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<mods:title id="422EAE72E0C6D7F3B8750D9352FCC470">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<taxonomicName id="4C1CADD6A614060BFDBCF8E6FC0EE35B" authority="Nomura, 1977" authorityName="Nomura" authorityYear="1977" box="[583,1004,1871,1898]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A614060BFDBCF8E6FCD9E35B" bold="true" box="[583,827,1871,1898]" italics="true" pageId="33" pageNumber="34">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A614060BFCB9F8F9FC0EE35B" author="Nomura S." box="[834,1004,1872,1898]" pageId="33" pageNumber="34" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura, 1977</bibRefCitation>
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<paragraph id="8BA3D655A614060BFD23F8DAFCB9E3BF" blockId="33.[583,1004,1871,1934]" box="[728,859,1907,1934]" pageId="33" pageNumber="34">
<figureCitation id="1327CAD0A614060BFD23F8DAFCB9E3BF" box="[728,859,1907,1934]" captionStart="Figs 4750" captionStartId="34.[190,242,1890,1916]" captionTargetBox="[280,1312,1066,1839]" captionTargetId="figure-293@34.[264,1323,1059,1852]" captionTargetPageId="34" captionText="Figs 4750. Miridiba kuraruana Nomura, 1977. 4749. holotype (NSMT). 50. Paratype (NSMT). 47. Habitus in dorsal view (holotype). 48. Parameres in dorsal (A) and lateral (B) views. 49. Endophallus and temones in dorsal view. 50. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="33" pageNumber="34">Figs 4750</figureCitation>
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<taxonomicName id="4C1CADD6A614060BFF46F81DFD64E3FE" authority="Nomura, 1977: 89" authorityName="Nomura" authorityPageNumber="89" authorityYear="1977" box="[189,646,1972,1999]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A614060BFF46F81DFE4BE3FF" box="[189,425,1972,1998]" italics="true" pageId="33" pageNumber="34">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A614060BFE4BF81CFD64E3FE" author="Nomura S." box="[432,646,1973,1999]" pageId="33" pageNumber="34" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura, 1977: 89</bibRefCitation>
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(
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loc.: Kenting,
<collectingCountry id="F30B96C5A614060BFC83F81DFC33E3FF" box="[888,977,1972,1998]" name="Taiwan" pageId="33" pageNumber="34">Taiwan</collectingCountry>
,
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).
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<taxonomicName id="4C1CADD6A6170608FF46FEA0FE4AE512" authorityName="Nomura" authorityYear="1977" box="[189,424,265,291]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6170608FF46FEA0FE4AE512" box="[189,424,265,291]" italics="true" pageId="34" pageNumber="35">Miridiba kuraruana</emphasis>
</taxonomicName>
<treatmentCitation id="0ABDF044A6170608FE38FEA0FD19E512" author="Smetana A. &amp; Kral D." box="[451,763,265,291]" page="222" pageId="34" pageNumber="35" year="2006">
<bibRefCitation id="EF8DABA4A6170608FE38FEA0FD19E512" author="Smetana A. &amp; Kral D." box="[451,763,265,291]" pageId="34" pageNumber="35" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A6170608FC4DFEA0FB56E512" author="Coca-Abia M. M." box="[950,1204,265,291]" page="680" pageId="34" pageNumber="35" year="2008">
<bibRefCitation id="EF8DABA4A6170608FC4DFEA0FB56E512" author="Coca-Abia M. M." box="[950,1204,265,291]" pageId="34" pageNumber="35" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 680</bibRefCitation>
</treatmentCitation>
, 683 (in key). —
<treatmentCitation id="0ABDF044A6170608FF12FE85FE21E576" ID-DOI="http://doi.org/10.5281/zenodo.6115517" ID-Zenodo-Dep="6115517" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[233,451,300,327]" httpUri="http://treatment.plazi.org/id/03CA87D3FFBBFF8A98808D9EFD31CECE" page="523" pageId="34" pageNumber="35" year="2015">
