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<document id="CE11F6D7296700DFDC6C83144E0C2902" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A61E0607FDD7FEA1FD1CE5CD" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba tuberculipennis M. Coca-Abia" docType="treatment" docVersion="11" lastPageNumber="46" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="44" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DF96CE96617CD005AC56A3DD79D7A476">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="02A00E04014805C06B3C2367027FAE80">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="A62C4F4DBBD1C6A55658F589B87F5A8A">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="816D396BAA6D18511C217D2926CE0768">Coca-Abia, María Milagro</mods:namePart>
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<treatment id="03B56743A61E0607FDD7FEA1FD1CE5CD" ID-DOI="http://doi.org/10.5281/zenodo.4773259" ID-GBIF-Taxon="182237232" ID-Zenodo-Dep="4773259" LSID="urn:lsid:plazi:treatment:03B56743A61E0607FDD7FEA1FD1CE5CD" httpUri="http://treatment.plazi.org/id/03B56743A61E0607FDD7FEA1FD1CE5CD" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection id="C30685DEA61E0601FDD7FEA1FBE5E512" box="[556,1031,264,291]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8BA3D655A61E0601FDD7FEA1FBE5E512" blockId="43.[556,1031,264,326]" box="[556,1031,264,291]" pageId="43" pageNumber="44">
<heading id="D0EB6139A61E0601FDD7FEA1FBE5E512" box="[556,1031,264,291]" centered="true" fontSize="11" level="2" pageId="43" pageNumber="44" reason="2">
<taxonomicName id="4C1CADD6A61E0601FDD7FEA1FBE5E512" authority="(Moser, 1913)" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[556,1031,264,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A61E0601FDD7FEA1FCB8E512" bold="true" box="[556,858,264,291]" italics="true" pageId="43" pageNumber="44">Miridiba tuberculipennis</emphasis>
(Moser, 1913)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA61E0601FD23FE85FCB9E577" box="[728,859,300,326]" pageId="43" pageNumber="44" type="description">
<paragraph id="8BA3D655A61E0601FD23FE85FCB9E577" blockId="43.[556,1031,264,326]" box="[728,859,300,326]" pageId="43" pageNumber="44">
<figureCitation id="1327CAD0A61E0601FD23FE85FCB9E577" box="[728,859,300,326]" captionStart="Figs 6062" captionStartId="43.[190,242,1920,1946]" captionTargetBox="[287,1310,1023,1877]" captionTargetId="figure-196@43.[264,1323,1016,1881]" captionTargetPageId="43" captionText="Figs 6062. Miridiba tuberculipennis (Moser, 1913). 6061. Paralectotype, ♂ (MFNB). 62. Non-type, ♀ (BMNH). 60. Habitus in dorsal view. 61. Male genitalia: parameres and endophallus in lateral view. 62. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="43" pageNumber="44">Figs 6062</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA61E0601FF46FEC4FE12E68E" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8BA3D655A61E0601FF46FEC4FC02E5B9" blockId="43.[189,992,365,427]" box="[189,992,365,392]" pageId="43" pageNumber="44">
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<treatmentCitation id="0ABDF044A61E0601FF46FEC4FD35E5B9" author="Moser J." box="[189,727,365,392]" page="55" pageId="43" pageNumber="44" year="1913">
<taxonomicName id="4C1CADD6A61E0601FF46FEC4FD35E5B9" authority="Moser, 1913 a: 55" authorityName="Moser" authorityPageNumber="55" authorityYear="1913" box="[189,727,365,392]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A61E0601FF46FEC4FDE0E5B6" box="[189,514,365,391]" italics="true" pageId="43" pageNumber="44">Holotrichia tuberculipennis</emphasis>
<bibRefCitation id="EF8DABA4A61E0601FDF1FEC7FD35E5B9" author="Moser J." box="[522,727,365,392]" pageId="43" pageNumber="44" pagination="49 - 70" refId="ref45347" refString="Moser J. 1913 a. Neue Indische Melolonthiden. (Col.). Deutsche Entomologische Zeitschrift 1913: 49 - 70." type="journal article" year="1913">Moser, 1913a: 55</bibRefCitation>
</taxonomicName>
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(
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loc.:
<collectingCountry id="F30B96C5A61E0601FCA7FEC7FC36E5B9" box="[860,980,366,392]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A61E0601FF46FE39FCE1E59A" blockId="43.[189,992,365,427]" box="[189,771,400,427]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB0CF17BA61E0601FF46FE39FCE1E59A" box="[189,771,400,427]" pageId="43" pageNumber="44">
<treatmentCitation id="0ABDF044A61E0601FF46FE39FD7AE59A" author="Frey G." box="[189,664,400,427]" page="245" pageId="43" pageNumber="44" year="1970">
<taxonomicName id="4C1CADD6A61E0601FF46FE39FD7AE59A" authority="Frey, 1970: 245" authorityName="Frey" authorityPageNumber="245" authorityYear="1970" box="[189,664,400,427]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A61E0601FF46FE39FE3AE59B" box="[189,472,400,426]" italics="true" pageId="43" pageNumber="44">Neodontocnema ardoini</emphasis>
<bibRefCitation id="EF8DABA4A61E0601FE1BFE38FD7AE59A" author="Frey G." box="[480,664,400,427]" pageId="43" pageNumber="44" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970: 245</bibRefCitation>
</taxonomicName>
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(in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A61E0601FF46FE7BFC27E5DC" blockId="43.[189,1398,466,703]" box="[189,965,466,493]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB0CF17BA61E0601FF46FE7BFC27E5DC" box="[189,965,466,493]" pageId="43" pageNumber="44">
<taxonomicName id="4C1CADD6A61E0601FF46FE7BFDDAE5DD" box="[189,568,466,492]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A61E0601FF46FE7BFDDAE5DD" box="[189,568,466,492]" italics="true" pageId="43" pageNumber="44">Neodontocnema tuberculipennis</emphasis>
</taxonomicName>
<treatmentCitation id="0ABDF044A61E0601FDAFFE7BFCEDE5DC" author="Arrow G. J." box="[596,783,466,493]" page="51" pageId="43" pageNumber="44" year="1948">
<bibRefCitation id="EF8DABA4A61E0601FDAFFE7BFCEDE5DC" author="Arrow G. J." box="[596,783,466,493]" pageId="43" pageNumber="44" pagination="49 - 54" refId="ref43061" refString="Arrow G. J. 1948. Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. The Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 17: 49 - 54. https: // doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow 1948: 51</bibRefCitation>
</treatmentCitation>
(combination).
