, maximum width 5458, posterior margin rounded, shield smooth almost throughout, except two diagonal lines in posterior part that fuse with another horizontal line and forming a faintly semi-triangular shape; bearing a pair of simple setae
<emphasis id="B968EA81DE001E3B5CC87754FCCE8203" box="[763,796,801,827]" italics="true" pageId="2" pageNumber="153">st5</emphasis>
(1921); paragenital poroids
<emphasis id="B968EA81DE001E3B5A407754FB478203" box="[1139,1173,801,827]" italics="true" pageId="2" pageNumber="153">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="B968EA81DE001E3B5FF27730FE338267" box="[449,481,837,863]" italics="true" pageId="2" pageNumber="153">st5</emphasis>
. Shield flanked by two pairs of irregular small platelets. Anal shield subtriangular, rounded anteriorly, length 5557, width 4044, anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (1416), cribrum consisting of a terminal tuft and pair of anterior arms reaching about to midlevel of distance between post-anal and para-anal setae; anal poroids
<emphasis id="B968EA81DE001E3B5D8677C4FC0F82F3" box="[949,989,945,971]" italics="true" pageId="2" pageNumber="153">gv3</emphasis>
on anterolateral margin of anal shield. Soft opisthogastric cuticle surrounding genital and anal shields with 12 pairs of subequal setae (1216) (
<emphasis id="B968EA81DE001E3B5B0977A0FA4482D7" box="[1338,1430,981,1007]" italics="true" pageId="2" pageNumber="153">Jv1Jv5</emphasis>
,
<emphasis id="B968EA81DE001E3B5EA4778CFF2B852B" box="[151,249,1017,1043]" italics="true" pageId="2" pageNumber="153">Zv1Zv5</emphasis>
,
<emphasis id="B968EA81DE001E3B5F34778CFEE8852B" box="[263,314,1017,1043]" italics="true" pageId="2" pageNumber="153">UR1</emphasis>
,
<emphasis id="B968EA81DE001E3B5F74778CFEA9852B" box="[327,379,1017,1043]" italics="true" pageId="2" pageNumber="153">UR2</emphasis>
). Setae
<emphasis id="B968EA81DE001E3B5FE7778CFE29852B" box="[468,507,1017,1043]" italics="true" pageId="2" pageNumber="153">Jv5</emphasis>
,
<emphasis id="B968EA81DE001E3B5C3B778CFDE0852B" box="[520,562,1017,1043]" italics="true" pageId="2" pageNumber="153">Zv5</emphasis>
and
<emphasis id="B968EA81DE001E3B5C5B778CFD49852B" box="[616,667,1017,1043]" italics="true" pageId="2" pageNumber="153">UR2</emphasis>
spatulate, metapodal plates absent. Exopodal and parapodal platelets not fused, strip-like, extending narrowly behind coxae IV. Peritreme extending anteriorly to mid-level of coxa I; peritrematal shield narrow, free from exopodal shields, protrusion band of cuticle lateral to the peritreme at the level of coxae IIIII, a lyrifissure
<emphasis id="B968EA81DE001E3B5FE47013FE3F8547" box="[471,493,1126,1151]" italics="true" pageId="2" pageNumber="153">ip</emphasis>
at level of coxa II, post-stigmatic section bearing two pore-like structures, a lyrifissures
<emphasis id="B968EA81DE001E3B5EEB70FFFF3C859B" box="[216,238,1162,1187]" italics="true" pageId="2" pageNumber="153">ip</emphasis>
and a gland pore
<emphasis id="B968EA81DE001E3B5F8670FFFE03859B" box="[437,465,1162,1187]" italics="true" pageId="2" pageNumber="153">gp</emphasis>
(
<figureCitation id="13272A16DE001E3B5FD370FCFDF5859B" box="[480,551,1161,1187]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 2</figureCitation>
&amp;
<figureCitation id="13272A16DE001E3B5C7870FCFD89859B" box="[587,603,1161,1187]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">8</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE001E3B5EF470D9FBF08734" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="153">
<taxonomicName id="4C1C4D10DE001E3B5EF470D9FE8985FF" authorityName="Krefft" authorityYear="1871" box="[199,347,1196,1223]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="153" phylum="Nematoda" rank="genus">
<emphasis id="B968EA81DE001E3B5EF470D9FE8985FF" bold="true" box="[199,347,1196,1223]" italics="true" pageId="2" pageNumber="153">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13272A16DE001E3B5F5A70D8FE1C85FF" box="[361,462,1197,1223]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 46</figureCitation>
,
<figureCitation id="13272A16DE001E3B5FEB70D8FDF185FF" box="[472,547,1197,1223]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">1012</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with about 1125 small teeth (
<figureCitation id="13272A16DE001E3B5F4470A4FE1385D3" box="[375,449,1233,1259]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 4</figureCitation>
&amp;
<figureCitation id="13272A16DE001E3B5FDF70A4FDD885D3" box="[492,522,1233,1259]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">10</figureCitation>
), groove wider anteriorly. Hypostome with four pairs of setae,
<emphasis id="B968EA81DE001E3B5ADC70A4FAD985D3" box="[1263,1291,1233,1259]" italics="true" pageId="2" pageNumber="153">h1</emphasis>
(2830),
<emphasis id="B968EA81DE001E3B5BB370A4FA4F85D3" box="[1408,1437,1233,1259]" italics="true" pageId="2" pageNumber="153">h2</emphasis>
(1012),
<emphasis id="B968EA81DE001E3B5F327080FECF8437" box="[257,285,1269,1295]" italics="true" pageId="2" pageNumber="153">h3</emphasis>
(2830), palpcoxal setae (
<emphasis id="B968EA81DE001E3B5C677083FDBC8437" box="[596,622,1270,1295]" italics="true" pageId="2" pageNumber="153">pc</emphasis>
) (2426) (
<figureCitation id="13272A16DE001E3B5CDE7080FCE58437" box="[749,823,1269,1295]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 4</figureCitation>
&amp;
<figureCitation id="13272A16DE001E3B5D527080FC528437" box="[865,896,1269,1295]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">10</figureCitation>
). Epistome trapezoidal, central part irregularly denticulate (
<figureCitation id="13272A16DE001E3B5F11716CFEBE840B" box="[290,364,1304,1331]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Figs 5</figureCitation>
,
<figureCitation id="13272A16DE001E3B5F49716CFE45840B" box="[378,407,1305,1331]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">11</figureCitation>
). Corniculi robust and horn-like, extending slightly beyond palp trochanter. Internal malae with median and lateral projections, fringed, inner lobes touching with outer lobes; labrum with pilose surface. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except
<emphasis id="B968EA81DE001E3B5B577114FAA88443" box="[1380,1402,1377,1403]" italics="true" pageId="2" pageNumber="153">al</emphasis>
on palp femur and genu thickened and apically spatulate, palp tarsal apotele two-tined (
<figureCitation id="13272A16DE001E3B5A0471F0FB5B84A7" box="[1079,1161,1413,1439]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 12</figureCitation>