<bibRefCitation id="EF8DABA4A6170608FF12FE85FE21E576" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[233,451,300,327]" pageId="34" pageNumber="35" pagination="521 - 536" refId="ref44907" refString="Li C. - L., Yang P. - S. &amp; Wang C. - C. 2015. A review of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) of Taiwan. Zootaxa 3955 (4): 521 - 536. https: // doi. org / 10.11646 / zootaxa. 3955.4.4" type="journal article" year="2015">
Li
<emphasis id="B9680A47A6170608FEF0FE84FEDFE577" box="[267,317,300,326]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2015: 523
</bibRefCitation>
</treatmentCitation>
, figs 1, 7, 13, 19, 2526 (redescription; in key). —
<treatmentCitation id="0ABDF044A6170608FBEAFE85FB07E577" author="Bezdek A." box="[1041,1253,300,326]" page="272" pageId="34" pageNumber="35" year="2016">
<bibRefCitation id="EF8DABA4A6170608FBEAFE85FB07E577" author="Bezdek A." box="[1041,1253,300,326]" pageId="34" pageNumber="35" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 272</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A6170608FEF4FEF9FE12E55B" ID-DOI="http://doi.org/10.5281/zenodo.5971120" ID-Zenodo-Dep="5971120" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[271,496,335,362]" httpUri="http://treatment.plazi.org/id/03B587C5ED38F34F28EDFF415C36DC6C" page="14" pageId="34" pageNumber="35" year="2018">
<bibRefCitation id="EF8DABA4A6170608FEF4FEF9FE12E55B" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[271,496,335,362]" pageId="34" pageNumber="35" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A6170608FEBCFEF9FE99E558" box="[327,379,335,361]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2018: 14
</bibRefCitation>
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(catalogue;
<typeStatus id="54A768F7A6170608FD79FEF9FD57E55B" box="[642,693,336,362]" pageId="34" pageNumber="35">type</typeStatus>
material); 2019: 461 (species list; in key).
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</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6170609FF46FE38FABDE5BA" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BA3D655A6170608FF46FE38FED4E59A" blockId="34.[189,310,401,427]" box="[189,310,401,427]" pageId="34" pageNumber="35">
<heading id="D0EB6139A6170608FF46FE38FED4E59A" bold="true" box="[189,310,401,427]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B9680A47A6170608FF46FE38FED4E59A" bold="true" box="[189,310,401,427]" pageId="34" pageNumber="35">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6170609FF46FE16FABDE5BA" blockId="34.[188,1401,447,993]" lastBlockId="35.[187,1400,265,395]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
External morphology of adult (
<figureCitation id="1327CAD0A6170608FDDBFE16FD97E5E8" box="[544,629,447,473]" captionStart="Figs 4750" captionStartId="34.[190,242,1890,1916]" captionTargetBox="[280,1312,1066,1839]" captionTargetId="figure-293@34.[264,1323,1059,1852]" captionTargetPageId="34" captionText="Figs 4750. Miridiba kuraruana Nomura, 1977. 4749. holotype (NSMT). 50. Paratype (NSMT). 47. Habitus in dorsal view (holotype). 48. Parameres in dorsal (A) and lateral (B) views. 49. Endophallus and temones in dorsal view. 50. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="34" pageNumber="35">Fig. 47</figureCitation>
). Body size 18.920.0 mm. Dorsal surface shiny. Clypeus concave, with oblique sides, anterior edge moderately emarginate at middle and pubescence short. Frons with conspicuous pubescence; punctures thicker than those of clypeus; angular strong carina with a midvertex. Pronotum with hardly visible short setae in each puncture; pronotal anterior margin sloped forward, glabrous, at most, some short setae; posterior margin glabrous; lateral margins strongly serrated, with conspicuous short pubescence; anterior angles obtuse, not projected forward; posterior angles obtuse and rounded. Prosternal process tongue-shaped. Scutellum punctate, with short setae. Elytra with punctures regularly distributed, disc with very short setae in each puncture hardly visible; basal and lateral part with conspicuous long pubescence; epipleuron with pubescence except on apex. Foretibia with dorsal strong carina, insertion of inner spur in male equidistant between second and third outer tooth. Meso- and metatibia with complete transverse carina, inner margin of dorsal surface with spines. Prepygidium regularly punctate, with short but conspicuous pubescence. Pygidium irregularly punctate, with conspicuous pubescence, longer on apex; apical margin moderately raised. Ventrite 5 moderately depressed at posterior half, with strong punctures and pubescence, longer on depressed part. Ventrite 6 with punctures irregularly distributed, conspicuous pubescence, bulky at anterior margin, bulge without punctures. Male genitalia: parameres (
