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<document id="5D8FB3045F5CA5C9ECBCCCC3777D1472" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A6730662FDFAF902FE93E160" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba rugaticollis Gao &amp; Coca-Abia 2021, comb. nov." docType="treatment" docVersion="8" lastPageNumber="73" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="71" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8A4829ECFEAA9A58B3547D9B78A28E2F">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="292708D18130F7002CC56BDB2B1432F1">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="604B2712C482634202FC4D6AC6B9BABA">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="18B4CA3DED8FCB9856F04E3919CDA88F">Coca-Abia, María Milagro</mods:namePart>
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<mods:date id="5B97F003FB5CBFDF518ED89E60DEB750">2021</mods:date>
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<subSubSection id="C30685DEA673066CFDFAF902FBD1E2F6" box="[513,1075,1707,1735]" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="8BA3D655A673066CFDFAF902FBD1E2F6" blockId="70.[513,1075,1707,1769]" box="[513,1075,1707,1735]" pageId="70" pageNumber="71">
<heading id="D0EB6139A673066CFDFAF902FBD1E2F6" box="[513,1075,1707,1735]" centered="true" fontSize="11" level="2" pageId="70" pageNumber="71" reason="2">
<taxonomicName id="4C1CADD6A673066CFDFAF902FC4BE2F7" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[513,937,1707,1734]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rugaticollis" status="comb. nov.">
<emphasis id="B9680A47A673066CFDFAF902FD1EE2F7" bold="true" box="[513,764,1707,1734]" italics="true" pageId="70" pageNumber="71">Miridiba rugaticollis</emphasis>
(Moser, 1913)
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA673066CFC4AF905FBD1E2F6" box="[945,1075,1708,1735]" pageId="70" pageNumber="71" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C30685DEA673066CFD39F966FC93E2D8" box="[706,881,1743,1769]" pageId="70" pageNumber="71" type="description">
<paragraph id="8BA3D655A673066CFD39F966FC93E2D8" blockId="70.[513,1075,1707,1769]" box="[706,881,1743,1769]" pageId="70" pageNumber="71">
<figureCitation id="1327CAD0A673066CFD39F966FCA5E2D8" box="[706,839,1743,1769]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="70" pageNumber="71">Figs 18–19</figureCitation>
,
<figureCitation id="1327CAD0A673066CFCAFF966FC93E2D8" box="[852,881,1743,1769]" captionStart="Figs 90–93. 90" captionStartId="71.[187,239,1888,1914]" captionTargetBox="[277,1309,700,1826]" captionTargetId="figure-140@71.[264,1324,685,1849]" captionTargetPageId="71" captionText="Figs 90–93. 90. Miridiba rugaticollis (Moser, 1913), lectotype, ♂ (MFNB). Habitus in dorsal view. 91–93. M. nigrescens (Moser, 1916), lectotype, ♂ (MFNB). 91. Habitus in dorsal view. 92. Parameres in dorsal (A) and lateral (B) views. 93. Endophallus and temones in lateral view. Scale bars: 90–91 = 5.0 mm; 92–93 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770407" httpUri="https://zenodo.org/record/4770407/files/figure.png" pageId="70" pageNumber="71">90</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA673066CFF46F8B8FC53E35C" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="8BA3D655A673066CFF46F8B8FC3DE31D" blockId="70.[189,991,1808,1836]" box="[189,991,1808,1836]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="AB0CF17BA673066CFF46F8B8FC3DE31D" box="[189,991,1808,1836]" pageId="70" pageNumber="71">
<treatmentCitation id="0ABDF044A673066CFF46F8B8FD4AE31A" author="Moser J." box="[189,680,1808,1835]" page="53" pageId="70" pageNumber="71" year="1913">