). Fixed digit of chelicera with an offset and most distal large tooth (gabelzhan) followed by 910 variously sized teeth, 4nd, 6th, 9th larger, a setaceous pilus dentilis, dorsal cheliceral setae prostrate, arthrodial membrane with a rounded flap and normal filaments and cheliceral lyrifissures indistinct, movable digit with two teeth (
<figureCitation id="13272A16DE001E3B5DFC7184FBC68734" box="[975,1044,1521,1548]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="2" pageNumber="153">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BA33693DE001E3F5EF47260FAD681EF" blockId="2.[151,1437,153,2015]" lastBlockId="6.[151,1437,153,611]" lastPageId="6" lastPageNumber="157" pageId="2" pageNumber="153">
<emphasis id="B968EA81DE001E3B5EF47260FF2F8717" bold="true" box="[199,253,1557,1583]" italics="true" pageId="2" pageNumber="153">Legs</emphasis>
(
<figureCitation id="13272A16DE001E3B5F3F7260FE5D8717" box="[268,399,1557,1583]" captionStart-0="FIGURES 714" captionStart-1="FIGURE 1518" captionStartId-0="4.[151,264,1945,1970]" captionStartId-1="5.[151,250,1910,1935]" captionTargetBox-0="[180,1407,182,1918]" captionTargetBox-1="[162,1425,192,1867]" captionTargetId-0="figure@4.[179,1409,181,1920]" captionTargetId-1="figure@5.[151,1436,181,1885]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." captionText-1="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.3687708" figureDoi-1="http://doi.org/10.5281/zenodo.3687710" httpUri-0="https://zenodo.org/record/3687708/files/figure.png" httpUri-1="https://zenodo.org/record/3687710/files/figure.png" pageId="2" pageNumber="153">Figs 1318</figureCitation>
). Legs II and III short (213217, 195204), I and IV longer (273296, 294301). Chaetotaxy normal for free-living
<taxonomicName id="4C1C4D10DE001E3B5FAA724CFDCA876B" box="[409,536,1593,1619]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="153" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="13272A16DE001E3B5C46724CFD19876B" box="[629,715,1592,1619]" captionStart="FIGURE 1518" captionStartId="5.[151,250,1910,1935]" captionTargetBox="[162,1425,192,1867]" captionTargetId="figure@5.[151,1436,181,1885]" captionTargetPageId="5" captionText="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687710" httpUri="https://zenodo.org/record/3687710/files/figure.png" pageId="2" pageNumber="153">Fig. 15</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/2, 1/1-1, femur 2-2/1, 3/3-2 (
<emphasis id="B968EA81DE001E3B5E927228FF1B874F" box="[161,201,1629,1655]" italics="true" pageId="2" pageNumber="153">pv1</emphasis>
,
<emphasis id="B968EA81DE001E3B5EEA7228FED3874F" box="[217,257,1629,1655]" italics="true" pageId="2" pageNumber="153">pv2</emphasis>
,
<emphasis id="B968EA81DE001E3B5F287228FE97874F" box="[283,325,1629,1655]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5F657228FE52874F" box="[342,384,1629,1655]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5FA37228FE68874F" box="[400,442,1629,1655]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
,
<emphasis id="B968EA81DE001E3B5FF87228FE26874F" box="[459,500,1629,1655]" italics="true" pageId="2" pageNumber="153">pd3</emphasis>
thickened, all dorsal setae with minute barbs), genu 2-3/2, 3/1-2 (all dorsal setae feather-like, lateral and ventral setae thickened except
<emphasis id="B968EA81DE001E3B5D3072F4FCF987A3" box="[771,811,1665,1691]" italics="true" pageId="2" pageNumber="153">av2</emphasis>
), tibia 2-3/2, 3/1-2 (all dorsal setae feather-like, lateral setae thickened). Leg II (
<figureCitation id="13272A16DE001E3B5F8372D0FDD687F8" box="[432,516,1701,1728]" captionStart="FIGURE 1518" captionStartId="5.[151,250,1910,1935]" captionTargetBox="[162,1425,192,1867]" captionTargetId="figure@5.[151,1436,181,1885]" captionTargetPageId="5" captionText="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687710" httpUri="https://zenodo.org/record/3687710/files/figure.png" pageId="2" pageNumber="153">Fig. 16</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1, femur 2-3/1, 2/2-1 (
<emphasis id="B968EA81DE001E3B5AE172D0FB2E8787" box="[1234,1276,1701,1727]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
truncate,
<emphasis id="B968EA81DE001E3B5B5872D0FA448787" box="[1387,1430,1701,1727]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5EA472BCFF1387DB" box="[151,193,1737,1763]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
falcate and with minute barbs), genu 2-3/1, 2/1-2 (
<emphasis id="B968EA81DE001E3B5CCF72BCFCF487DB" box="[764,806,1737,1763]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5D0772BCFC8D87DB" box="[820,863,1737,1763]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
,
<emphasis id="B968EA81DE001E3B5D5F72BCFC4487DB" box="[876,918,1737,1763]" italics="true" pageId="2" pageNumber="153">ad3</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5A1F72BCFB8487DB" box="[1068,1110,1737,1763]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5A5772BCFB5C87DB" box="[1124,1166,1737,1763]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
falcate and with minute barbs), tibia 2-2/1, 2/1-2 (
<emphasis id="B968EA81DE001E3B5F8A7298FE31863F" box="[441,483,1773,1799]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5FC27298FDCE863F" box="[497,540,1773,1799]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5C827298FD0E863F" box="[689,732,1773,1799]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5CDA7298FCC1863F" box="[745,787,1773,1799]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
thickened and with minute barbs). Leg III (
<figureCitation id="13272A16DE001E3B5ACC7298FA80863F" box="[1279,1362,1773,1799]" captionStart="FIGURE 1518" captionStartId="5.[151,250,1910,1935]" captionTargetBox="[162,1425,192,1867]" captionTargetId="figure@5.[151,1436,181,1885]" captionTargetPageId="5" captionText="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687710" httpUri="https://zenodo.org/record/3687710/files/figure.png" pageId="2" pageNumber="153">Fig. 17</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1, femur 1-2/1, 1/0-1 (
<emphasis id="B968EA81DE001E3B5D397364FCE18613" box="[778,819,1809,1835]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5D737364FC8E8613" box="[832,860,1809,1835]" italics="true" pageId="2" pageNumber="153">pd</emphasis>
,
<emphasis id="B968EA81DE001E3B5D5A7364FCAD8613" box="[873,895,1809,1835]" italics="true" pageId="2" pageNumber="153">pl</emphasis>
thickened,
<emphasis id="B968EA81DE001E3B5DCC7364FBFB8613" box="[1023,1065,1809,1835]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
apically blunt), genu 2-2/1, 2/1-1 (
<emphasis id="B968EA81DE001E3B5E927340FF198677" box="[161,203,1845,1871]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5EEA7340FED18677" box="[217,259,1845,1871]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5FA97340FE168677" box="[410,452,1845,1871]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5FE07340FE2C8677" box="[467,510,1845,1871]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