<figureCitation id="1327CAD0A6170608FD7BFC6EFD34E7D0" box="[640,726,967,993]" captionStart="Figs 4750" captionStartId="34.[190,242,1890,1916]" captionTargetBox="[280,1312,1066,1839]" captionTargetId="figure-293@34.[264,1323,1059,1852]" captionTargetPageId="34" captionText="Figs 4750. Miridiba kuraruana Nomura, 1977. 4749. holotype (NSMT). 50. Paratype (NSMT). 47. Habitus in dorsal view (holotype). 48. Parameres in dorsal (A) and lateral (B) views. 49. Endophallus and temones in dorsal view. 50. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="34" pageNumber="35">Fig. 48</figureCitation>
) with setae on ventral side of dorsal branches and dorsal side of ventral ones; in lateral view, moderately bent towards apices at two stretches; apices of ventral branches thin and slightly curved outwards and upward, with a small apical tip. Endophallus (
<figureCitation id="1327CAD0A6160609FB1FFE85FAD5E577" box="[1252,1335,300,326]" captionStart="Figs 4750" captionStartId="34.[190,242,1890,1916]" captionTargetBox="[280,1312,1066,1839]" captionTargetId="figure-293@34.[264,1323,1059,1852]" captionTargetPageId="34" captionText="Figs 4750. Miridiba kuraruana Nomura, 1977. 4749. holotype (NSMT). 50. Paratype (NSMT). 47. Habitus in dorsal view (holotype). 48. Parameres in dorsal (A) and lateral (B) views. 49. Endophallus and temones in dorsal view. 50. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="35" pageNumber="36">Fig. 49</figureCitation>
) with spines at distal end; all proximal part of sac covered with trichodea sensilla. Female genitalia (
<figureCitation id="1327CAD0A6160609FB00FEE7FAB2E558" box="[1275,1360,334,361]" captionStart="Figs 4750" captionStartId="34.[190,242,1890,1916]" captionTargetBox="[280,1312,1066,1839]" captionTargetId="figure-293@34.[264,1323,1059,1852]" captionTargetPageId="34" captionText="Figs 4750. Miridiba kuraruana Nomura, 1977. 4749. holotype (NSMT). 50. Paratype (NSMT). 47. Habitus in dorsal view (holotype). 48. Parameres in dorsal (A) and lateral (B) views. 49. Endophallus and temones in dorsal view. 50. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="35" pageNumber="36">Fig. 50</figureCitation>
) as that of morphotype I, with two dark rounded sclerotized areas on dorsal membrane of median oviduct.
</paragraph>
</subSubSection>
<caption id="DF6386DDA6170608FF45F8CBFEFCE3D7" ID-DOI="http://doi.org/10.5281/zenodo.4770360" ID-Zenodo-Dep="4770360" httpUri="https://zenodo.org/record/4770360/files/figure.png" pageId="34" pageNumber="35" startId="34.[190,242,1890,1916]" targetBox="[280,1312,1066,1839]" targetPageId="34">
<paragraph id="8BA3D655A6170608FF45F8CBFEFCE3D7" blockId="34.[187,1399,1890,2022]" pageId="34" pageNumber="35">
<emphasis id="B9680A47A6170608FF45F8CBFEB2E34D" bold="true" box="[190,336,1890,1916]" pageId="34" pageNumber="35">Figs 4750.</emphasis>
<taxonomicName id="4C1CADD6A6170608FEA0F8CBFCE9E34D" authority="Nomura, 1977" authorityName="Nomura" authorityYear="1977" box="[347,779,1890,1917]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6170608FEA0F8CBFDAEE34D" box="[347,588,1890,1916]" italics="true" pageId="34" pageNumber="35">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A6170608FDA2F8CAFCE9E34D" author="Nomura S." box="[601,779,1890,1917]" pageId="34" pageNumber="35" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura, 1977</bibRefCitation>
</taxonomicName>
.
<emphasis id="B9680A47A6170608FCE7F8CBFC87E34D" bold="true" box="[796,869,1890,1916]" pageId="34" pageNumber="35">4749</emphasis>
. holotype (NSMT).