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</paragraph>
<paragraph id="8BA3D655A61E0601FF46FE5CFC5DE621" blockId="43.[189,1398,466,703]" box="[189,959,501,528]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB0CF17BA61E0601FF46FE5CFC5DE621" box="[189,959,501,528]" pageId="43" pageNumber="44">
<emphasis id="B9680A47A61E0601FF46FE5CFDCFE63E" box="[189,557,501,527]" italics="true" pageId="43" pageNumber="44">Neodontcnema tuberculiventris</emphasis>
<treatmentCitation id="0ABDF044A61E0601FDB1FE5CFC5DE621" author="Frey" box="[586,959,501,528]" page="88" pageId="43" pageNumber="44" year="1970">
<bibRefCitation id="EF8DABA4A61E0601FDB1FE5CFD1CE621" author="Frey G." box="[586,766,501,528]" pageId="43" pageNumber="44" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey 1970: 245</bibRefCitation>
(lapsus calami).
</treatmentCitation>
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</paragraph>
<paragraph id="8BA3D655A61E0601FF46FDB1FBECE648" blockId="43.[189,1398,466,703]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB0CF17BA61E0601FF46FDB1FBECE648" pageId="43" pageNumber="44">
<taxonomicName id="4C1CADD6A61E0601FF46FDB1FE03E603" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[189,481,536,562]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A61E0601FF46FDB1FE03E603" box="[189,481,536,562]" italics="true" pageId="43" pageNumber="44">Miridiba tuberculipennis</emphasis>
</taxonomicName>
<treatmentCitation id="0ABDF044A61E0601FE06FDB0FD29E603" author="Nomura S." box="[509,715,536,563]" page="88" pageId="43" pageNumber="44" year="1977">
<bibRefCitation id="EF8DABA4A61E0601FE06FDB0FD29E603" author="Nomura S." box="[509,715,536,563]" pageId="43" pageNumber="44" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977: 88</bibRefCitation>
</treatmentCitation>
(combination). —
<treatmentCitation id="0ABDF044A61E0601FC51FDB1FBB4E603" author="Itoh T." box="[938,1110,536,563]" page="199" pageId="43" pageNumber="44" year="1995">
<bibRefCitation id="EF8DABA4A61E0601FC51FDB1FBB4E603" author="Itoh T." box="[938,1110,536,563]" pageId="43" pageNumber="44" pagination="195 - 204" refId="ref44436" refString="Itoh T. 1995. Melolonthidae (Coleoptera) from Thailand II. Elytra, Tokyo 23 (2): 195 - 204." type="journal article" year="1995">Itoh 1995: 199</bibRefCitation>
</treatmentCitation>
(combination). —
<treatmentCitation id="0ABDF044A61E0601FACEFDB1FEB2E667" author="Keith D." page="45" pageId="43" pageNumber="44" year="2006">
<bibRefCitation id="EF8DABA4A61E0601FACEFDB1FEB2E667" author="Keith D." pageId="43" pageNumber="44" pagination="43 - 58" refId="ref44636" refString="Keith D. 2006. Especes nouvelles ou meconnues de Melolonthidae asiatiques (Coleoptera, Scarabaeidae). Symbioses Nouvelle Serie 17: 43 - 58." type="journal article" year="2006">Keith 2006: 45</bibRefCitation>
</treatmentCitation>
(nom. nud.). —
<treatmentCitation id="0ABDF044A61E0601FDEBFD92FCAAE664" author="Coca-Abia M. M." box="[528,840,571,597]" page="683" pageId="43" pageNumber="44" year="2008">
<bibRefCitation id="EF8DABA4A61E0601FDEBFD92FCAAE664" author="Coca-Abia M. M." box="[528,840,571,597]" pageId="43" pageNumber="44" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 683684</bibRefCitation>
</treatmentCitation>
, figs 1920 (
<typeStatus id="54A768F7A61E0601FC21FD92FBECE664" box="[986,1038,571,597]" pageId="43" pageNumber="44">type</typeStatus>
material; in key). —
<treatmentCitation id="0ABDF044A61E0601FAFEFD92FEB1E648" ID-DOI="http://doi.org/10.5281/zenodo.5971154" ID-Zenodo-Dep="5971154" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." httpUri="http://treatment.plazi.org/id/03B587C5ED25F35228EDFB9A5DF2D820" page="17" pageId="43" pageNumber="44" year="2018">
<bibRefCitation id="EF8DABA4A61E0601FAFEFD92FEB1E648" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." pageId="43" pageNumber="44" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A61E0601FAC6FD95FA8DE664" box="[1341,1391,571,597]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
. 2018: 17
</bibRefCitation>
</treatmentCitation>
, figs 5b, d, f, 6 (catalogue); 2019: 462 (species list; in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A61E0601FF46FD2BFE12E68E" blockId="43.[189,1398,466,703]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB0CF17BA61E0601FF46FD2BFE12E68E" pageId="43" pageNumber="44">
<taxonomicName id="4C1CADD6A61E0601FF46FD2BFE61E6AD" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[189,387,642,668]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A61E0601FF46FD2BFE61E6AD" box="[189,387,642,668]" italics="true" pageId="43" pageNumber="44">Miridiba ardoini</emphasis>
</taxonomicName>
<treatmentCitation id="0ABDF044A61E0601FE5FFD2BFDB4E6AC" author="Keith D." box="[420,598,642,669]" page="45" pageId="43" pageNumber="44" year="2006">
<bibRefCitation id="EF8DABA4A61E0601FE5FFD2BFDB4E6AC" author="Keith D." box="[420,598,642,669]" pageId="43" pageNumber="44" pagination="43 - 58" refId="ref44636" refString="Keith D. 2006. Especes nouvelles ou meconnues de Melolonthidae asiatiques (Coleoptera, Scarabaeidae). Symbioses Nouvelle Serie 17: 43 - 58." type="journal article" year="2006">Keith 2006: 45</bibRefCitation>
</treatmentCitation>
(as combination of
<taxonomicName id="4C1CADD6A61E0601FCB3FD2AFC22E6AD" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[840,960,642,668]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A61E0601FCB3FD2AFC22E6AD" box="[840,960,642,668]" italics="true" pageId="43" pageNumber="44">N. ardoini</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A61E0601FBFBFD2BFAE1E6AD" author="Coca-Abia M. M." box="[1024,1283,642,668]" page="676" pageId="43" pageNumber="44" year="2008">