lanceolate, with minute barbs,
<emphasis id="B968EA81DE001E3B5D537340FCA48677" box="[864,886,1845,1871]" italics="true" pageId="2" pageNumber="153">pl</emphasis>
thickened,
<figureCitation id="13272A16DE001E3B5DCA7340FB9E8677" box="[1017,1100,1845,1871]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 13</figureCitation>
), tibia: 2-1/1, 2/1-1 (
<emphasis id="B968EA81DE001E3B5B0A7340FAB68677" box="[1337,1380,1845,1871]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5B417340FA4E8677" box="[1394,1436,1845,1871]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5F17732CFE9C864B" box="[292,334,1881,1907]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
lanceolate, with minute barbs,
<emphasis id="B968EA81DE001E3B5C95732CFD6E864B" box="[678,700,1881,1907]" italics="true" pageId="2" pageNumber="153">pl</emphasis>
thickened,
<figureCitation id="13272A16DE001E3B5D08732CFC5E864B" box="[827,908,1881,1907]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 13</figureCitation>
). Leg IV (
<figureCitation id="13272A16DE001E3B5DC8732CFB9F864B" box="[1019,1101,1881,1907]" captionStart="FIGURE 1518" captionStartId="5.[151,250,1910,1935]" captionTargetBox="[162,1425,192,1867]" captionTargetId="figure@5.[151,1436,181,1885]" captionTargetPageId="5" captionText="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687710" httpUri="https://zenodo.org/record/3687710/files/figure.png" pageId="2" pageNumber="153">Fig. 17</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/1-1 (
<emphasis id="B968EA81DE001E3B5F1E7308FE9B86AF" box="[301,329,1917,1943]" italics="true" pageId="2" pageNumber="153">ad</emphasis>
spatulate), femur 1-2/1, 1/0-1 (
<emphasis id="B968EA81DE001E3B5C9D7308FD0586AF" box="[686,727,1917,1943]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5CD57308FCC386AF" box="[742,785,1917,1943]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
spatulate,
<emphasis id="B968EA81DE001E3B5DB97308FC7486AF" box="[906,934,1917,1943]" italics="true" pageId="2" pageNumber="153">pd</emphasis>
thickened with minute barbs,
<figureCitation id="13272A16DE001E3B5ACD7308FA8286AF" box="[1278,1360,1917,1943]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 14</figureCitation>
), genu 2-2/1, 3/0-1 (
<emphasis id="B968EA81DE001E3B5F1F73D4FE848683" box="[300,342,1953,1979]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5F5773D4FE5C8683" box="[356,398,1953,1979]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
,
<emphasis id="B968EA81DE001E3B5FAF73D4FE148683" box="[412,454,1953,1979]" italics="true" pageId="2" pageNumber="153">pd3</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5C6F73D4FD548683" box="[604,646,1953,1979]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5CA773D4FD6C8683" box="[660,702,1953,1979]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
lanceolate, with minute barbs,
<emphasis id="B968EA81DE001E3B5A2D73D7FBEA8683" box="[1054,1080,1954,1979]" italics="true" pageId="2" pageNumber="153">av</emphasis>
thickened,
<figureCitation id="13272A16DE001E3B5A8973D4FADE8683" box="[1210,1292,1953,1979]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 14</figureCitation>
), tibia 2-1/1, 3/1-2 (
<emphasis id="B968EA81DE001E3B5ED073B0FEDF86E7" box="[227,269,1989,2015]" italics="true" pageId="2" pageNumber="153">pd1</emphasis>
,
<emphasis id="B968EA81DE001E3B5F2873B0FE9786E7" box="[283,325,1989,2015]" italics="true" pageId="2" pageNumber="153">pd2</emphasis>
,
<emphasis id="B968EA81DE001E3B5F6073B0FEAF86E7" box="[339,381,1989,2015]" italics="true" pageId="2" pageNumber="153">pd3</emphasis>
feather-like,
<emphasis id="B968EA81DE001E3B5C2073B0FDEF86E7" box="[531,573,1989,2015]" italics="true" pageId="2" pageNumber="153">ad1</emphasis>
,
<emphasis id="B968EA81DE001E3B5C7873B0FDA786E7" box="[587,629,1989,2015]" italics="true" pageId="2" pageNumber="153">ad2</emphasis>
lanceolate, with minute barbs,
<emphasis id="B968EA81DE001E3B5DE673B3FC3D86E7" box="[981,1007,1990,2015]" italics="true" pageId="2" pageNumber="153">av</emphasis>
,
<emphasis id="B968EA81DE001E3B5DCE73B0FBC186E7" box="[1021,1043,1989,2015]" italics="true" pageId="2" pageNumber="153">al</emphasis>
,
<emphasis id="B968EA81DE001E3B5A1273B3FBE986E7" box="[1057,1083,1990,2015]" italics="true" pageId="2" pageNumber="153">pv</emphasis>
thickened,
<figureCitation id="13272A16DE001E3B5A8D73B0FAC286E7" box="[1214,1296,1989,2015]" captionStart="FIGURES 714" captionStartId="4.[151,264,1945,1970]" captionTargetBox="[180,1407,182,1918]" captionTargetId="figure@4.[179,1409,181,1920]" captionTargetPageId="4" captionText="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." figureDoi="http://doi.org/10.5281/zenodo.3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="2" pageNumber="153">Fig. 14</figureCitation>
). Tarsi II-IV with 18 setae (3-3/2, 3/2-3 +
<emphasis id="B968EA81DE041E3F5FEA74EFFE28818B" box="[473,506,154,179]" italics="true" pageId="6" pageNumber="157">mv</emphasis>
,
<emphasis id="B968EA81DE041E3F5C3474ECFDFB818B" box="[519,553,153,179]" italics="true" pageId="6" pageNumber="157">md</emphasis>
). All pretarsi with well-developed paired claws and rounded pulvilli and a long thin stalk, ventral and lateral setae on tarsus II and III thickened,
<emphasis id="B968EA81DE041E3F5D5C74C8FC4B81EF" box="[879,921,189,215]" italics="true" pageId="6" pageNumber="157">pd2</emphasis>
,
<emphasis id="B968EA81DE041E3F5D9474C8FC0381EF" box="[935,977,189,215]" italics="true" pageId="6" pageNumber="157">pd3</emphasis>
on tarsi III-IV feather-like.
</paragraph>
<caption id="DF63661BDE011E3A5EA473ECFD9786EE" ID-DOI="http://doi.org/10.5281/zenodo.3687706" ID-Zenodo-Dep="3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="3" pageNumber="154" startId="3.[151,264,1945,1970]" targetBox="[182,1399,191,1905]" targetPageId="3">
<paragraph id="8BA33693DE011E3A5EA473ECFD9786EE" blockId="3.[151,1436,1945,2006]" pageId="3" pageNumber="154">
<emphasis id="B968EA81DE011E3A5EA473ECFEE8868A" bold="true" box="[151,314,1945,1970]" pageId="3" pageNumber="154">FIGURES 16.</emphasis>
<taxonomicName id="4C1C4D10DE011E3A5F7373EFFD98868A" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[320,586,1946,1970]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE011E3A5F7373EFFD98868A" box="[320,586,1946,1970]" italics="true" pageId="3" pageNumber="154">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE011E3A5C6373EFFD72868A" bold="true" box="[592,672,1946,1970]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="A25B57FADE011E3A5C6373EFFD72868A" box="[592,672,1946,1970]" pageId="3" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera.