<emphasis id="B9680A47A6170608FB9EF8CBFB60E34D" bold="true" box="[1125,1154,1890,1916]" pageId="34" pageNumber="35">50</emphasis>
. Paratype (NSMT).
<emphasis id="B9680A47A6170608FF46F82CFF38E3AE" bold="true" box="[189,218,1925,1951]" pageId="34" pageNumber="35">47</emphasis>
. Habitus in dorsal view (holotype).
<emphasis id="B9680A47A6170608FD8CF82CFD76E3AE" bold="true" box="[631,660,1925,1951]" pageId="34" pageNumber="35">48</emphasis>
. Parameres in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A6170608FB43F82CFB37E3AE" bold="true" box="[1208,1237,1925,1951]" pageId="34" pageNumber="35">49</emphasis>
. Endophallus and temones in dorsal view.
<emphasis id="B9680A47A6170608FDE2F801FDD4E3F3" bold="true" box="[537,566,1960,1986]" pageId="34" pageNumber="35">50</emphasis>
. Female genitalia in dorsal view. Scale bars: 47 = 5.0 mm; 4850 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA6160609FF46FE03FAD5E72E" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BA3D655A6160609FF46FE03FE48E5F5" blockId="35.[189,426,426,452]" box="[189,426,426,452]" pageId="35" pageNumber="36">
<heading id="D0EB6139A6160609FF46FE03FE48E5F5" bold="true" box="[189,426,426,452]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B9680A47A6160609FF46FE03FE48E5F5" bold="true" box="[189,426,426,452]" pageId="35" pageNumber="36">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6160609FF11FE71FEBEE5C3" blockId="35.[189,1398,472,568]" box="[234,348,472,498]" pageId="35" pageNumber="36">
<emphasis id="B9680A47A6160609FF11FE71FEBEE5C3" bold="true" box="[234,348,472,498]" pageId="35" pageNumber="36">
<typeStatus id="54A768F7A6160609FF11FE71FEBEE5C3" box="[234,348,472,498]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6160609FF46FE52FB43E609" blockId="35.[189,1398,472,568]" pageId="35" pageNumber="36">
<materialsCitation id="3B74DC08A6160609FF46FE52FB43E609" ID-GBIF-Occurrence="3114121301" collectingDate="1971-06-12" collectionCode="NSMT" collectorName="Y. Maeda" country="China" county="Kenting Park" location="Taiwan" municipality="Taiwan" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F30B96C5A6160609FF46FE52FEFAE624" box="[189,280,507,533]" name="China" pageId="35" pageNumber="36">CHINA</collectingCountry>
<specimenCount id="9D1A1DDCA6160609FED4FE55FEAAE627" box="[303,328,508,534]" pageId="35" pageNumber="36" type="male"></specimenCount>
; “Formosa [
<collectingMunicipality id="6BC74C2FA6160609FE2FFE53FDD1E624" box="[468,563,506,533]" pageId="35" pageNumber="36">Taiwan</collectingMunicipality>
],
<collectingCounty id="62C2AED9A6160609FDBEFE52FD01E624" box="[581,739,507,533]" pageId="35" pageNumber="36">Kenting Park</collectingCounty>
;
<date id="FFA2F095A6160609FD16FE52FC67E624" box="[749,901,507,534]" pageId="35" pageNumber="36" value="1971-06-12">
<collectingDate id="EFE6097DA6160609FD16FE52FC67E624" box="[749,901,507,534]" pageId="35" pageNumber="36" value="1971-06-12">12 Jun. 1971</collectingDate>
</date>
;
<collectorName id="26E9B383A6160609FC74FE55FC1CE624" box="[911,1022,507,534]" pageId="35" pageNumber="36">Y. Maeda</collectorName>
leg.”; “light”; “Sizumu Nomura 1981 Bequest, 1981”; “♂”; “Holotype
<taxonomicName id="4C1CADD6A6160609FD7FFDB7FBD0E609" authority="Nomura (1977)" authorityName="Nomura" authorityYear="1977" box="[644,1074,542,568]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FD7FFDB7FC92E609" box="[644,880,542,568]" italics="true" pageId="35" pageNumber="36">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A6160609FC8CFDB7FBD0E609" author="Nomura S." box="[887,1074,542,568]" pageId="35" pageNumber="36" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura (1977)</bibRefCitation>
</taxonomicName>
”;