<bibRefCitation id="EF8DABA4A61E0601FBFBFD2BFAE1E6AD" author="Coca-Abia M. M." box="[1024,1283,642,668]" pageId="43" pageNumber="44" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 676</bibRefCitation>
</treatmentCitation>
. — Keith 2010: 232 (synonym).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA61E0606FF46FD4FFCB3E702" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BA3D655A61E0601FF46FD4FFED4E731" blockId="43.[189,310,742,768]" box="[189,310,742,768]" pageId="43" pageNumber="44">
<heading id="D0EB6139A61E0601FF46FD4FFED4E731" bold="true" box="[189,310,742,768]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B9680A47A61E0601FF46FD4FFED4E731" bold="true" box="[189,310,742,768]" pageId="43" pageNumber="44">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A61E0606FF46FCBCFCB3E702" blockId="43.[189,1399,789,956]" lastBlockId="44.[187,1400,265,819]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
External morphology of adult (
<figureCitation id="1327CAD0A61E0601FDCEFCBCFD72E701" box="[565,656,789,816]" captionStart="Figs 6062" captionStartId="43.[190,242,1920,1946]" captionTargetBox="[287,1310,1023,1877]" captionTargetId="figure-196@43.[264,1323,1016,1881]" captionTargetPageId="43" captionText="Figs 6062. Miridiba tuberculipennis (Moser, 1913). 6061. Paralectotype, ♂ (MFNB). 62. Non-type, ♀ (BMNH). 60. Habitus in dorsal view. 61. Male genitalia: parameres and endophallus in lateral view. 62. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="43" pageNumber="44">Fig. 60</figureCitation>
). Body size 20.024.0 mm. Dorsal surface glabrous, at most, with tiny setae in each puncture hardly visible. Clypeus shorter than frons, concave, with anterior edge emarginate at middle. Frons very sloping, strong carina sunk at middle. Pronotal anterior margin with a perpendicular anterior face with sparse and short setae; posterior margin glabrous, finely flanged and with a row of punctures, except at middle where is a little bulky; lateral margins smooth and glabrous, strongly convergent and elevated at anterior half; anterior angles acute, not projected forward; posterior angles obtuse and rounded. Prosternal process cone-shaped. Scutellum glabrous, with punctures except at sides. Each elytron with a wart on apex only in females; epipleuron with setae at basal half. Foretibia with insertion of inner spur closer to third outer tooth than second one in females, in males almost equidistant between both outer teeth; with moderate carina. Mesotibia with transverse carina interrupted at middle in females, complete in males. Metatibia with transverse carina complete in both sexes; inner margin of dorsal surface with spines. Prepygidium roughly punctate and pubescent. Pygidium pubescent, irregularly punctate, truncated cone shaped, blunt, and apical margin moderately raised. Ventrite 5 moderately depressed at posterior half with pubescence longer on depressed part. Ventrite 6 moderately concave and bulging at anterior margin. Male genitalia: parameres (
<figureCitation id="1327CAD0A6190606FC2FFDEFFBCAE650" box="[980,1064,582,609]" captionStart="Figs 6062" captionStartId="43.[190,242,1920,1946]" captionTargetBox="[287,1310,1023,1877]" captionTargetId="figure-196@43.[264,1323,1016,1881]" captionTargetPageId="43" captionText="Figs 6062. Miridiba tuberculipennis (Moser, 1913). 6061. Paralectotype, ♂ (MFNB). 62. Non-type, ♀ (BMNH). 60. Habitus in dorsal view. 61. Male genitalia: parameres and endophallus in lateral view. 62. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="44" pageNumber="45">Fig. 61</figureCitation>
) without pubescence, dorsal branches starting slightly below collum, long, thin and ventrally curved, apices of ventral branches bent outwards ninety degrees. Endophallus (
<figureCitation id="1327CAD0A6190606FD74FD25FD07E696" box="[655,741,652,679]" captionStart="Figs 6062" captionStartId="43.[190,242,1920,1946]" captionTargetBox="[287,1310,1023,1877]" captionTargetId="figure-196@43.[264,1323,1016,1881]" captionTargetPageId="43" captionText="Figs 6062. Miridiba tuberculipennis (Moser, 1913). 6061. Paralectotype, ♂ (MFNB). 62. Non-type, ♀ (BMNH). 60. Habitus in dorsal view. 61. Male genitalia: parameres and endophallus in lateral view. 62. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="44" pageNumber="45">Fig. 61</figureCitation>
) asymmetrical in lateral view, with a dorsal part armed with a very strong spine and a ventral part with numerous setae. Apophysis of temones with distal ends separated from each other. Female genitalia (
<figureCitation id="1327CAD0A6190606FD29FD7BFCCEE6DC" box="[722,812,722,749]" captionStart="Figs 6062" captionStartId="43.[190,242,1920,1946]" captionTargetBox="[287,1310,1023,1877]" captionTargetId="figure-196@43.[264,1323,1016,1881]" captionTargetPageId="43" captionText="Figs 6062. Miridiba tuberculipennis (Moser, 1913). 6061. Paralectotype, ♂ (MFNB). 62. Non-type, ♀ (BMNH). 60. Habitus in dorsal view. 61. Male genitalia: parameres and endophallus in lateral view. 62. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="44" pageNumber="45">Fig. 62</figureCitation>
): genital chamber with vestigial sternites reduced to an area with a few sensillae. Median oviduct reduced with epithelium not sclerotized placed behind peduncle of bursa copulatrix that is strongly sclerotized.