</paragraph>
</caption>
<caption id="DF63661BDE061E3D5EA473ECFD1386C1" ID-DOI="http://doi.org/10.5281/zenodo.3687708" ID-Zenodo-Dep="3687708" httpUri="https://zenodo.org/record/3687708/files/figure.png" pageId="4" pageNumber="155" startId="4.[151,264,1945,1970]" targetBox="[180,1407,182,1918]" targetPageId="4">
<paragraph id="8BA33693DE061E3D5EA473ECFD1386C1" blockId="4.[151,1437,1945,2042]" pageId="4" pageNumber="155">
<emphasis id="B968EA81DE061E3D5EA473ECFE95868A" bold="true" box="[151,327,1945,1970]" pageId="4" pageNumber="155">FIGURES 714.</emphasis>
DIC micrographs of
<taxonomicName id="4C1C4D10DE061E3D5C1373EFFCF9868A" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[544,811,1946,1970]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE061E3D5C1373EFFCF9868A" box="[544,811,1946,1970]" italics="true" pageId="4" pageNumber="155">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE061E3D5D0273EFFC53868A" bold="true" box="[817,897,1946,1970]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="A25B57FADE061E3D5D0273EFFC53868A" box="[817,897,1946,1970]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV.
</paragraph>
</caption>
<caption id="DF63661BDE071E3C5EA47303FAB486B7" ID-DOI="http://doi.org/10.5281/zenodo.3687710" ID-Zenodo-Dep="3687710" box="[151,1382,1910,1935]" httpUri="https://zenodo.org/record/3687710/files/figure.png" pageId="5" pageNumber="156" startId="5.[151,250,1910,1935]" targetBox="[162,1425,192,1867]" targetPageId="5">
<paragraph id="8BA33693DE071E3C5EA47303FAB486B7" blockId="5.[151,1382,1910,1935]" box="[151,1382,1910,1935]" pageId="5" pageNumber="156">
<emphasis id="B968EA81DE071E3C5EA47303FE9486B7" bold="true" box="[151,326,1910,1935]" pageId="5" pageNumber="156">FIGURE 1518.</emphasis>
<taxonomicName id="4C1C4D10DE071E3C5F7F7302FD8586B7" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[332,599,1911,1935]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE071E3C5F7F7302FD8586B7" box="[332,599,1911,1935]" italics="true" pageId="5" pageNumber="156">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE071E3C5C6E7302FD7C86B7" bold="true" box="[605,686,1911,1935]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A25B57FADE071E3C5C6E7302FD7C86B7" box="[605,686,1911,1935]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV.
</paragraph>
</caption>
<paragraph id="8BA33693DE041E3F5EF47494FD7F807B" blockId="6.[151,1437,153,611]" pageId="6" pageNumber="157">
<emphasis id="B968EA81DE041E3F5EF47494FE0E81C3" bold="true" box="[199,476,225,251]" italics="true" pageId="6" pageNumber="157">Insemination structures</emphasis>
(
<figureCitation id="13272A16DE041E3F5FDE7494FDE181C3" box="[493,563,225,251]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="6" pageNumber="157">Fig. 3</figureCitation>
). Laelapid-type sperm access system, tubulus long, wider at the solenostome level of coxa III and entering sacculus via a pair of circular openings. Sacculus reniform, the proximal ends of the tubulus slightly swollen at junction with ramus.
</paragraph>
<paragraph id="8BA33693DE041E3F5EF47538FE7480B3" blockId="6.[151,1437,153,611]" pageId="6" pageNumber="157">
<emphasis id="B968EA81DE041E3F5EF47538FE82805F" bold="true" box="[199,336,332,359]" pageId="6" pageNumber="157">Male (n=7).</emphasis>
(
<figureCitation id="13272A16DE041E3F5F6C7538FE33805F" box="[351,481,333,359]" captionStart-0="FIGURES 1920" captionStart-1="FIGURES 2124" captionStartId-0="6.[151,264,1987,2012]" captionStartId-1="7.[151,264,1945,1970]" captionTargetBox-0="[164,1421,650,1948]" captionTargetBox-1="[175,1409,187,1920]" captionTargetId-0="figure@6.[151,1435,633,1962]" captionTargetId-1="figure@7.[175,1413,181,1920]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 1920. Cosmolaelaps ceylonensis sp. nov., male. 19, ventral idiosoma; 20, chelicera." captionText-1="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi-0="http://doi.org/10.5281/zenodo.3687712" figureDoi-1="http://doi.org/10.5281/zenodo.3687714" httpUri-0="https://zenodo.org/record/3687712/files/figure.png" httpUri-1="https://zenodo.org/record/3687714/files/figure.png" pageId="6" pageNumber="157">Figs 1924</figureCitation>
).
<emphasis id="B968EA81DE041E3F5FC17539FD7C805F" bold="true" box="[498,686,332,359]" italics="true" pageId="6" pageNumber="157">Dorsal idiosoma</emphasis>
. Dorsal shield (268295) long, (154172) wide; ornamentation and chaetotaxy as in female.
</paragraph>
<paragraph id="8BA33693DE041E3F5EF475E1FBB3835B" blockId="6.[151,1437,153,611]" pageId="6" pageNumber="157">
<emphasis id="B968EA81DE041E3F5EF475E1FE598097" bold="true" box="[199,395,404,431]" italics="true" pageId="6" pageNumber="157">Ventral idiosoma</emphasis>
(
<figureCitation id="13272A16DE041E3F5FA875E0FE208097" box="[411,498,405,431]" captionStart="FIGURES 1920" captionStartId="6.[151,264,1987,2012]" captionTargetBox="[164,1421,650,1948]" captionTargetId="figure@6.[151,1435,633,1962]" captionTargetPageId="6" captionText="FIGURES 1920. Cosmolaelaps ceylonensis sp. nov., male. 19, ventral idiosoma; 20, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687712" httpUri="https://zenodo.org/record/3687712/files/figure.png" pageId="6" pageNumber="157">Figs 19</figureCitation>
,
<figureCitation id="13272A16DE041E3F5FCC75E0FDB28097" box="[511,608,405,431]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="6" pageNumber="157">21 &amp; 23</figureCitation>
). Sternal, genital, endopodal, ventral and anal shields fused into holoventral shield, weakly reticulate, more distinct behind
<emphasis id="B968EA81DE041E3F5CE575CCFCD680EB" box="[726,772,441,467]" italics="true" pageId="6" pageNumber="157">ZV1</emphasis>
, bearing
<emphasis id="B968EA81DE041E3F5D5C75CCFC7E80EB" box="[879,940,441,467]" italics="true" pageId="6" pageNumber="157">st15</emphasis>
, five pairs of opisthogastric setae (
<emphasis id="B968EA81DE041E3F5B0975CCFA4780EB" box="[1338,1429,441,467]" italics="true" pageId="6" pageNumber="157">Jv1Jv3</emphasis>
,
<emphasis id="B968EA81DE041E3F5EA475A8FF1380CF" box="[151,193,477,503]" italics="true" pageId="6" pageNumber="157">Zv1</emphasis>
,
<emphasis id="B968EA81DE041E3F5EFC75A8FF2B80CF" box="[207,249,477,503]" italics="true" pageId="6" pageNumber="157">Zv2</emphasis>
) in addition to circumanal setae; six pairs of poroids and a pore-like (
<emphasis id="B968EA81DE041E3F5A3275A8FBFB80CF" box="[1025,1065,477,503]" italics="true" pageId="6" pageNumber="157">gv3</emphasis>
) laterad of para-anal setae, gland pore
<emphasis id="B968EA81DE041E3F5EE17674FF288323" box="[210,250,513,539]" italics="true" pageId="6" pageNumber="157">gv2</emphasis>
behind coxa IV not discerned; shield with a strongly sclerotised median circular shape projection (
<figureCitation id="13272A16DE041E3F5B5C7674FF648307" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="6" pageNumber="157">Fig. 22</figureCitation>
), cribrum with 34 irregular rows of spicules and a pair of anterior arms reaching about to mid-level of distance between post-anal and para-anal setae. Soft opisthogastric cuticle with five pairs setae.
