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<document id="146623006072A690C5778CEA6B2CB06D" ID-DOI="10.5852/ejt.2020.615" ID-GBIF-Dataset="b2465730-6237-4a14-a98d-7fd2d1096cb0" ID-Zenodo-Dep="3711000" ID-ZooBank="637FC87F-13B5-4B32-BC52-11A9B30ECF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584210958022" checkinUser="plazi" docAuthor="Galea, Horia R." docDate="2020" docId="03B520481A5EFFAB9551716F59DF809B" docLanguage="en" docName="ejt-615_galea.pdf.imf" docOrigin="European Journal of Taxonomy 615" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cladocarpus asymmetricus Galea 2020, sp. nov." docType="treatment" docUuid="765A957F-FEF6-47A6-B09A-314F31F0440E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="9" masterDocId="FF8C58301A5AFFA3971777095A2A8450" masterDocTitle="Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program" masterLastPageNumber="47" masterPageNumber="1" pageNumber="5" updateTime="1698728323888" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="3445A5DF347491A2CC1D95D8C91E3AF3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FBE13C94EA2773E395BD194F0609F7B9">
<mods:title id="7240857543371656574E91D50B4886D1">Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program</mods:title>
</mods:titleInfo>
<mods:name id="463D73F26ED83A012E6A643169A3A00E" type="personal">
<mods:role id="167E76CCD964CAA7955A9FF24A626D36">
<mods:roleTerm id="979720F94EEC0A350F03DF3F3D5FB83E">Author</mods:roleTerm>
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<mods:namePart id="199610090F182F9D34858CF5EBE52158">Galea, Horia R.</mods:namePart>
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<mods:title id="833D31FD0DF840512C8D5F9E19051C5C">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="8EBFFAB75D921F84F5C56F53B6573C63">
<mods:date id="0775ABDB3FA956C0DB6B0D002AD57621">2020</mods:date>
<mods:detail id="76D16CDA56378F5D5A1ADD031A485895" type="pubDate">
<mods:number id="B3FB6A1DB2054707DD5C9FF1C163B10A">2020-03-13</mods:number>
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<mods:number id="625638583FBD083C55F0DB06F93DA7A8">615</mods:number>
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<mods:start id="BFFF95566B3CE29673CBAB7C01ECA522">1</mods:start>
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<mods:classification id="4ACB89C702F9A72408571A50239338BA">journal article</mods:classification>
<mods:identifier id="FD2862DCA78C54FC102729F700493082" type="DOI">10.5852/ejt.2020.615</mods:identifier>
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<mods:identifier id="33D32CBE7BEF7678BA400CC91C209E3A" type="Zenodo-Dep">3711000</mods:identifier>
<mods:identifier id="3B1B59E1FCC08228C56FF45CB1552B03" type="ZooBank">637FC87F-13B5-4B32-BC52-11A9B30ECF1D</mods:identifier>
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<treatment id="03B520481A5EFFAB9551716F59DF809B" ID-DOI="http://doi.org/10.5281/zenodo.3718327" ID-GBIF-Taxon="162505290" ID-Zenodo-Dep="3718327" LSID="urn:lsid:zoobank.org:act:765A957F-FEF6-47A6-B09A-314F31F0440E" httpUri="http://treatment.plazi.org/id/03B520481A5EFFAB9551716F59DF809B" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<subSubSection id="C306C2D51A5EFFA79551716F5E9582F5" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BA3915E1A5EFFA79551716F59C482D1" blockId="4.[373,1215,1638,1736]" box="[582,1006,1638,1665]" pageId="4" pageNumber="5">
<heading id="D0EB26321A5EFFA79551716F59C482D1" box="[582,1006,1638,1665]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C1CEADD1A5EFFA79551716F59A682D0" ID-CoL="847LQ" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[582,908,1638,1665]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B9684D4C1A5EFFA79551716F59A682D0" bold="true" box="[582,908,1638,1665]" italics="true" pageId="4" pageNumber="5">Cladocarpus asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A5EFFA79483716E59C482D1" box="[916,1006,1639,1665]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA3915E1A5EFFA7966271835E9582F5" blockId="4.[373,1215,1638,1736]" box="[373,1215,1674,1701]" pageId="4" pageNumber="5">