</paragraph>
</subSubSection>
<caption id="DF6386DDA61E0601FF45F829FAB1E3D0" ID-DOI="http://doi.org/10.5281/zenodo.4770374" ID-Zenodo-Dep="4770374" httpUri="https://zenodo.org/record/4770374/files/figure.png" pageId="43" pageNumber="44" startId="43.[190,242,1920,1946]" targetBox="[287,1310,1023,1877]" targetPageId="43">
<paragraph id="8BA3D655A61E0601FF45F829FAB1E3D0" blockId="43.[190,1399,1920,2018]" pageId="43" pageNumber="44">
<emphasis id="B9680A47A61E0601FF45F829FEA8E3AB" bold="true" box="[190,330,1920,1946]" pageId="43" pageNumber="44">Figs 6062.</emphasis>
<taxonomicName id="4C1CADD6A61E0601FEA8F829FCC9E3AA" authority="(Moser, 1913)" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[339,811,1920,1947]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A61E0601FEA8F829FD9AE3AB" box="[339,632,1920,1946]" italics="true" pageId="43" pageNumber="44">Miridiba tuberculipennis</emphasis>
(Moser, 1913)
</taxonomicName>
.
<emphasis id="B9680A47A61E0601FCC3F829FC62E3AB" bold="true" box="[824,896,1920,1946]" pageId="43" pageNumber="44">6061</emphasis>
. Paralectotype, ♂ (MFNB).
<emphasis id="B9680A47A61E0601FB28F829FB12E3AB" bold="true" box="[1235,1264,1920,1946]" pageId="43" pageNumber="44">62</emphasis>
. Non-type, ♀ (BMNH).
<emphasis id="B9680A47A61E0601FEADF80AFE91E38C" bold="true" box="[342,371,1955,1981]" pageId="43" pageNumber="44">60</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A61E0601FD60F80AFD5AE38C" bold="true" box="[667,696,1955,1981]" pageId="43" pageNumber="44">61</emphasis>
. Male genitalia: parameres and endophallus in lateral view.
<emphasis id="B9680A47A61E0601FF45F86EFF39E3D0" bold="true" box="[190,219,1991,2017]" pageId="43" pageNumber="44">62</emphasis>
. Female genitalia in dorsal (A) and ventral (B) views. Scale bars: 60 = 5.0 mm; 6162 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA6190606FF46FCF3FBDEE242" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BA3D655A6190606FF46FCF3FE48E745" blockId="44.[189,426,858,884]" box="[189,426,858,884]" pageId="44" pageNumber="45">
<heading id="D0EB6139A6190606FF46FCF3FE48E745" bold="true" box="[189,426,858,884]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B9680A47A6190606FF46FCF3FE48E745" bold="true" box="[189,426,858,884]" pageId="44" pageNumber="45">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6190606FF11FC20FD91E792" blockId="44.[189,1399,904,1073]" box="[234,627,904,931]" pageId="44" pageNumber="45">
<emphasis id="B9680A47A6190606FF11FC20FD91E792" bold="true" box="[234,627,904,931]" pageId="44" pageNumber="45">
<typeStatus id="54A768F7A6190606FF11FC20FE84E792" box="[234,358,905,931]" pageId="44" pageNumber="45" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C1CADD6A6190606FE76FC20FD91E792" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[397,627,904,931]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE76FC20FD91E792" bold="true" box="[397,627,904,931]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6190606FF46FC05FD49E001" blockId="44.[189,1399,904,1073]" pageId="44" pageNumber="45">
<materialsCitation id="3B74DC08A6190606FF46FC05FD49E001" ID-GBIF-Occurrence="3114121350" collectionCode="MFNB" collectorName="A. K. Weld Downing" country="Myanmar" determinerName="M. Coca-Abia" location="Burma" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F30B96C5A6190606FF46FC05FEB4E7F7" box="[189,342,940,966]" name="Myanmar" pageId="44" pageNumber="45">MYANMAR</collectingCountry>
<specimenCount id="9D1A1DDCA6190606FE89FC04FE69E7F6" box="[370,395,941,967]" pageId="44" pageNumber="45" type="male"></specimenCount>
; “
<location id="8EC3808EA6190606FE58FC05FE1AE7F7" LSID="urn:lsid:plazi:treatment:03B56743A61E0607FDD7FEA1FD1CE5CD:8EC3808EA6190606FE58FC05FE1AE7F7" box="[419,504,940,966]" country="Myanmar" name="Burma" pageId="44" pageNumber="45">Burma</location>
” [Myanmar];
<collectorName id="26E9B383A6190606FD5BFC05FC76E7F7" box="[672,916,940,966]" pageId="44" pageNumber="45">A.K. Weld Downing</collectorName>
leg.; “Zool. Mus. Berlin” [
<location id="8EC3808EA6190606FB2CFC05FE55E7DB" LSID="urn:lsid:plazi:treatment:03B56743A61E0607FDD7FEA1FD1CE5CD:8EC3808EA6190606FB2CFC05FE55E7DB" country="Myanmar" name="Zoologisches Museum in Berlin" pageId="44" pageNumber="45">Zoologisches Museum in Berlin</location>
]; “
<taxonomicName id="4C1CADD6A6190606FE1EFC66FC6EE7DB" authority="Mos." authorityName="Moser" authorityYear="1913" box="[485,908,975,1002]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE1EFC66FCDBE7D8" box="[485,825,975,1001]" italics="true" pageId="44" pageNumber="45">Holotrichia tuberculipennis</emphasis>
Mos.