<taxonomicName id="4C1CADD6A673066CFF46F8B8FD4AE31A" authority="Moser, 1913 a: 53" authorityName="Moser" authorityPageNumber="53" authorityYear="1913" box="[189,680,1808,1835]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A673066CFF46F8B8FE36E31A" box="[189,468,1809,1835]" italics="true" pageId="70" pageNumber="71">Holotrichia rugaticollis</emphasis>
<bibRefCitation id="EF8DABA4A673066CFE27F8B8FD4AE31A" author="Moser J." box="[476,680,1808,1835]" pageId="70" pageNumber="71" pagination="49 - 70" refId="ref45347" refString="Moser J. 1913 a. Neue Indische Melolonthiden. (Col.). Deutsche Entomologische Zeitschrift 1913: 49 - 70." type="journal article" year="1913">Moser, 1913a: 53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A673066CFD43F8B8FD0EE31A" box="[696,748,1809,1835]" pageId="70" pageNumber="71">type</typeStatus>
loc.: Kanara,
<collectingCountry id="F30B96C5A673066CFC69F8B8FC31E31D" box="[914,979,1809,1836]" name="India" pageId="70" pageNumber="71">India</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A673066CFF46F8FBFC53E35C" blockId="70.[189,945,1874,1901]" box="[189,945,1874,1901]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="AB0CF17BA673066CFF46F8FBFC53E35C" box="[189,945,1874,1901]" pageId="70" pageNumber="71">
<taxonomicName id="4C1CADD6A673066CFF46F8FBFE36E35D" authorityName="Moser" authorityYear="1913" box="[189,468,1874,1900]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A673066CFF46F8FBFE36E35D" box="[189,468,1874,1900]" italics="true" pageId="70" pageNumber="71">Holotrichia rugaticollis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A673066CFE09F8FAFD00E35D" author="Frey G." box="[498,738,1874,1901]" page="220" pageId="70" pageNumber="71" year="1971">
<bibRefCitation id="EF8DABA4A673066CFE09F8FAFD00E35D" author="Frey G." box="[498,738,1874,1901]" pageId="70" pageNumber="71" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey 1971: 220–221</bibRefCitation>
</treatmentCitation>
, fig. 29a (in key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6720662FF46FEA1FBD1E651" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="8BA3D655A672066DFF46FEA1FED4E513" blockId="71.[189,310,264,290]" box="[189,310,264,290]" pageId="71" pageNumber="72">
<heading id="D0EB6139A672066DFF46FEA1FED4E513" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="71" pageNumber="72" reason="3">
<emphasis id="B9680A47A672066DFF46FEA1FED4E513" bold="true" box="[189,310,264,290]" pageId="71" pageNumber="72">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6720662FF46FE9EFBD1E651" blockId="71.[187,1400,311,619]" lastBlockId="72.[188,1399,265,608]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
External morphology of adult (
<figureCitation id="1327CAD0A672066DFDE6FE9EFD91E560" box="[541,627,311,337]" captionStart="Figs 90–93. 90" captionStartId="71.[187,239,1888,1914]" captionTargetBox="[277,1309,700,1826]" captionTargetId="figure-140@71.[264,1324,685,1849]" captionTargetPageId="71" captionText="Figs 90–93. 90. Miridiba rugaticollis (Moser, 1913), lectotype, ♂ (MFNB). Habitus in dorsal view. 91–93. M. nigrescens (Moser, 1916), lectotype, ♂ (MFNB). 91. Habitus in dorsal view. 92. Parameres in dorsal (A) and lateral (B) views. 93. Endophallus and temones in lateral view. Scale bars: 90–91 = 5.0 mm; 92–93 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770407" httpUri="https://zenodo.org/record/4770407/files/figure.png" pageId="71" pageNumber="72">Fig. 90</figureCitation>
). Body size
<quantity id="4CE47BB0A672066DFD00FE9EFC87E563" box="[763,869,311,338]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="71" pageNumber="72" unit="mm" value="14.7">14.7 mm</quantity>