<uri id="FF8D9D5C1A5EFFA7966271835E9582F5" box="[373,1215,1674,1701]" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
<uuid id="FFBAAB8B1A5EFFA7958571835E9582F5" box="[658,1215,1674,1701]" pageId="4" pageNumber="5">765A957F-FEF6-47A6-B09A-314F31F0440E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A5EFFA7958E71A759B18298" box="[665,923,1710,1736]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BA3915E1A5EFFA7958E71A759B18298" blockId="4.[373,1215,1638,1736]" box="[665,923,1710,1736]" pageId="4" pageNumber="5">
<figureCitation id="13278DDB1A5EFFA7958E71A758F78298" box="[665,733,1710,1736]" captionStart="Fig" captionStartId="2.[189,228,1862,1886]" captionTargetBox="[189,1399,532,1814]" captionTargetId="figure@2.[189,1399,532,1814]" captionTargetPageId="2" captionText="Fig. 1. A–B. Aglaophenia howensis Briggs, 1918, fertile cormoid (A) and corbula (B), both from sample MNHN-IK-2015-544. — C. Aglaophenia sinuosa Bale, 1888, cormoid from sample MNHN- IK-2015-545. — D–E. Cladocarpus asymmetricus sp. nov., fertile cormoid (D) and phylactocarp (E) from sample MNHN-IK-2015-530 (holotype). — F. Cladocarpus keiensis Schuchert, 2003, colony from sample MNHN-IK-2015-534. Scale bars: F = 2 cm; A, C–D = 1 cm; E = 1 mm; B = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3711002" httpUri="https://zenodo.org/record/3711002/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
D–E, 4;
<tableCitation id="C69EA4E51A5EFFA7945371A759B18298" box="[836,923,1710,1736]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionText="Table 1. Nominal species of Cladocarpus Allman, 1874 with (occasionally asymmetrical) hydrothecae provided with sigmoid (or nearly so) lateral ridges, similar to those of C. asymmetricus sp. nov., and their morphological differences with the new species." pageId="4" pageNumber="5">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A5EFFA797AA71E658B283BB" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BA3915E1A5EFFA797AA71E65B1C8359" blockId="4.[189,310,1775,1801]" box="[189,310,1775,1801]" pageId="4" pageNumber="5">
<heading id="D0EB26321A5EFFA797AA71E65B1C8359" bold="true" box="[189,310,1775,1801]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B9684D4C1A5EFFA797AA71E65B1C8359" bold="true" box="[189,310,1775,1801]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A5EFFA797AA701758B283BB" blockId="4.[189,1399,1822,2027]" pageId="4" pageNumber="5">
<taxonomicName id="4C1CEADD1A5EFFA797AA70175B7F8368" authorityName="Allman" authorityYear="1874" box="[189,341,1822,1848]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="4" pageNumber="5" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A5EFFA797AA70175B7F8368" box="[189,341,1822,1848]" italics="true" pageId="4" pageNumber="5">Cladocarpus</emphasis>
</taxonomicName>
with lightly fascicled, unbranched stems divided into fairly long internodes, each with 2–3 frontal nematothecae, a mamelon and an axillar nematotheca. Cladia close to one another. Cormidia with up to 7 intranodal, adaxial septa. Hydrothecae elongated, asymmetrical in frontal view, with two lateral transverse ridges projecting into the lumen, rim with 9 rounded cusps. Phylactocarps given off from first cormidium, gonothecae concrescent with the rachis basally, crescent-shaped, with three
<typeStatus id="54A72FFC1A5EFFA7922E70A75F5D8398" box="[1337,1399,1966,1992]" pageId="4" pageNumber="5">types</typeStatus>
of nematothecae on both body and apex.