</taxonomicName>
Type ♂”; “SYNTYPUS
<taxonomicName id="4C1CADD6A6190606FB15FC66FDEFE03C" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FB15FC66FE90E03D" italics="true" pageId="44" pageNumber="45">Holotrichia tuberculipennis</emphasis>
Moser, 1913
</taxonomicName>
labelled by MNHUB 2004”; “Lectotype
<taxonomicName id="4C1CADD6A6190606FC14FC5BFE51E001" authority="(Moser, 1913) M. Coca-Abia" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FC14FC5BFAF6E03D" box="[1007,1300,1010,1036]" italics="true" pageId="44" pageNumber="45">Miridiba tuberculipennis</emphasis>
(Moser, 1913)
<determinerName id="7E7F3A91A6190606FEF3FBBFFE51E001" box="[264,435,1046,1072]" pageId="44" pageNumber="45">M. Coca-Abia</determinerName>
</taxonomicName>
, det. 2005”;
<collectionCode id="ED0D4E90A6190606FDABFBBFFD45E001" box="[592,679,1046,1072]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="44" pageNumber="45">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A6190606FF11FBFEFD46E043" blockId="44.[189,1198,1111,1173]" box="[234,676,1111,1138]" pageId="44" pageNumber="45">
<emphasis id="B9680A47A6190606FF11FBFEFD46E043" bold="true" box="[234,676,1111,1138]" pageId="44" pageNumber="45">
<typeStatus id="54A768F7A6190606FF11FBFEFE75E040" box="[234,407,1111,1137]" pageId="44" pageNumber="45" type="paralectotype">Paralectotype</typeStatus>
of
<taxonomicName id="4C1CADD6A6190606FE45FBFEFD46E043" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[446,676,1111,1138]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE45FBFEFD46E043" bold="true" box="[446,676,1111,1138]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6190606FF46FBD2FB4CE0A4" blockId="44.[189,1198,1111,1173]" box="[189,1198,1146,1173]" pageId="44" pageNumber="45">
<materialsCitation id="3B74DC08A6190606FF46FBD2FB48E0A4" ID-GBIF-Occurrence="3114121368" box="[189,1194,1146,1173]" collectionCode="MFNB" collectorName="A. K. Weld Downing" country="Myanmar" location="MYANMAR" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<collectingCountry id="F30B96C5A6190606FF46FBD2FEB4E0A4" box="[189,342,1147,1173]" name="Myanmar" pageId="44" pageNumber="45">MYANMAR</collectingCountry>
<specimenCount id="9D1A1DDCA6190606FE94FBD2FE64E0A4" box="[367,390,1147,1173]" pageId="44" pageNumber="45" type="female"></specimenCount>
; same collection data as for lectotype of
<taxonomicName id="4C1CADD6A6190606FC9CFBD2FBA1E0A5" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[871,1091,1146,1172]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FC9CFBD2FBA1E0A5" box="[871,1091,1146,1172]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
;
<collectionCode id="ED0D4E90A6190606FBA8FBD2FB48E0A4" box="[1107,1194,1147,1173]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="44" pageNumber="45">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A6190606FF11FB15FC92E0E6" blockId="44.[189,1399,1211,1379]" box="[234,880,1211,1239]" pageId="44" pageNumber="45">
<emphasis id="B9680A47A6190606FF11FB15FD4DE0E7" bold="true" box="[234,687,1211,1238]" pageId="44" pageNumber="45">
<typeStatus id="54A768F7A6190606FF11FB15FE40E0E7" box="[234,418,1212,1238]" pageId="44" pageNumber="45" type="paralectotype">Paralectotypes</typeStatus>
of
<taxonomicName id="4C1CADD6A6190606FE31FB15FD4DE0E7" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[458,687,1211,1238]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE31FB15FD4DE0E7" bold="true" box="[458,687,1211,1238]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
</emphasis>
(here indicated)
</paragraph>
<paragraph id="8BA3D655A6190606FF46FB76FC08E152" blockId="44.[189,1399,1211,1379]" pageId="44" pageNumber="45">
<materialsCitation id="3B74DC08A6190606FF46FB76FEF7E152" ID-GBIF-Occurrence="3114121330" collectionCode="BMNH" collectorName="A. K. Weld Downing Burma" country="Myanmar" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="MYANMAR" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<collectingCountry id="F30B96C5A6190606FF46FB76FEB4E0C8" box="[189,342,1247,1273]" name="Myanmar" pageId="44" pageNumber="45">MYANMAR</collectingCountry>
<specimenCount id="9D1A1DDCA6190606FE8AFB76FE7DE0CB" box="[369,415,1247,1274]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
; “Burma” [Myanmar]; “
<collectorName id="26E9B383A6190606FD38FB76FBFCE0C8" box="[707,1054,1247,1274]" pageId="44" pageNumber="45">A. K. Weld Downing Burma</collectorName>
”; “781”; “Andrewes Bequest B.M. 1922-221”; “
<taxonomicName id="4C1CADD6A6190606FE59FAABFD09E12D" authorityName="Moser" authorityYear="1913" box="[418,747,1282,1308]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE59FAABFD09E12D" box="[418,747,1282,1308]" italics="true" pageId="44" pageNumber="45">Holotrichia tuberculipennis</emphasis>
</taxonomicName>
Type Mos.”; “Type” [round label]; “Type” [red print label]; “Paralectotype
<taxonomicName id="4C1CADD6A6190606FE38FA8CFC77E171" authority="(Moser, 1913)" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[451,917,1317,1344]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE38FA8CFD05E10E" box="[451,743,1317,1343]" italics="true" pageId="44" pageNumber="45">Miridiba tuberculipennis</emphasis>
(Moser, 1913)
</taxonomicName>
<determinerName id="7E7F3A91A6190606FC66FA8CFBABE171" box="[925,1097,1317,1344]" pageId="44" pageNumber="45">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A6190606FB95FA8CFB0FE10E" box="[1134,1261,1317,1343]" pageId="44" pageNumber="45">Coca-Abia</determinerName>
det. 2019”;
<collectionCode id="ED0D4E90A6190606FF46FAE0FEF7E152" box="[189,277,1353,1379]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6190606FED5FAE0FC04E152" ID-GBIF-Occurrence="3114121305" box="[302,998,1353,1379]" collectionCode="BMNH" collectorName="A. K. Weld Downing Burma" country="Myanmar" determinerName="Coca-Abia" location="MYANMAR" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="9D1A1DDCA6190606FED5FAE0FEBFE152" box="[302,349,1353,1379]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