. Dorsal surface shiny, glabrous. Male antennal club longer than funiculus and scape. Clypeus longer than frons, almost flat, arcuate laterally, anterior edge emarginate at middle; surface with conspicuous and deep punctures. Frons with punctures similar to those of clypeus, wrinkled appearance; frontal carina blunt. Pronotal surface with strong and deep punctures, wrinkled appearance; anterior margin glabrous, widely flanged; posterior margin glabrous; lateral margins widely serrated with short setae, moderately elevated at anterior half; anterior angles almost right, posterior angles obtuse and marked. Prosternal process slender, apex spade-shaped. Scutellum glabrous, with punctures uniformly distributed. Elytral surface with punctures evenly distributed and softer than those of pronotum; glabrous, at most with short setae in each puncture hardly visible; epipleuron glabrous, with a few short setae at basal part. Foretibia with a dorsal carina moderately developed. Meso- and metatibia with transverse carina complete, inner margin of dorsal surface with a few spines. Prepygidium with punctures regularly distributed. Pygidium with punctures irregularly distributed, glabrous; apex with sparse and short pubescence. Ventrites 1 and 2 with conspicuous short pubescence reclined and regularly distributed. Ventrites 3 and 4 glabrous. Ventrite 5 moderately depressed at posterior half, with sparse long pubescence irregularly distributed. Ventrite 6 moderately bulging at anterior margin, with strong and deep punctures, wrinkled appearance; long pubescence on posterior margin. Male genitalia fits into morphotype VII “
<emphasis id="B9680A47A67D0662FE68FE56FDFDE628" box="[403,543,511,537]" italics="true" pageId="72" pageNumber="73">Rugaticollis</emphasis>
”. Parameres (
<figureCitation id="1327CAD0A67D0662FD30FE56FCC0E628" box="[715,802,511,537]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="72" pageNumber="73">Fig. 18</figureCitation>
) with notch of dorsal branch well developed and circular-shaped. Endophallus (
<figureCitation id="1327CAD0A67D0662FDE4FD8BFD91E60C" box="[543,627,546,573]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="72" pageNumber="73">Fig. 19</figureCitation>
) with an area of soft sensillae at distal half. Apophysis of temones (
<figureCitation id="1327CAD0A67D0662FF3DFDEFFEF8E651" box="[198,282,582,608]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="72" pageNumber="73">Fig. 19</figureCitation>
) separated from each other at both ends. Female genitalia unknown.
</paragraph>
</subSubSection>
<caption id="DF6386DDA672066DFF40F8C9FE10E3D5" ID-DOI="http://doi.org/10.5281/zenodo.4770407" ID-Zenodo-Dep="4770407" httpUri="https://zenodo.org/record/4770407/files/figure.png" pageId="71" pageNumber="72" startId="71.[187,239,1888,1914]" targetBox="[277,1309,700,1826]" targetPageId="71">
<paragraph id="8BA3D655A672066DFF40F8C9FE10E3D5" blockId="71.[187,1399,1888,2021]" pageId="71" pageNumber="72">
<emphasis id="B9680A47A672066DFF40F8C9FE90E34B" bold="true" box="[187,370,1888,1914]" pageId="71" pageNumber="72">Figs 90–93. 90</emphasis>
.
<taxonomicName id="4C1CADD6A672066DFE7DF8C9FCDAE34B" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[390,824,1888,1914]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A672066DFE7DF8C9FD62E34B" box="[390,640,1888,1914]" italics="true" pageId="71" pageNumber="72">Miridiba rugaticollis</emphasis>
(Moser, 1913)
</taxonomicName>
, lectotype, ♂ (MFNB). Habitus in dorsal view.
<emphasis id="B9680A47A672066DFF46F82AFEE4E3AC" bold="true" box="[189,262,1923,1949]" pageId="71" pageNumber="72">91–93</emphasis>
.