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A5FFFA697AA76025EED8665" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BA3915E1A5FFFA697AA76025B6E8575" blockId="5.[189,324,267,293]" box="[189,324,267,293]" pageId="5" pageNumber="6">
<heading id="D0EB26321A5FFFA697AA76025B6E8575" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9684D4C1A5FFFA697AA76025B6E8575" bold="true" box="[189,324,267,293]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A5FFFA697AA76325BD38529" blockId="5.[189,1399,315,444]" pageId="5" pageNumber="6">
From the Latin ‘
<emphasis id="B9684D4C1A5FFFA696917632582F8505" box="[390,517,315,341]" italics="true" pageId="5" pageNumber="6">ăsymmĕtĕr</emphasis>
,
<emphasis id="B9684D4C1A5FFFA69501763258688504" box="[534,578,315,340]" italics="true" pageId="5" pageNumber="6">-tra</emphasis>
,
<emphasis id="B9684D4C1A5FFFA69544763258BF8504" box="[595,661,315,340]" italics="true" pageId="5" pageNumber="6">-trum</emphasis>
’, meaning ‘asymmetrical’, with reference to the shape of the hydrotheca in frontal view.
</paragraph>
<paragraph id="8BA3915E1A5FFFA697AA76AB5B8085EC" blockId="5.[189,1399,315,444]" box="[189,426,418,444]" pageId="5" pageNumber="6">
<emphasis id="B9684D4C1A5FFFA697AA76AB5B8085EC" bold="true" box="[189,426,418,444]" pageId="5" pageNumber="6">Material examined</emphasis>
</paragraph>
<paragraph id="8BA3915E1A5FFFA697FD76DB5B7685BC" blockId="5.[189,1398,466,565]" box="[234,348,466,492]" pageId="5" pageNumber="6">
<emphasis id="B9684D4C1A5FFFA697FD76DB5B7685BC" bold="true" box="[234,348,466,492]" pageId="5" pageNumber="6">
<typeStatus id="54A72FFC1A5FFFA697FD76DB5B7685BC" box="[234,348,466,492]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA3915E1A5FFFA697AA76FF5EED8665" blockId="5.[189,1398,466,565]" pageId="5" pageNumber="6">
<materialsCitation id="3B749B031A5FFFA697AA76FF5EED8665" ID-GBIF-Occurrence="2575926600" collectingDate="2017-09-09" collectionCode="MNHN" collectorName="KANADEEP" country="New Caledonia" elevation="295" latitude="-19.7" location="stn DW 4979" longLatPrecision="1271" longitude="158.61667" pageId="5" pageNumber="6" specimenCode="MNHN-IK-2015-530." specimenCount="1" typeStatus="holotype">
PACIFIC OCEAN •
<specimenCount id="9D1A5AD71A5FFFA696BF76FF58788640" box="[424,594,502,528]" pageId="5" pageNumber="6">1 single plume</specimenCount>
, ca
<quantity id="4CE43CBB1A5FFFA6956E76FF58ED8640" box="[633,711,502,529]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="6" unit="cm" value="8.5">8.5 cm</quantity>
high, hydrorhiza missing; off
<collectingCountry id="F30BD1CE1A5FFFA6930A76FF5EF98640" box="[1053,1235,502,528]" name="New Caledonia" pageId="5" pageNumber="6">New Caledonia</collectingCountry>
,
<location id="8EC3C7851A5FFFA693CB76FF5F588640" LSID="urn:lsid:plazi:treatment:03B520481A5EFFAB9551716F59DF809B:8EC3C7851A5FFFA693CB76FF5F588640" box="[1244,1394,502,528]" country="New Caledonia" latitude="-19.7" longLatPrecision="1271" longitude="158.61667" name="stn DW 4979" pageId="5" pageNumber="6">stn DW4979</location>
;
<geoCoordinate id="EE28F7991A5FFFA697AA75105B098664" box="[189,291,537,565]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-19.7">19°42ʹ S</geoCoordinate>
,
<geoCoordinate id="EE28F7991A5FFFA6962775105B828665" box="[304,424,537,565]" degrees="158" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="158.61667">158°37ʹ E</geoCoordinate>
; 315–
<quantity id="4CE43CBB1A5FFFA696F8751358168665" box="[495,572,538,565]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="5" pageNumber="6" unit="m" value="295.0">
<elevation id="0031766D1A5FFFA696F8751358168665" box="[495,572,538,565]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="5" pageNumber="6" unit="m" value="295.0">295 m</elevation>
</quantity>
;
<date id="FFA2B79E1A5FFFA6955E751358FD8664" box="[585,727,538,564]" pageId="5" pageNumber="6" value="2017-09-09">