; same collection data as for preceding; cotype;
<collectionCode id="ED0D4E90A6190606FC71FAE0FC04E152" box="[906,998,1353,1379]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A6190606FF11FA23FDE6E195" blockId="44.[189,1399,1417,1515]" box="[234,516,1417,1444]" pageId="44" pageNumber="45">
<emphasis id="B9680A47A6190606FF11FA23FDE6E195" bold="true" box="[234,516,1417,1444]" pageId="44" pageNumber="45">
<typeStatus id="54A768F7A6190606FF11FA23FEBEE195" box="[234,348,1418,1444]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C1CADD6A6190606FE7FFA23FDE6E195" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[388,516,1417,1444]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FE7FFA23FDE6E195" bold="true" box="[388,516,1417,1444]" italics="true" pageId="44" pageNumber="45">M. ardoini</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6190606FF46FA04FB06E1DA" blockId="44.[189,1399,1417,1515]" pageId="44" pageNumber="45">
<collectingCountry id="F30B96C5A6190606FF46FA04FEA6E1F6" box="[189,324,1453,1479]" name="Vietnam" pageId="44" pageNumber="45">VIETNAM</collectingCountry>
<specimenCount id="9D1A1DDCA6190606FE98FA07FE98E1F9" box="[355,378,1454,1480]" pageId="44" pageNumber="45" type="female"></specimenCount>
; “Vietnam; 1960”; “Type”; “
<taxonomicName id="4C1CADD6A6190606FD27FA04FC1BE1F6" authority="Gao &amp; Coca-Abia, 2021" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[732,1017,1453,1479]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini" status="sp. nov.">
<emphasis id="B9680A47A6190606FD27FA04FC1BE1F6" box="[732,1017,1453,1479]" italics="true" pageId="44" pageNumber="45">Neodontocnema ardoini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BB73CA6190606FBFFFA07FBDCE1F9" box="[1028,1086,1454,1480]" pageId="44" pageNumber="45" rank="species">n sp.</taxonomicNameLabel>
det G.
<bibRefCitation id="EF8DABA4A6190606FB66FA07FAFCE1F6" author="Frey G." box="[1181,1310,1453,1480]" pageId="44" pageNumber="45" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
Type”; “
<typeStatus id="54A768F7A6190606FF3CFA79FED5E1DB" box="[199,311,1488,1514]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C1CADD6A6190606FEC5FA79FD79E1DA" authority="(Frey, 1970)" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[318,667,1488,1515]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FEC5FA79FDE1E1DB" box="[318,515,1488,1514]" italics="true" pageId="44" pageNumber="45">Miridiba ardoini</emphasis>
(
<bibRefCitation id="EF8DABA4A6190606FDE8FA78FD71E1DA" author="Frey G." box="[531,659,1489,1515]" pageId="44" pageNumber="45" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
)
</taxonomicName>
Chuan-bu Gao &amp; Coca-Abia det. 2017”;
<collectionCode id="ED0D4E90A6190606FB7FFA78FB02E1DA" box="[1156,1248,1489,1515]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="44" pageNumber="45">NHMB</collectionCode>
.
</paragraph>
<paragraph id="8BA3D655A6190606FF11F9BBFD51E21D" blockId="44.[189,1398,1553,1651]" box="[234,691,1553,1580]" pageId="44" pageNumber="45">
<emphasis id="B9680A47A6190606FF11F9BBFD51E21D" bold="true" box="[234,691,1553,1580]" pageId="44" pageNumber="45">
Other material of
<taxonomicName id="4C1CADD6A6190606FE36F9BBFD51E21D" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[461,691,1553,1580]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE36F9BBFD51E21D" bold="true" box="[461,691,1553,1580]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6190606FF46F99CFBDEE242" blockId="44.[189,1398,1553,1651]" pageId="44" pageNumber="45">
<materialsCitation id="3B74DC08A6190606FF46F99CFCF6E261" ID-GBIF-Occurrence="3114121378" box="[189,788,1589,1616]" collectionCode="BMNH" collectorName="A. K. Weld" country="Myanmar" location="Burma" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F30B96C5A6190606FF46F99CFEB4E27E" box="[189,342,1589,1615]" name="Myanmar" pageId="44" pageNumber="45">MYANMAR</collectingCountry>
<specimenCount id="9D1A1DDCA6190606FE97F99CFE78E261" box="[364,410,1589,1616]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
;
<location id="8EC3808EA6190606FE5EF99FFE1BE261" LSID="urn:lsid:plazi:treatment:03B56743A61E0607FDD7FEA1FD1CE5CD:8EC3808EA6190606FE5EF99FFE1BE261" box="[421,505,1590,1616]" country="Myanmar" name="Burma" pageId="44" pageNumber="45">Burma</location>
;
<collectorName id="26E9B383A6190606FDFAF99CFD9FE27E" box="[513,637,1589,1615]" pageId="44" pageNumber="45">A.K. Weld</collectorName>
leg.;
<collectionCode id="ED0D4E90A6190606FD47F99FFCF6E261" box="[700,788,1590,1616]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6190606FCD0F99CFA94E261" ID-GBIF-Occurrence="3114121335" box="[811,1398,1589,1616]" collectionCode="BMNH" collectorName="Burma &amp; A. K. Weld" country="Myanmar" location="MYANMAR" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1A1DDCA6190606FCD0F99CFCB4E261" box="[811,854,1589,1616]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0D4E90A6190606FAE5F99FFA94E261" box="[1310,1398,1590,1616]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6190606FF34F9F1FBDAE242" ID-GBIF-Occurrence="3114121371" box="[207,1080,1624,1651]" collectingDate="1916-04" collectionCode="BMNH" collectorName="F. M. Mackwood" country="Myanmar" elevation="1219" location="Kalaw" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Shan State">
<specimenCount id="9D1A1DDCA6190606FF34F9F1FF1EE242" box="[207,252,1624,1651]" pageId="44" pageNumber="45" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49D818B7A6190606FEF1F9F1FE68E243" box="[266,394,1624,1650]" country="Myanmar" name="Shan" pageId="44" pageNumber="45">Shan State</collectingRegion>
,