<collectionCode id="ED0D4E90A6160609FBB2FDB7FB7EE609" box="[1097,1180,542,568]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="35" pageNumber="36">NSMT</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A6160609FF11FDFEFE85E640" blockId="35.[189,1399,599,799]" box="[234,359,599,625]" pageId="35" pageNumber="36">
<emphasis id="B9680A47A6160609FF11FDFEFE85E640" bold="true" box="[234,359,599,625]" pageId="35" pageNumber="36">
<typeStatus id="54A768F7A6160609FF11FDFEFE85E640" box="[234,359,599,625]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A6160609FF46FDD3FAD5E72E" blockId="35.[189,1399,599,799]" pageId="35" pageNumber="36">
<materialsCitation id="3B74DC08A6160609FF46FDD3FC0DE687" ID-GBIF-Occurrence="3114121339" collectingDate="1977-05-07" collectionCode="NSMT" collectorName="M. Sakaino" country="China" location="Taiwan" municipality="Kenting" pageId="35" pageNumber="36" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F30B96C5A6160609FF46FDD3FEFAE6A5" box="[189,280,634,660]" name="China" pageId="35" pageNumber="36">CHINA</collectingCountry>
<specimenCount id="9D1A1DDCA6160609FEC9FDD3FEBDE6A5" box="[306,351,634,660]" pageId="35" pageNumber="36" type="female">1 ♀</specimenCount>
; “
<location id="8EC3808EA6160609FE83FDD3FE36E6A5" LSID="urn:lsid:plazi:treatment:03B56743A6140609FDBCF8E6FE9EE067:8EC3808EA6160609FE83FDD3FE36E6A5" box="[376,468,634,660]" country="China" municipality="Kenting" name="Taiwan" pageId="35" pageNumber="36">Taiwan</location>
,
<collectingMunicipality id="6BC74C2FA6160609FE24FDD3FDA3E6A5" box="[479,577,634,660]" pageId="35" pageNumber="36">Kenting</collectingMunicipality>
;
<date id="FFA2F095A6160609FDB5FDD3FD3DE6A5" box="[590,735,634,660]" pageId="35" pageNumber="36" value="1977-05-07">
<collectingDate id="EFE6097DA6160609FDB5FDD3FD3DE6A5" box="[590,735,634,660]" pageId="35" pageNumber="36" value="1977-05-07">7 May 1977</collectingDate>
</date>
;
<collectorName id="26E9B383A6160609FD17FDD3FC96E6A5" box="[748,884,634,660]" pageId="35" pageNumber="36">M. Sakaino</collectorName>
leg.”; “Sizumu NOMURA Bequest, 1981”; “♀”; “PARATYPE
<taxonomicName id="4C1CADD6A6160609FE51FD35FC61E687" authority="NOMURA (1977)" authorityName="Nomura" authorityYear="1977" box="[426,899,668,694]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FE51FD35FD7BE687" box="[426,665,668,694]" italics="true" pageId="35" pageNumber="36">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A6160609FD5FFD35FC61E687" author="Nomura S." box="[676,899,668,694]" pageId="35" pageNumber="36" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">NOMURA (1977)</bibRefCitation>
</taxonomicName>
”;
<collectionCode id="ED0D4E90A6160609FC65FD35FC0DE687" box="[926,1007,668,694]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="35" pageNumber="36">NSMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6160609FBF4FD35FB46E6E8" ID-GBIF-Occurrence="3114121379" collectingDate="1970-04-02" collectionCode="NSMT" collectorName="S. Cho" country="China" location="Taiwan" municipality="Kenting" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1A1DDCA6160609FBF4FD35FBA0E686" box="[1039,1090,668,695]" pageId="35" pageNumber="36" type="male">1 ♂</specimenCount>
;
<location id="8EC3808EA6160609FBAAFD35FB48E687" LSID="urn:lsid:plazi:treatment:03B56743A6140609FDBCF8E6FE9EE067:8EC3808EA6160609FBAAFD35FB48E687" box="[1105,1194,668,694]" country="China" municipality="Kenting" name="Taiwan" pageId="35" pageNumber="36">Taiwan</location>
,
<collectingMunicipality id="6BC74C2FA6160609FB42FD35FAF9E687" box="[1209,1307,668,694]" pageId="35" pageNumber="36">Kenting</collectingMunicipality>
;
<date id="FFA2F095A6160609FAD0FD35FF19E6E8" pageId="35" pageNumber="36" value="1970-04-02">