<collectingDate id="EFE64E761A5FFFA6955E751358FD8664" box="[585,727,538,564]" pageId="5" pageNumber="6" value="2017-09-09">9 Sep. 2017</collectingDate>
</date>
;
<collectorName id="26E9F4881A5FFFA695F4751259AA8665" box="[739,896,539,565]" pageId="5" pageNumber="6">KANADEEP</collectorName>
leg.;
<specimenCode id="DBBA39251A5FFFA694D575135EED8665" box="[962,1223,538,565]" pageId="5" pageNumber="6">MNHN-IK-2015-530.</specimenCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A5FFFA597AA755458628518" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8BA3915E1A5FFFA697AA75545B648627" blockId="5.[189,334,605,631]" box="[189,334,605,631]" pageId="5" pageNumber="6">
<heading id="D0EB26321A5FFFA697AA75545B648627" bold="true" box="[189,334,605,631]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9684D4C1A5FFFA697AA75545B648627" bold="true" box="[189,334,605,631]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A5FFFA597AA758458628518" blockId="5.[189,1399,653,2027]" lastBlockId="6.[189,1399,267,329]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Colony erect, at least
<quantity id="4CE43CBB1A5FFFA696A47584582B86F7" box="[435,513,653,680]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="6" unit="cm" value="8.5">8.5 cm</quantity>
high, hydrorhiza missing. Stem unbranched, fairly fascicled, accessory tubes forming a beam around the main tube; no deeply-cut, proximal, oblique nodes; elsewhere division into internodes indistinct, except for distalmost, monosiphonic part, with thinner perisarc, where transverse nodes (
<quantity id="4CE43CBB1A5FFFA6960775F35B8C8745" box="[272,422,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="5" pageNumber="6" unit="mm" value="172.5" valueMax="180.0" valueMin="165.0">165–180 µm</quantity>
wide) delimit rather long (
<quantity id="4CE43CBB1A5FFFA695CE75F359558745" box="[729,895,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="9.25" metricValueMax="10.3" metricValueMin="8.2" pageId="5" pageNumber="6" unit="mm" value="925.0" valueMax="1030.0" valueMin="820.0">820–1030 µm</quantity>
) internodes composed of 2–3 (occasionally 4) proximal nematothecae in a row, a lateral cladial apophysis with its associated conical nematotheca (with rounded, apical aperture), and an axillar nematotheca rather lateral to the apophysis; dorsal axillar nematotheca absent; all stem nematothecae sac-shaped, adnate for ⅔ of their length, with apical, guttershaped aperture facing adaxially; each accessory tube of the stem with a row of saccate, deeply immersed nematothecae, with only their ovoid apertures slightly protruding from the perisarc. Cladial apophyses alternate along the stem, ending distally in a deeply oblique node; cladia close to one another, up to
<quantity id="4CE43CBB1A5FFFA697AA74F05B248044" box="[189,270,1017,1044]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="6" unit="cm" value="1.9">1.9 cm</quantity>
long, divided into rather long internodes (
<quantity id="4CE43CBB1A5FFFA6941474F059908044" box="[771,954,1017,1044]" metricMagnitude="0" metricUnit="m" metricValue="1.0574999999999999" metricValueMax="1.09" metricValueMin="1.025" pageId="5" pageNumber="6" unit="mm" value="1057.5" valueMax="1090.0" valueMin="1025.0">
<date id="FFA2B79E1A5FFFA6941474F059A28044" box="[771,904,1017,1044]" pageId="5" pageNumber="6" value="1025" valueMax="1090">1025–1090</date>
µm
</quantity>
) by means of slight, transverse (
<quantity id="4CE43CBB1A5FFFA6922C74F05B338068" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" pageId="5" pageNumber="6" unit="mm" value="117.5" valueMax="120.0" valueMin="115.0">115– 120 µm</quantity>
wide) constrictions of the perisarc, each accommodating a hydrotheca and its 3 nematothecae: one mesial and a pair of laterals flanking the aperture. Hydrotheca elongated,