<location id="8EC3808EA6190606FE6EF9F1FE07E243" LSID="urn:lsid:plazi:treatment:03B56743A61E0607FDD7FEA1FD1CE5CD:8EC3808EA6190606FE6EF9F1FE07E243" box="[405,485,1624,1650]" country="Myanmar" name="Kalaw" pageId="44" pageNumber="45" stateProvince="Shan State">Kalaw</location>
;
<quantity id="4CE47BB0A6190606FE0AF9F1FDAAE243" box="[497,584,1624,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="44" pageNumber="45" unit="ft" value="4000.0">
<elevation id="00313166A6190606FE0AF9F1FDAAE243" box="[497,584,1624,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="44" pageNumber="45" unit="ft" value="4000.0">4000 ft</elevation>
</quantity>
;
<date id="FFA2F095A6190606FDA8F9F1FD2EE243" box="[595,716,1624,1650]" pageId="44" pageNumber="45" value="1916-04">
<collectingDate id="EFE6097DA6190606FDA8F9F1FD2EE243" box="[595,716,1624,1650]" pageId="44" pageNumber="45" value="1916-04">Apr. 1916</collectingDate>
</date>
;
<collectorName id="26E9B383A6190606FD23F9F0FC78E243" box="[728,922,1624,1651]" pageId="44" pageNumber="45">F.M. Mackwood</collectorName>
leg.;
<collectionCode id="ED0D4E90A6190606FC27F9F0FBDAE242" box="[988,1080,1625,1651]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6190607FF46F933FB80E5BC" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BA3D655A6190606FF46F933FECDE285" blockId="44.[189,303,1690,1716]" box="[189,303,1690,1716]" pageId="44" pageNumber="45">
<heading id="D0EB6139A6190606FF46F933FECDE285" bold="true" box="[189,303,1690,1716]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B9680A47A6190606FF46F933FECDE285" bold="true" box="[189,303,1690,1716]" pageId="44" pageNumber="45">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6190607FF46F961FB80E5BC" blockId="44.[189,1401,1736,2009]" lastBlockId="45.[189,1398,265,397]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<taxonomicName id="4C1CADD6A6190606FF46F961FE3DE2D3" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[189,479,1736,1762]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FF46F961FE3DE2D3" box="[189,479,1736,1762]" italics="true" pageId="44" pageNumber="45">Neodontocnema ardoini</emphasis>
</taxonomicName>
was described based on only
<specimenCount id="9D1A1DDCA6190606FC92F960FC10E2D3" box="[873,1010,1736,1763]" pageId="44" pageNumber="45" type="female">one female</specimenCount>
specimen from
<collectingCountry id="F30B96C5A6190606FB3EF961FACAE2D3" box="[1221,1320,1736,1762]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
(
<bibRefCitation id="EF8DABA4A6190606FAC4F961FF1EE337" author="Frey G." pageId="44" pageNumber="45" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey 1970</bibRefCitation>
). The external morphology described by
<bibRefCitation id="EF8DABA4A6190606FD09F945FC62E337" author="Frey G." box="[754,896,1772,1798]" pageId="44" pageNumber="45" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey (1970)</bibRefCitation>
led
<bibRefCitation id="EF8DABA4A6190606FC41F942FBB6E337" author="Keith D." box="[954,1108,1771,1798]" pageId="44" pageNumber="45" pagination="43 - 58" refId="ref44636" refString="Keith D. 2006. Especes nouvelles ou meconnues de Melolonthidae asiatiques (Coleoptera, Scarabaeidae). Symbioses Nouvelle Serie 17: 43 - 58." type="journal article" year="2006">Keith (2006)</bibRefCitation>
to consider the transfer of
<taxonomicName id="4C1CADD6A6190606FF24F8A6FEBBE318" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[223,345,1807,1833]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FF24F8A6FEBBE318" box="[223,345,1807,1833]" italics="true" pageId="44" pageNumber="45">N. ardoini</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A6190606FE7EF8A6FE09E318" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[389,491,1807,1833]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6190606FE7EF8A6FE09E318" box="[389,491,1807,1833]" italics="true" pageId="44" pageNumber="45">Miridiba</emphasis>
</taxonomicName>
. Later, Keith (2010) synonymized
<taxonomicName id="4C1CADD6A6190606FC6EF8A6FBF1E318" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[917,1043,1807,1833]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FC6EF8A6FBF1E318" box="[917,1043,1807,1833]" italics="true" pageId="44" pageNumber="45">M. ardoini</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A6190606FBA0F8A6FADBE318" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[1115,1337,1807,1833]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FBA0F8A6FADBE318" box="[1115,1337,1807,1833]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
after studying the type of
<taxonomicName id="4C1CADD6A6190606FE44F89AFDA2E37D" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[447,576,1842,1868]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FE44F89AFDA2E37D" box="[447,576,1842,1868]" italics="true" pageId="44" pageNumber="45">M. ardoini</emphasis>
</taxonomicName>
and two non-type specimens of
<taxonomicName id="4C1CADD6A6190606FC2CF89BFBDDE37D" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[983,1087,1842,1868]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6190606FC2CF89BFBDDE37D" box="[983,1087,1842,1868]" italics="true" pageId="44" pageNumber="45">Miridiba</emphasis>
</taxonomicName>
from
<collectingCountry id="F30B96C5A6190606FB74F89BFB10E37D" box="[1167,1266,1842,1868]" name="Vietnam" pageId="44" pageNumber="45">Vietnam</collectingCountry>
(male and female), concluding that the features of these
<specimenCount id="9D1A1DDCA6190606FD15F8FFFC40E35E" box="[750,930,1877,1904]" pageId="44" pageNumber="45" type="generic">two specimens</specimenCount>
, female and male, matched well those of