<collectingDate id="EFE6097DA6160609FAD0FD35FF19E6E8" pageId="35" pageNumber="36" value="1970-04-02">2 Apr. 1970</collectingDate>
</date>
;
<collectorName id="26E9B383A6160609FEFEFD16FEB6E6E8" box="[261,340,703,729]" pageId="35" pageNumber="36">S. Cho</collectorName>
leg.; “♂”; “PARATYPE
<taxonomicName id="4C1CADD6A6160609FD8FFD16FBDFE6E8" authority="NOMURA (1977)" authorityName="Nomura" authorityYear="1977" box="[628,1085,703,729]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FD8FFD16FCBCE6E8" box="[628,862,703,729]" italics="true" pageId="35" pageNumber="36">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A6160609FC9FFD16FBDFE6E8" author="Nomura S." box="[868,1085,703,729]" pageId="35" pageNumber="36" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">NOMURA (1977)</bibRefCitation>
</taxonomicName>
”;
<collectionCode id="ED0D4E90A6160609FBA8FD16FB46E6E8" box="[1107,1188,703,729]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="35" pageNumber="36">NSMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6160609FB41FD16FD62E72E" ID-GBIF-Occurrence="3114121391" collectingDate="1973-05-19" collectionCode="BMNH" collectorName="K. Masumoto" country="China" location="Ken ding" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1A1DDCA6160609FB41FD16FB05E6EB" box="[1210,1255,703,730]" pageId="35" pageNumber="36" type="male">1 ♂</specimenCount>
; “
<location id="8EC3808EA6160609FB07FD16FA8EE6E8" LSID="urn:lsid:plazi:treatment:03B56743A6140609FDBCF8E6FE9EE067:8EC3808EA6160609FB07FD16FA8EE6E8" box="[1276,1388,703,729]" country="China" name="Ken ding" pageId="35" pageNumber="36">Ken ding</location>
” [Taiwan, Kenting];
<date id="FFA2F095A6160609FE53FD4BFDAEE6CD" box="[424,588,738,764]" pageId="35" pageNumber="36" value="1973-05-19">
<collectingDate id="EFE6097DA6160609FE53FD4BFDAEE6CD" box="[424,588,738,764]" pageId="35" pageNumber="36" value="1973-05-19">19 May 1973</collectingDate>
</date>
;
<collectorName id="26E9B383A6160609FDA1FD4BFD1FE6CD" box="[602,765,738,764]" pageId="35" pageNumber="36">K. Masumoto</collectorName>
leg.; “♂”; “B. n. 1985-385”; “PARATYPE
<taxonomicName id="4C1CADD6A6160609FAF5FD4BFDF3E72F" authority="NOMURA, 1977" authorityName="Nomura" authorityYear="1977" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FAF5FD4BFEDBE72F" italics="true" pageId="35" pageNumber="36">Miridiba kuraruana</emphasis>
<bibRefCitation id="EF8DABA4A6160609FEBBFCADFDF3E72F" author="Nomura S." box="[320,529,772,798]" pageId="35" pageNumber="36" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">NOMURA, 1977</bibRefCitation>
</taxonomicName>
”;
<collectionCode id="ED0D4E90A6160609FDD3FCACFD62E72E" box="[552,640,773,799]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="35" pageNumber="36">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B74DC08A6160609FD62FCADFAD6E72E" ID-GBIF-Occurrence="3114121315" box="[665,1332,772,799]" collectingDate="1973-05-19" collectionCode="BMNH" collectorName="K. Masumoto" country="China" location="Ken" pageId="35" pageNumber="36" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1A1DDCA6160609FD62FCADFD24E72E" box="[665,710,772,799]" pageId="35" pageNumber="36" type="female">1 ♀</specimenCount>
; same collection data as for preceding; “♀”;
<collectionCode id="ED0D4E90A6160609FB23FCACFAD6E72E" box="[1240,1332,773,799]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="35" pageNumber="36">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6160609FF46FC97FBD7E7DF" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BA3D655A6160609FF46FC97FECDE769" blockId="35.[189,303,830,856]" box="[189,303,830,856]" pageId="35" pageNumber="36">