</paragraph>
<caption id="DF63661BDE041E3F5EA473B6FBBE86E4" ID-DOI="http://doi.org/10.5281/zenodo.3687712" ID-Zenodo-Dep="3687712" box="[151,1132,1987,2012]" httpUri="https://zenodo.org/record/3687712/files/figure.png" pageId="6" pageNumber="157" startId="6.[151,264,1987,2012]" targetBox="[164,1421,650,1948]" targetPageId="6">
<paragraph id="8BA33693DE041E3F5EA473B6FBBE86E4" blockId="6.[151,1132,1987,2012]" box="[151,1132,1987,2012]" pageId="6" pageNumber="157">
<emphasis id="B968EA81DE041E3F5EA473B6FE8686E4" bold="true" box="[151,340,1987,2012]" pageId="6" pageNumber="157">FIGURES 1920.</emphasis>
<taxonomicName id="4C1C4D10DE041E3F5F6973B6FDB786E3" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[346,613,1987,2011]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE041E3F5F6973B6FDB786E3" box="[346,613,1987,2011]" italics="true" pageId="6" pageNumber="157">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE041E3F5C5873B1FD6E86E4" bold="true" box="[619,700,1988,2012]" pageId="6" pageNumber="157">
<taxonomicNameLabel id="A25B57FADE041E3F5C5873B1FD6E86E4" box="[619,700,1988,2012]" pageId="6" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 19, ventral idiosoma; 20, chelicera.
</paragraph>
</caption>
<caption id="DF63661BDE051E3E5EA473ECFBA486ED" ID-DOI="http://doi.org/10.5281/zenodo.3687714" ID-Zenodo-Dep="3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,264,1945,1970]" targetBox="[175,1409,187,1920]" targetPageId="7">
<paragraph id="8BA33693DE051E3E5EA473ECFBA486ED" blockId="7.[151,1436,1945,2006]" pageId="7" pageNumber="158">
<emphasis id="B968EA81DE051E3E5EA473ECFE84868A" bold="true" box="[151,342,1945,1970]" pageId="7" pageNumber="158">FIGURES 2124.</emphasis>
DIC micrographs of
<taxonomicName id="4C1C4D10DE051E3E5C0473EFFC96868A" authority="Joharchi &amp; Ermilov &amp; Khaustov, 2020" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[567,836,1946,1970]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="B968EA81DE051E3E5C0473EFFC96868A" box="[567,836,1946,1970]" italics="true" pageId="7" pageNumber="158">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
<emphasis id="B968EA81DE051E3E5D7E73EFFC4D868A" bold="true" box="[845,927,1946,1970]" pageId="7" pageNumber="158">
<taxonomicNameLabel id="A25B57FADE051E3E5D7E73EFFC4D868A" box="[845,927,1946,1970]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II.
</paragraph>
</caption>
<paragraph id="8BA33693DE0A1E315EF474EDFC578026" blockId="8.[151,1437,152,1403]" pageId="8" pageNumber="159">
<taxonomicName id="4C1C4D10DE0A1E315EF474EDFE89818B" authorityName="Krefft" authorityYear="1871" box="[199,347,152,179]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="159" phylum="Nematoda" rank="genus">
<emphasis id="B968EA81DE0A1E315EF474EDFE89818B" bold="true" box="[199,347,152,179]" italics="true" pageId="8" pageNumber="159">Gnathosoma</emphasis>
</taxonomicName>
. Epistome and subcapitulum similar to female. Fixed digit of chelicera with seven teeth of various sizes, pilus dentilis setaceous. Movable digit of chelicera with one tooth, spermatodactyl short and apically curved dorsally, with blunt tip, free portion of spermatodactyl shorter than movable digit, fringed hyaline arthrodial process at base of movable digit (
<figureCitation id="13272A16DE0A1E315F877571FDDA8027" box="[436,520,260,287]" captionStart="FIGURES 1920" captionStartId="6.[151,264,1987,2012]" captionTargetBox="[164,1421,650,1948]" captionTargetId="figure@6.[151,1435,633,1962]" captionTargetPageId="6" captionText="FIGURES 1920. Cosmolaelaps ceylonensis sp. nov., male. 19, ventral idiosoma; 20, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687712" httpUri="https://zenodo.org/record/3687712/files/figure.png" pageId="8" pageNumber="159">Fig. 20</figureCitation>
). Palps similar to those of female.