<taxonomicName id="4C1CADD6A6190606FF19F8D0FE81E3A3" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[226,355,1912,1938]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FF19F8D0FE81E3A3" box="[226,355,1912,1938]" italics="true" pageId="44" pageNumber="45">M. ardoini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1CADD6A6190606FE5CF8D0FD6AE3A3" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[423,648,1912,1938]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FE5CF8D0FD6AE3A3" box="[423,648,1912,1938]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
, respectively. However, to establish this synonymy, types of
<taxonomicName id="4C1CADD6A6190606FF46F835FE79E384" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[189,411,1947,1973]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FF46F835FE79E384" box="[189,411,1947,1973]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
should also have been studied. In this work, we have studied types of
<taxonomicName id="4C1CADD6A6190606FB01F835FA9BE384" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[1274,1401,1947,1973]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6190606FB01F835FA9BE384" box="[1274,1401,1947,1973]" italics="true" pageId="44" pageNumber="45">M. ardoini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1CADD6A6190606FF0BF816FE2FE3E8" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[240,461,1983,2009]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6190606FF0BF816FE2FE3E8" box="[240,461,1983,2009]" italics="true" pageId="44" pageNumber="45">M. tuberculipennis</emphasis>
</taxonomicName>
and verified that features of the external morphology and female genitalia that characterize
<taxonomicName id="4C1CADD6A6180607FEACFEA3FDD5E512" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[343,567,265,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6180607FEACFEA3FDD5E512" box="[343,567,265,291]" italics="true" pageId="45" pageNumber="46">M. tuberculipennis</emphasis>
</taxonomicName>
are present in
<taxonomicName id="4C1CADD6A6180607FD0CFEA3FC9AE512" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[759,888,265,291]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6180607FD0CFEA3FC9AE512" box="[759,888,265,291]" italics="true" pageId="45" pageNumber="46">M. ardoini</emphasis>
</taxonomicName>
. Therefore, we confirm the synonymy of
<taxonomicName id="4C1CADD6A6180607FF46FE84FED9E577" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[189,315,300,326]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6180607FF46FE84FED9E577" box="[189,315,300,326]" italics="true" pageId="45" pageNumber="46">M. ardoini</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A6180607FE78FE84FD82E577" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[387,608,300,326]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6180607FE78FE84FD82E577" box="[387,608,300,326]" italics="true" pageId="45" pageNumber="46">M. tuberculipennis</emphasis>
</taxonomicName>
, and we include this species into morphotype I “
<taxonomicName id="4C1CADD6A6180607FB55FE85FAA3E577" box="[1198,1345,300,326]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6180607FB55FE85FAA3E577" box="[1198,1345,300,326]" italics="true" pageId="45" pageNumber="46">Trichophora</emphasis>
</taxonomicName>
”. In addition, two further former
<typeStatus id="54A768F7A6180607FDEBFEF9FD95E55B" box="[528,631,336,362]" pageId="45" pageNumber="46" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C1CADD6A6180607FD66FEF9FC9BE558" authorityName="M. Coca-Abia" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[669,889,335,361]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tuberculipennis">
<emphasis id="B9680A47A6180607FD66FEF9FC9BE558" box="[669,889,335,361]" italics="true" pageId="45" pageNumber="46">M. tuberculipennis</emphasis>
</taxonomicName>
are recognised as
<typeStatus id="54A768F7A6180607FBA8FEE6FB1EE558" box="[1107,1276,335,361]" pageId="45" pageNumber="46" type="paralectotype">paralectotypes</typeStatus>
(BMNH), and the unique female specimen of
<taxonomicName id="4C1CADD6A6180607FDA5FEDAFD38E5BD" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" box="[606,730,370,396]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="B9680A47A6180607FDA5FEDAFD38E5BD" box="[606,730,370,396]" italics="true" pageId="45" pageNumber="46">M. ardoini</emphasis>
</taxonomicName>
is labelled as
<typeStatus id="54A768F7A6180607FC7AFEDBFC0AE5BD" box="[897,1000,370,396]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
(NHMB).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6180607FF46FE1DFD1CE5CD" box="[189,766,436,509]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BA3D655A6180607FF46FE1DFEB4E5FF" blockId="45.[189,342,436,462]" box="[189,342,436,462]" pageId="45" pageNumber="46">
<heading id="D0EB6139A6180607FF46FE1DFEB4E5FF" bold="true" box="[189,342,436,462]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B9680A47A6180607FF46FE1DFEB4E5FF" bold="true" box="[189,342,436,462]" pageId="45" pageNumber="46">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6180607FF46FE4AFD1CE5CD" blockId="45.[189,766,482,509]" box="[189,766,482,509]" pageId="45" pageNumber="46">
<collectingCountry id="F30B96C5A6180607FF46FE4AFF1BE5CC" box="[189,249,483,509]" name="Laos" pageId="45" pageNumber="46">Laos</collectingCountry>
,
<collectingCountry id="F30B96C5A6180607FEFEFE4AFE95E5CC" box="[261,375,483,509]" name="Myanmar" pageId="45" pageNumber="46">Myanmar</collectingCountry>
(
<collectingRegion id="49D818B7A6180607FE7DFE4BFDEBE5CC" box="[390,521,482,509]" country="Myanmar" name="Shan" pageId="45" pageNumber="46">Shan State</collectingRegion>
),
<collectingCountry id="F30B96C5A6180607FDE7FE4BFD65E5CD" box="[540,647,482,508]" name="Thailand" pageId="45" pageNumber="46">Thailand</collectingCountry>
,
<collectingCountry id="F30B96C5A6180607FD68FE4BFD18E5CD" box="[659,762,482,508]" name="Vietnam" pageId="45" pageNumber="46">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>