<heading id="D0EB6139A6160609FF46FC97FECDE769" bold="true" box="[189,303,830,856]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B9680A47A6160609FF46FC97FECDE769" bold="true" box="[189,303,830,856]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6160609FF46FCC5FBD7E7DF" blockId="35.[189,1399,876,1006]" pageId="35" pageNumber="36">
<bibRefCitation id="EF8DABA4A6160609FF46FCC5FE95E7B7" author="Nomura S." box="[189,375,876,902]" pageId="35" pageNumber="36" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura (1977)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A6160609FE05FCC5FD0CE7B7" authorityName="Nomura" authorityYear="1977" box="[510,750,876,902]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FE05FCC5FD0CE7B7" box="[510,750,876,902]" italics="true" pageId="35" pageNumber="36">Miridiba kuraruana</emphasis>
</taxonomicName>
based on a male
<typeStatus id="54A768F7A6160609FC35FCC5FBD7E7B7" box="[974,1077,876,902]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
and thirty-three
<typeStatus id="54A768F7A6160609FAFDFCC5FA94E7B7" box="[1286,1398,876,902]" pageId="35" pageNumber="36" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D1A1DDCA6160609FF3DFC27FED7E799" box="[198,309,910,936]" pageId="35" pageNumber="36" type="male">18 males</specimenCount>
and
<specimenCount id="9D1A1DDCA6160609FE81FC27FDE4E798" box="[378,518,910,937]" pageId="35" pageNumber="36" type="female">15 females</specimenCount>
). After studying the
<typeStatus id="54A768F7A6160609FCF0FC27FC90E799" box="[779,882,910,936]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D1A1DDCA6160609FC4DFC27FB86E798" box="[950,1124,910,937]" pageId="35" pageNumber="36" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
of
<taxonomicName id="4C1CADD6A6160609FB6CFC26FAA2E799" authorityName="Nomura" authorityYear="1977" box="[1175,1344,910,936]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FB6CFC26FAA2E799" box="[1175,1344,910,936]" italics="true" pageId="35" pageNumber="36">M. kuraruana</emphasis>
</taxonomicName>
, we conclude that
<taxonomicName id="4C1CADD6A6160609FE9BFC1BFDE1E7FA" authorityName="Nomura" authorityYear="1977" box="[352,515,945,971]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuraruana">
<emphasis id="B9680A47A6160609FE9BFC1BFDE1E7FA" box="[352,515,945,971]" italics="true" pageId="35" pageNumber="36">M. kuraruana</emphasis>
</taxonomicName>
shows features of male and female genitalia that characterize morphotype I “
<taxonomicName id="4C1CADD6A6160609FF31FC7DFEBFE7DF" box="[202,349,980,1006]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6160609FF31FC7DFEBFE7DF" box="[202,349,980,1006]" italics="true" pageId="35" pageNumber="36">Trichophora</emphasis>
</taxonomicName>
”. Hence, this species is included in this genital morphotype I.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6160609FF46FBA4FE9EE067" box="[189,380,1037,1110]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BA3D655A6160609FF46FBA4FEB4E016" blockId="35.[189,342,1037,1063]" box="[189,342,1037,1063]" pageId="35" pageNumber="36">
<heading id="D0EB6139A6160609FF46FBA4FEB4E016" bold="true" box="[189,342,1037,1063]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B9680A47A6160609FF46FBA4FEB4E016" bold="true" box="[189,342,1037,1063]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6160609FF46FB92FE9EE067" blockId="35.[189,380,1083,1110]" box="[189,380,1083,1110]" pageId="35" pageNumber="36">
<collectingCountry id="F30B96C5A6160609FF46FB92FEE1E064" box="[189,259,1083,1109]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
(
<collectingRegion id="49D818B7A6160609FEE8FB92FE95E067" box="[275,375,1083,1110]" country="China" name="Taiwan)" pageId="35" pageNumber="36">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>