</paragraph>
<paragraph id="8BA33693DE0A1E315EF4755CFED280B3" blockId="8.[151,1437,152,1403]" pageId="8" pageNumber="159">
<emphasis id="B968EA81DE0A1E315EF4755CFF2F807B" bold="true" box="[199,253,297,323]" italics="true" pageId="8" pageNumber="159">Legs</emphasis>
. Chaetotaxy as in female. Seta
<emphasis id="B968EA81DE0A1E315C6F755CFD56807B" box="[604,644,297,323]" italics="true" pageId="8" pageNumber="159">av1</emphasis>
on femur of leg II lanceolate (
<figureCitation id="13272A16DE0A1E315DE4755DFBF8807B" box="[983,1066,296,323]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="8" pageNumber="159">Fig. 24</figureCitation>
),
<emphasis id="B968EA81DE0A1E315A0E755CFB85807A" box="[1085,1111,297,322]" italics="true" pageId="8" pageNumber="159">pv</emphasis>
on tibia IV (
<figureCitation id="13272A16DE0A1E315AD6755DFAE4807B" box="[1253,1334,296,323]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="8" pageNumber="159">Fig. 23</figureCitation>
) and
<emphasis id="B968EA81DE0A1E315B47755CFA4E807B" box="[1396,1436,297,323]" italics="true" pageId="8" pageNumber="159">av2</emphasis>
on tarsus II, IV,
<emphasis id="B968EA81DE0A1E315F637538FEAB805F" box="[336,377,333,359]" italics="true" pageId="8" pageNumber="159">pv2</emphasis>
on tarsus III, IV spur-like (apically blunt) and tochanter II bearing a large dome-shaped spur (
<figureCitation id="13272A16DE0A1E315E937505FF2680B3" box="[160,244,368,395]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1945,1970]" captionTargetBox="[175,1409,187,1920]" captionTargetId="figure@7.[175,1413,181,1920]" captionTargetPageId="7" captionText="FIGURES 2124. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., male. 21, ventral idiosoma; 22, ventral strongly sclerotised median circular shape projection, 23, ventral view of leg IV; 24, ventral view of leg II." figureDoi="http://doi.org/10.5281/zenodo.3687714" httpUri="https://zenodo.org/record/3687714/files/figure.png" pageId="8" pageNumber="159">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0A1E315EF475E0FBB88096" box="[199,1130,404,431]" pageId="8" pageNumber="159" type="etymology">
<paragraph id="8BA33693DE0A1E315EF475E0FBB88096" blockId="8.[151,1437,152,1403]" box="[199,1130,404,431]" pageId="8" pageNumber="159">
<emphasis id="B968EA81DE0A1E315EF475E0FE9C8097" bold="true" box="[199,334,405,431]" pageId="8" pageNumber="159">Etymology.</emphasis>
Named after the
<typeStatus id="54A78831DE0A1E315C2375E0FD928097" box="[528,576,405,431]" pageId="8" pageNumber="159">type</typeStatus>
locality,
<collectingCountry id="F30B7603DE0A1E315C9475E1FCCA8096" box="[679,792,404,430]" name="Sri Lanka" pageId="8" pageNumber="159">Sri Lanka</collectingCountry>
, previously known as
<collectingCountry id="F30B7603DE0A1E315A2175E1FBB58096" box="[1042,1127,404,430]" name="Sri Lanka" pageId="8" pageNumber="159">Ceylon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066518DE0A1E315EF475CDFDAE8442" pageId="8" pageNumber="159" type="diagnosis">
<paragraph id="8BA33693DE0A1E315EF475CDFC4E82BA" blockId="8.[151,1437,152,1403]" pageId="8" pageNumber="159">
<emphasis id="B968EA81DE0A1E315EF475CDFE1F80EB" bold="true" box="[199,461,440,467]" pageId="8" pageNumber="159">Differential diagnosis.</emphasis>
The dorsal setae in
<taxonomicName id="4C1C4D10DE0A1E315C9A75CDFC1C80EA" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[681,974,440,466]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315C9A75CDFC1C80EA" box="[681,974,440,466]" italics="true" pageId="8" pageNumber="159">Cosmolaelaps ceylonensis</emphasis>
</taxonomicName>
are similar in shape to those of
<taxonomicName id="4C1C4D10DE0A1E315B1E75CCFEAD80CE" authority="Sheals, 1962" authorityName="Sheals" authorityYear="1962" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B968EA81DE0A1E315B1E75CCFA9280EB" box="[1325,1344,441,467]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315B7F75CCFF3480CE" italics="true" pageId="8" pageNumber="159">rectangularis</emphasis>
<bibRefCitation id="EF8D4B62DE0A1E315EDE75A9FEAD80CE" author="Sheals, J. G." box="[237,383,476,502]" pageId="8" pageNumber="159" pagination="83 - 110" refId="ref8673" refString="Sheals, J. G. (1962) Mesostigmata: Gamasida (Acari). Biologie de L'Amerique Australe, 1, 83 - 110." type="journal article" year="1962">Sheals, 1962</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315FBA75A8FD6880CF" authority="Tragardh, 1952" authorityName="Tragardh" authorityYear="1952" box="[393,698,476,503]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B968EA81DE0A1E315FBA75A8FE4E80CF" box="[393,412,477,503]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315F9975A8FDD480CE" box="[426,518,477,502]" italics="true" pageId="8" pageNumber="159">serratus</emphasis>
Trägårdh, 1952
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315CF675A8FC2780CF" authority="(Berlese, 1883)" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[709,1013,476,503]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="B968EA81DE0A1E315CF675A8FD0A80CF" box="[709,728,477,503]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315CD675A9FC9380CE" box="[741,833,476,502]" italics="true" pageId="8" pageNumber="159">claviger</emphasis>
(
<bibRefCitation id="EF8D4B62DE0A1E315D7C75A9FC3F80CF" author="Berlese, A." box="[847,1005,476,503]" pageId="8" pageNumber="159" refId="ref7131" refString="Berlese, A. (1883) Acari, Myriapoda et Scorpiones Hucusque in Italia Reperta, 4. 12 text pages + Plates 1 - 10. (Reprint by Junk, The Hague, 1979)." type="book" year="1883">Berlese, 1883</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315A3275A8FAC780CF" authority="(Karg, 1994)" baseAuthorityName="Karg" baseAuthorityYear="1994" box="[1025,1301,476,503]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B968EA81DE0A1E315A3275A8FBC680CF" box="[1025,1044,477,503]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315A1275A9FBAC80CE" box="[1057,1150,476,502]" italics="true" pageId="8" pageNumber="159">diversus</emphasis>
(
<bibRefCitation id="EF8D4B62DE0A1E315ABE75A8FADE80CF" author="Karg, W." box="[1165,1292,477,503]" pageId="8" pageNumber="159" pagination="179 - 216" refId="ref8226" refString="Karg, W. (1994) Raubmilben der cohors Gamasina Leach (Acarina, Parasitiformes) vom Galapagos-Archipel. Mitteilungen aus dem Zoologischen Museum in Berlin, 70, 179 - 216. http: // dx. doi. org / 10.1002 / mmnz. 19940700202" type="journal article" year="1994">Karg, 1994</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1C4D10DE0A1E315B7975A8FE278323" authority="(Bhattacharyya, 1968)" baseAuthorityName="Bhattacharyya" baseAuthorityYear="1968" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B968EA81DE0A1E315B7975A8FA8F80CF" box="[1354,1373,477,503]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315B5975A9FF218322" italics="true" pageId="8" pageNumber="159">bengalensis</emphasis>
(
<bibRefCitation id="EF8D4B62DE0A1E315F327675FE3E8323" author="Bhattacharyya, S. K." box="[257,492,512,539]" pageId="8" pageNumber="159" pagination="527 - 549" refId="ref7255" refString="Bhattacharyya, S. K. (1968) Studies on Indian mites (Acarina: Mesostigmata). 6. Six records and descriptions of nine new species. Acarologia, 10, 527 - 549." type="journal article" year="1968">Bhattacharyya, 1968</bibRefCitation>
)
</taxonomicName>
, described from
<collectingCountry id="F30B7603DE0A1E315C9E7675FCCD8322" box="[685,799,512,538]" name="Argentina" pageId="8" pageNumber="159">Argentina</collectingCountry>
,
<collectingCountry id="F30B7603DE0A1E315D1A7675FC7E8322" box="[809,940,512,538]" name="Saint Lucia" pageId="8" pageNumber="159">Saint Lucia</collectingCountry>
, Tahiti, Galápagos Islands and
<collectingCountry id="F30B7603DE0A1E315B367675FA938322" box="[1285,1345,512,538]" name="India" pageId="8" pageNumber="159">India</collectingCountry>
, respectively. It differs from all these species by the different shape of
<emphasis id="B968EA81DE0A1E315D6A7650FCBD8307" box="[857,879,549,575]" italics="true" pageId="8" pageNumber="159">j1</emphasis>
,
<emphasis id="B968EA81DE0A1E315D4E7650FC478307" box="[893,917,549,575]" italics="true" pageId="8" pageNumber="159">z1</emphasis>
and
<emphasis id="B968EA81DE0A1E315DFF7650FC3B8307" box="[972,1001,549,575]" italics="true" pageId="8" pageNumber="159">Z5</emphasis>
setae, foliate-dentate in
<taxonomicName id="4C1C4D10DE0A1E315AC87650FA4E8306" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[1275,1436,548,575]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315AC87650FADC8307" box="[1275,1294,549,575]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315B287651FA4E8306" box="[1307,1436,548,574]" italics="true" pageId="8" pageNumber="159">ceylonensis</emphasis>
</taxonomicName>
(
<figureCitation id="13272A16DE0A1E315E93763DFF32835B" box="[160,224,584,611]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="8" pageNumber="159">Fig. 1</figureCitation>
) (vs.