<taxonomicName id="4C1CADD6A672066DFEEDF82DFD8FE3AF" authority="(Moser, 1916)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1916" box="[278,621,1923,1950]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="B9680A47A672066DFEEDF82DFE58E3AC" box="[278,442,1924,1949]" italics="true" pageId="71" pageNumber="72">M. nigrescens</emphasis>
(
<bibRefCitation id="EF8DABA4A672066DFE30F82DFD86E3AF" author="Moser J." box="[459,612,1923,1950]" pageId="71" pageNumber="72" pagination="129 - 190" refId="ref45401" refString="Moser J. 1916. Beitrag zur Kenntnis der Melolonthiden. (Col.) V. Deutsche Entomologische Zeitschrift 1916: 129 - 190." type="journal article" year="1916">Moser, 1916</bibRefCitation>
)
</taxonomicName>
, lectotype, ♂ (MFNB).
<emphasis id="B9680A47A672066DFC72F82AFC44E3AC" bold="true" box="[905,934,1923,1949]" pageId="71" pageNumber="72">91</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A672066DFB2BF82AFB0EE3AC" bold="true" box="[1232,1260,1923,1949]" pageId="71" pageNumber="72">92</emphasis>
. Parameres in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A672066DFD90F80FFD65E3F1" bold="true" box="[619,647,1958,1984]" pageId="71" pageNumber="72">93</emphasis>
. Endophallus and temones in lateral view. Scale bars: 90–91 = 5.0 mm; 92–93 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA67D0662FF46FD2EFE46E7D4" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8BA3D655A67D0662FF46FD2EFE48E690" blockId="72.[189,426,647,673]" box="[189,426,647,673]" pageId="72" pageNumber="73">
<heading id="D0EB6139A67D0662FF46FD2EFE48E690" bold="true" box="[189,426,647,673]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B9680A47A67D0662FF46FD2EFE48E690" bold="true" box="[189,426,647,673]" pageId="72" pageNumber="73">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67D0662FF11FD1FFDDAE6E0" blockId="72.[189,1398,694,826]" box="[234,568,694,721]" pageId="72" pageNumber="73">
<emphasis id="B9680A47A67D0662FF11FD1FFE84E6E1" bold="true" box="[234,358,694,720]" pageId="72" pageNumber="73">
<typeStatus id="54A768F7A67D0662FF11FD1FFE84E6E1" box="[234,358,694,720]" pageId="72" pageNumber="73" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA3D655A67D0662FF46FD70FC45E70B" blockId="72.[189,1398,694,826]" pageId="72" pageNumber="73">
<materialsCitation id="3B74DC08A67D0662FF46FD70FC45E70B" ID-GBIF-Occurrence="3114121400" collectionCode="MFNB" country="India" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="INDIA" pageId="72" pageNumber="73" specimenCode="MNHUB 2016" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F30B96C5A67D0662FF46FD70FEECE6C2" box="[189,270,729,755]" name="India" pageId="72" pageNumber="73">INDIA</collectingCountry>
•
<specimenCount id="9D1A1DDCA67D0662FEC0FD70FEB6E6C2" box="[315,340,729,755]" pageId="72" pageNumber="73" type="male">♂</specimenCount>
; “Kanara”; “
<emphasis id="B9680A47A67D0662FE04FD70FD6FE6C2" box="[511,653,729,755]" italics="true" pageId="72" pageNumber="73">Rugaticollis</emphasis>
Brensk”; “
<taxonomicName id="4C1CADD6A67D0662FCDEFD70FBA4E6C2" authorityName="Moser" authorityYear="1913" box="[805,1094,729,755]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FCDEFD70FBA4E6C2" box="[805,1094,729,755]" italics="true" pageId="72" pageNumber="73">Holotrichia rugaticollis</emphasis>
</taxonomicName>
Type Mos”; “Syntypus
<taxonomicName id="4C1CADD6A67D0662FF46FD55FD9EE727" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[189,636,764,791]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FF46FD55FE3BE727" box="[189,473,764,790]" italics="true" pageId="72" pageNumber="73">Holotrichia rugaticollis</emphasis>
Moser, 1913
</taxonomicName>
labelled by
<specimenCode id="DBBA7E2EA67D0662FCE2FD54FC31E727" box="[793,979,764,791]" pageId="72" pageNumber="73">MNHUB 2016</specimenCode>
”; “Lectotype
<taxonomicName id="4C1CADD6A67D0662FB80FD55FE81E708" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FB80FD55FA94E727" box="[1147,1398,764,790]" italics="true" pageId="72" pageNumber="73">Miridiba rugaticollis</emphasis>
(Moser, 1913)
</taxonomicName>
<determinerName id="7E7F3A91A67D0662FE90FCB6FDF5E708" box="[363,535,799,825]" pageId="72" pageNumber="73">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A67D0662FDC7FCB6FD59E708" box="[572,699,799,825]" pageId="72" pageNumber="73">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A67D0662FCB7FC89FC41E70B" box="[844,931,800,826]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="72" pageNumber="73">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A67D0662FF11FCC8FDB5E74D" blockId="72.[187,1399,865,997]" box="[234,599,865,892]" pageId="72" pageNumber="73">
<emphasis id="B9680A47A67D0662FF11FCC8FE75E74A" bold="true" box="[234,407,865,891]" pageId="72" pageNumber="73">
<typeStatus id="54A768F7A67D0662FF11FCC8FE75E74A" box="[234,407,865,891]" pageId="72" pageNumber="73" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(here indicated)
</paragraph>
<paragraph id="8BA3D655A67D0662FF46FC2DFE46E7D4" blockId="72.[187,1399,865,997]" pageId="72" pageNumber="73">
<materialsCitation id="3B74DC08A67D0662FF46FC2DFE42E7D4" ID-GBIF-Occurrence="3114121375" collectionCode="MFNB" country="India" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="INDIA" pageId="72" pageNumber="73" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="F30B96C5A67D0662FF46FC2DFEECE7AF" box="[189,270,900,926]" name="India" pageId="72" pageNumber="73">INDIA</collectingCountry>
•
<specimenCount id="9D1A1DDCA67D0662FEDFFC2DFEB3E7AE" box="[292,337,900,927]" pageId="72" pageNumber="73" type="male">1 ♂</specimenCount>
; “Kanara”; “
<emphasis id="B9680A47A67D0662FE1EFC2DFD91E7AF" box="[485,627,900,926]" italics="true" pageId="72" pageNumber="73">Rugaticollis</emphasis>
” [in red]; “Syntypus
<taxonomicName id="4C1CADD6A67D0662FC9DFC2DFAF6E7AF" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[870,1300,900,927]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FC9DFC2DFB99E7AF" box="[870,1147,900,926]" italics="true" pageId="72" pageNumber="73">Holotrichia rugaticollis</emphasis>
Moser, 1913
</taxonomicName>
labelled by MNHUB 2016”; “Paralectotype
<taxonomicName id="4C1CADD6A67D0662FD97FC0EFBF7E7F0" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[620,1045,935,961]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FD97FC0EFC87E7F0" box="[620,869,935,961]" italics="true" pageId="72" pageNumber="73">Miridiba rugaticollis</emphasis>
(Moser, 1913)
</taxonomicName>
<determinerName id="7E7F3A91A67D0662FBE4FC0EFB2CE7F0" box="[1055,1230,935,961]" pageId="72" pageNumber="73">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A67D0662FB0CFC0EFA94E7F0" box="[1271,1398,935,961]" pageId="72" pageNumber="73">Coca-Abia</determinerName>
det. 2017”;
<collectionCode id="ED0D4E90A67D0662FEB2FC62FE42E7D4" box="[329,416,971,997]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="72" pageNumber="73">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67D0662FF46FBA5FE71E0D0" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8BA3D655A67D0662FF46FBA5FECDE017" blockId="72.[189,303,1036,1062]" box="[189,303,1036,1062]" pageId="72" pageNumber="73">
<heading id="D0EB6139A67D0662FF46FBA5FECDE017" bold="true" box="[189,303,1036,1062]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B9680A47A67D0662FF46FBA5FECDE017" bold="true" box="[189,303,1036,1062]" pageId="72" pageNumber="73">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67D0662FF46FB92FE71E0D0" blockId="72.[188,1400,1082,1249]" pageId="72" pageNumber="73">