<emphasis id="B968EA81DE0A1E315F10763CFEEA835B" box="[291,312,585,611]" italics="true" pageId="8" pageNumber="159">j1</emphasis>
,
<emphasis id="B968EA81DE0A1E315F75763CFE8D835B" box="[326,351,585,611]" italics="true" pageId="8" pageNumber="159">z1</emphasis>
and
<emphasis id="B968EA81DE0A1E315FA5763CFE66835B" box="[406,436,585,611]" italics="true" pageId="8" pageNumber="159">Z5</emphasis>
setae obovate-spatulate or simple and smooth in the other species, never foliate-dentate). The sternal and genital setae (
<emphasis id="B968EA81DE0A1E315FD87618FDE983BF" box="[491,571,621,647]" italics="true" pageId="8" pageNumber="159">st1st3</emphasis>
and
<emphasis id="B968EA81DE0A1E315C407618FD4683BF" box="[627,660,621,647]" italics="true" pageId="8" pageNumber="159">st5</emphasis>
) are simple and the shields are smooth, without distinct reticulation, except for two diagonal lines in the posterior part of the genital shield which fuse with another horizontal line to form a semi-triangular figure in posterior part of the genital shield surface) in
<taxonomicName id="4C1C4D10DE0A1E315A2476C0FB6883F6" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[1047,1210,692,719]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315A2476C0FBF883F7" box="[1047,1066,693,719]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315A0A76C1FB6883F6" box="[1081,1210,692,718]" italics="true" pageId="8" pageNumber="159">ceylonensis</emphasis>
</taxonomicName>
(
<figureCitation id="13272A16DE0A1E315AFF76C1FAC183F7" box="[1228,1299,692,719]" captionStart="FIGURES 16" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[182,1399,191,1905]" captionTargetId="figure@3.[173,1415,179,1917]" captionTargetPageId="3" captionText="FIGURES 16. Cosmolaelaps ceylonensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, insemination structures, 4, subcapitulum; 5, epistome; 6, chelicera." figureDoi="http://doi.org/10.5281/zenodo.3687706" httpUri="https://zenodo.org/record/3687706/files/figure.png" pageId="8" pageNumber="159">Fig. 2</figureCitation>
). Both or at least one of the sternal or genital shields have distinct reticulation in
<taxonomicName id="4C1C4D10DE0A1E315D9776ACFB8983CA" authorityName="Sheals" authorityYear="1962" box="[932,1115,728,755]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="B968EA81DE0A1E315D9776ACFC6583CB" box="[932,951,729,755]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315DF676ADFB8983CA" box="[965,1115,728,754]" italics="true" pageId="8" pageNumber="159">rectangularis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315A5976ACFB3A83CA" authorityName="Tragardh" authorityYear="1952" box="[1130,1256,729,755]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B968EA81DE0A1E315A5976ACFBAF83CB" box="[1130,1149,729,755]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315ABF76ACFB3A83CA" box="[1164,1256,729,754]" italics="true" pageId="8" pageNumber="159">serratus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315AC576ACFAA683CA" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[1270,1396,728,755]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="B968EA81DE0A1E315AC576ACFADB83CB" box="[1270,1289,729,755]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315B2B76ADFAA683CA" box="[1304,1396,728,754]" italics="true" pageId="8" pageNumber="159">claviger</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D10DE0A1E315BB076ACFF26822E" baseAuthorityName="Karg" baseAuthorityYear="1994" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B968EA81DE0A1E315BB076ACFA4483CB" box="[1411,1430,729,755]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315EA47689FF26822E" box="[151,244,764,790]" italics="true" pageId="8" pageNumber="159">diversus</emphasis>
</taxonomicName>
; and setae
<emphasis id="B968EA81DE0A1E315F467688FE17822F" box="[373,453,765,791]" italics="true" pageId="8" pageNumber="159">st1st3</emphasis>
and
<emphasis id="B968EA81DE0A1E315FCE7688FDCF822F" box="[509,541,765,791]" italics="true" pageId="8" pageNumber="159">st5</emphasis>
are spatulate and with some barbs respectively in
<taxonomicName id="4C1C4D10DE0A1E315A687688FAD3822E" baseAuthorityName="Bhattacharyya" baseAuthorityYear="1968" box="[1115,1281,764,791]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B968EA81DE0A1E315A687688FBBC822F" box="[1115,1134,765,791]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315A4E7689FAD3822E" box="[1149,1281,764,790]" italics="true" pageId="8" pageNumber="159">bengalensis</emphasis>
</taxonomicName>
. Some dorsal setae on the tibia and genu of legs IIV and
<emphasis id="B968EA81DE0A1E315CB27755FD798202" box="[641,683,800,826]" italics="true" pageId="8" pageNumber="159">pd2</emphasis>
,
<emphasis id="B968EA81DE0A1E315C8A7755FD318202" box="[697,739,800,826]" italics="true" pageId="8" pageNumber="159">pd3</emphasis>
on tarsi IIIIV are feather-like in
<taxonomicName id="4C1C4D10DE0A1E315A6E7754FB2C8202" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[1117,1278,800,827]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315A6E7754FBA28203" box="[1117,1136,801,827]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315A4E7755FB2C8202" box="[1149,1278,800,826]" italics="true" pageId="8" pageNumber="159">ceylonensis</emphasis>
</taxonomicName>
(
<figureCitation id="13272A16DE0A1E315B3E7755FA5C8203" box="[1293,1422,800,827]" captionStart-0="FIGURES 714" captionStart-1="FIGURE 1518" captionStartId-0="4.[151,264,1945,1970]" captionStartId-1="5.[151,250,1910,1935]" captionTargetBox-0="[180,1407,182,1918]" captionTargetBox-1="[162,1425,192,1867]" captionTargetId-0="figure@4.[179,1409,181,1920]" captionTargetId-1="figure@5.[151,1436,181,1885]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 714. DIC micrographs of Cosmolaelaps ceylonensis sp. nov., female. 7, idiosoma in dorsal view, enlarged section not to scale; 8, idiosoma in ventral view; 9, general view of some dorsal setae; 10, subcapitulum; 11, epistome; 12, palp; 13, genu and tibia leg III; 14, femur, genu and tibia leg IV." captionText-1="FIGURE 1518. Cosmolaelaps ceylonensis sp. nov., female. 15, leg I (trochanter-tibia); 16, leg II; 17, leg III; 18, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.3687708" figureDoi-1="http://doi.org/10.5281/zenodo.3687710" httpUri-0="https://zenodo.org/record/3687708/files/figure.png" httpUri-1="https://zenodo.org/record/3687710/files/figure.png" pageId="8" pageNumber="159">Figs 1318</figureCitation>
); while the dorsal setae on tibia and genu of legs IIV in all the other species mentioned are sometimes spatulate or thickened but never feather-like, and
<emphasis id="B968EA81DE0A1E315C0A771DFDB182BA" box="[569,611,872,898]" italics="true" pageId="8" pageNumber="159">pd2</emphasis>
,
<emphasis id="B968EA81DE0A1E315C42771DFD4882BA" box="[625,666,872,898]" italics="true" pageId="8" pageNumber="159">pd3</emphasis>
on tarsi IIIIV simple.
</paragraph>
<paragraph id="8BA33693DE0A1E315EF477F8FDAE8442" blockId="8.[151,1437,152,1403]" pageId="8" pageNumber="159">
In the classification of
<bibRefCitation id="EF8D4B62DE0A1E315FFB77F8FD86829F" author="Karg, W." box="[456,596,909,935]" pageId="8" pageNumber="159" pagination="213 - 232" refId="ref8030" refString="Karg, W. (1981) Die Untergattung Cosmolaelaps Berlese, 1903 der Raubmilbengattung Hypoaspis Canestrini, 1884. Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 213 - 232." type="journal article" year="1981">Karg (1981)</bibRefCitation>
,
<taxonomicName id="4C1C4D10DE0A1E315C5377F8FCD0829E" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[608,770,908,935]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315C5377F8FDA1829F" box="[608,627,909,935]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315CB277F9FCD0829E" box="[641,770,908,934]" italics="true" pageId="8" pageNumber="159">ceylonensis</emphasis>
</taxonomicName>
falls into the
<taxonomicName id="4C1C4D10DE0A1E315DAF77F9FC2A829E" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[924,1016,908,934]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="B968EA81DE0A1E315DAF77F9FC2A829E" box="[924,1016,908,934]" italics="true" pageId="8" pageNumber="159">claviger</emphasis>
</taxonomicName>
species group of
<taxonomicName id="4C1C4D10DE0A1E315A8D77F8FF2182F3" authority="" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Cosmolaelaps">
<emphasis id="B968EA81DE0A1E315A8D77F8FAE6829E" box="[1214,1332,909,934]" italics="true" pageId="8" pageNumber="159">Hypoaspis</emphasis>
(
<emphasis id="B968EA81DE0A1E315B7677F8FF3A82F2" italics="true" pageId="8" pageNumber="159">Cosmolaelaps</emphasis>
)
</taxonomicName>
, because almost all dorsal shield setae oblong-mucronate, but
<taxonomicName id="4C1C4D10DE0A1E315D8A77C4FB8E82F2" authorityName="Joharchi &amp; Ermilov &amp; Khaustov" authorityYear="2020" box="[953,1116,944,971]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B968EA81DE0A1E315D8A77C4FC1E82F3" box="[953,972,945,971]" italics="true" pageId="8" pageNumber="159">C</emphasis>
.
<emphasis id="B968EA81DE0A1E315DE877C5FB8E82F2" box="[987,1116,944,970]" italics="true" pageId="8" pageNumber="159">ceylonensis</emphasis>
</taxonomicName>
is distinguished easily from all species in the
<taxonomicName id="4C1C4D10DE0A1E315F6877A1FE6582D6" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[347,439,980,1006]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="B968EA81DE0A1E315F6877A1FE6582D6" box="[347,439,980,1006]" italics="true" pageId="8" pageNumber="159">claviger</emphasis>
</taxonomicName>
species group by the following combination of unique characters: (1) dorsal shield with well reticulate ornamentation over whole surface and narrowing from level of setae
<emphasis id="B968EA81DE0A1E315A71778CFB89852B" box="[1090,1115,1017,1043]" italics="true" pageId="8" pageNumber="159">r3</emphasis>
, progressively tapering until rounded posteriorly, bearing 39 pairs of oblong-mucronate setae and almost all setae with 24 minute barbs and uniform in length and thickness except
<emphasis id="B968EA81DE0A1E315C1A7034FDED8563" box="[553,575,1089,1115]" italics="true" pageId="8" pageNumber="159">j1</emphasis>
,
<emphasis id="B968EA81DE0A1E315C7F7034FDB78563" box="[588,613,1089,1115]" italics="true" pageId="8" pageNumber="159">z1</emphasis>
and
<emphasis id="B968EA81DE0A1E315CA87034FD6A8563" box="[667,696,1089,1115]" italics="true" pageId="8" pageNumber="159">Z5</emphasis>
foliate-dentate and shorter; (2) both sternal and genital shields obviously smooth and without distinct reticulation (except two diagonal lines in posterior part of genital shield which confluence with another horizontal line and forming semi-triangular in posterior part of genital shield surface) and sternal shield bearing two pairs of large slit shape pore-like structures and
<emphasis id="B968EA81DE0A1E315DD270D8FBD085FF" box="[993,1026,1197,1223]" italics="true" pageId="8" pageNumber="159">st1</emphasis>
length enough to well reach base of
<emphasis id="B968EA81DE0A1E315EA470A4FF6585D3" box="[151,183,1233,1259]" italics="true" pageId="8" pageNumber="159">st2</emphasis>
, (3) fixed digit of chelicera with 1011 various size of teeth; (4) some dorsal setae on tibia and genu of legs IIV (antero-dorsals and postero-dorsals setae on tibia and genu III and postero-dorsals setae on tibia and genu IIIIV) and
<emphasis id="B968EA81DE0A1E315EF5716DFF22840A" box="[198,240,1304,1330]" italics="true" pageId="8" pageNumber="159">pd2</emphasis>
,
<emphasis id="B968EA81DE0A1E315ECE716DFEF5840A" box="[253,295,1304,1330]" italics="true" pageId="8" pageNumber="159">pd3</emphasis>
on tarsi IIIIV feather-like; (5) Male with holoventral shield; shield with a strongly sclerotised median circular shape projection and spermatodactyl short and apically curved dorsally; (6)
<emphasis id="B968EA81DE0A1E315A767149FBB3846E" box="[1093,1121,1340,1366]" italics="true" pageId="8" pageNumber="159">ad</emphasis>
on tibia IV and
<emphasis id="B968EA81DE0A1E315B247149FA90846E" box="[1303,1346,1340,1366]" italics="true" pageId="8" pageNumber="159">ad2</emphasis>
,
<emphasis id="B968EA81DE0A1E315B7C7149FAAB846E" box="[1359,1401,1340,1366]" italics="true" pageId="8" pageNumber="159">pd2</emphasis>
on tarsus IV spur-like (apically blunt) in male.
</paragraph>
</subSubSection>
</treatment>
</document>