<bibRefCitation id="EF8DABA4A67D0662FF46FB92FE8FE065" author="Moser J." box="[189,365,1082,1109]" pageId="72" pageNumber="73" pagination="49 - 70" refId="ref45347" refString="Moser J. 1913 a. Neue Indische Melolonthiden. (Col.). Deutsche Entomologische Zeitschrift 1913: 49 - 70." type="journal article" year="1913">Moser (1913a)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A67D0662FE0BFB93FCEBE065" authorityName="Moser" authorityYear="1913" box="[496,777,1082,1108]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FE0BFB93FCEBE065" box="[496,777,1082,1108]" italics="true" pageId="72" pageNumber="73">Holotrichia rugaticollis</emphasis>
</taxonomicName>
from a series of specimens from
<collectingCountry id="F30B96C5A67D0662FB60FB93FB1CE065" box="[1179,1278,1082,1108]" name="India" pageId="72" pageNumber="73">Bombay</collectingCountry>
(Kanara). We have studied
<specimenCount id="9D1A1DDCA67D0662FE76FBF7FE1EE046" box="[397,508,1117,1144]" pageId="72" pageNumber="73" type="male">two male</specimenCount>
type specimens (MFNB) (one of them heavily damaged), which show the external features that characterize the genus
<taxonomicName id="4C1CADD6A67D0662FD28FB28FCD9E0AA" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[723,827,1153,1179]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67D0662FD28FB28FCD9E0AA" box="[723,827,1153,1179]" italics="true" pageId="72" pageNumber="73">Miridiba</emphasis>
</taxonomicName>
and the morphotype VII “
<emphasis id="B9680A47A67D0662FB82FB28FAE5E0AA" box="[1145,1287,1153,1179]" italics="true" pageId="72" pageNumber="73">Rugaticollis</emphasis>
”. Hence, we transfer
<taxonomicName id="4C1CADD6A67D0662FEB8FB0CFE0FE08F" authorityName="Moser" authorityYear="1913" box="[323,493,1188,1214]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A67D0662FEB8FB0CFE0FE08F" box="[323,493,1188,1214]" italics="true" pageId="72" pageNumber="73">H. rugaticollis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A67D0662FDEBFB0DFD9AE08F" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[528,632,1188,1214]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67D0662FDEBFB0DFD9AE08F" box="[528,632,1188,1214]" italics="true" pageId="72" pageNumber="73">Miridiba</emphasis>
</taxonomicName>
and to this genital morphotype. Moreover, we designate
<typeStatus id="54A768F7A67D0662FAF0FB0DFA95E08F" box="[1291,1399,1188,1214]" pageId="72" pageNumber="73" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54A768F7A67D0662FF16FB6EFE6DE0D0" box="[237,399,1223,1249]" pageId="72" pageNumber="73" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67D0662FF46FAA1FE93E160" box="[189,369,1288,1361]" pageId="72" pageNumber="73" type="distribution">
<paragraph id="8BA3D655A67D0662FF46FAA1FEB4E113" blockId="72.[189,342,1288,1314]" box="[189,342,1288,1314]" pageId="72" pageNumber="73">
<heading id="D0EB6139A67D0662FF46FAA1FEB4E113" bold="true" box="[189,342,1288,1314]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B9680A47A67D0662FF46FAA1FEB4E113" bold="true" box="[189,342,1288,1314]" pageId="72" pageNumber="73">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67D0662FF46FA9EFE93E160" blockId="72.[189,369,1335,1361]" box="[189,369,1335,1361]" pageId="72" pageNumber="73">
<collectingCountry id="F30B96C5A67D0662FF46FA9EFF1BE160" box="[189,249,1335,1361]" name="India" pageId="72" pageNumber="73">India</collectingCountry>
(Kanara).
</paragraph>
</subSubSection>
</treatment>
</document>
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