<quantity id="4CE43CBB1A5FFFA69328734B5EF9800C" box="[1087,1235,1090,1117]" metricMagnitude="-1" metricUnit="m" metricValue="9.225" metricValueMax="9.5" metricValueMin="8.95" pageId="5" pageNumber="6" unit="mm" value="922.5" valueMax="950.0" valueMin="895.0">895–950 µm</quantity>
deep, slightly compressed laterally below the rim, with straight adaxial and slightly curved abaxial walls, with an intrathecal septum given off from the lower ¼ of the adaxial wall, and projecting for a short distance into the lumen; in frontal view, lower part of hydrotheca asymmetrical, due to the presence of two conspicuous, semicircular, intrathecal plates given off at two different levels from the lateral walls; free edges slightly overlapping in the middle of the lumen; in a lateral view of the hydrotheca, the attachment sites of these septa are distinctly sigmoid. Mesial nematotheca long, tubular, adnate for about ¾ its length, leaving distally a short (
<quantity id="4CE43CBB1A5FFFA696CC72485873810B" box="[475,601,1345,1372]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="5" pageNumber="6" unit="mm" value="92.5" valueMax="95.0" valueMin="90.0">90–95 µm</quantity>
), gutter-shaped aperture; lateral nematothecae short (
<quantity id="4CE43CBB1A5FFFA693DA72485F42810B" box="[1229,1384,1345,1372]" metricMagnitude="-1" metricUnit="m" metricValue="1.975" metricValueMax="2.0" metricValueMin="1.95" pageId="5" pageNumber="6" unit="mm" value="197.5" valueMax="200.0" valueMin="195.0">195–200 µm</quantity>
), tubular, tapering towards the distal aperture, the latter with deeply scooped margin on adaxial side. Hydrothecal aperture
<quantity id="4CE43CBB1A5FFFA696D77283587281F4" box="[448,600,1418,1445]" metricMagnitude="-1" metricUnit="m" metricValue="3.025" metricValueMax="3.05" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="302.5" valueMax="305.0" valueMin="300.0">300–305 µm</quantity>
wide, rim with 9 cusps: one median, abaxial with squared outline, and 4 pairs of laterals, the latter triangular in shape, with rounded tips, becoming gradually lower and wider towards the adaxial side of the theca; up to 7 intranodal septa, all given off from the adaxial wall of the hydrothecae and projecting for a relatively short distance into the lumen of the internode; of these, 2 cusps are given off from the adaxial thecal wall below the intrahydrothecal septum, while up to 5 others from above it. Phylactocarp given off laterally from first cormidium, borne on distinct, athecate apophysis with distal, oblique node; phylactocarp up to
<quantity id="4CE43CBB1A5FFFA694DA716D5E3E822F" box="[973,1044,1636,1663]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="6" unit="mm" value="6.0">6 mm</quantity>
long, composed of a slightly geniculate rachis divided into up to 20 short (
<quantity id="4CE43CBB1A5FFFA695F2718059AA82F3" box="[741,896,1673,1700]" metricMagnitude="-1" metricUnit="m" metricValue="2.775" metricValueMax="3.05" metricValueMin="2.5" pageId="5" pageNumber="6" unit="mm" value="277.5" valueMax="305.0" valueMin="250.0">250–305 µm</quantity>
) internodes by means of slightly-marked, transverse nodes; each internode with a proximal nematotheca, a basally concrescent gonotheca, and a second nematotheca lateral to its insertion (both nematothecae elongated, with scooped rims adaxially); the proximalmost internode has an extra proximal nematotheca. Gonothecae crescent-shaped in lateral view, ca 850 ×
<quantity id="4CE43CBB1A5FFFA6969770135BC88365" box="[384,482,1818,1845]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="mm" value="250.0">250 µm</quantity>
, those with curvature on one side alternating with those curved oppositely from the other side; circular in transverse section, tapering gently below and gradually above to form a pointed apex bearing a nematotheca with scooped rim; two dissimilar nematothecae, placed at nearly the same level on the adaxial side of the thecal wall, at about ⅓ of the length above from the insertion of the gonotheca on the rachis: one nematotheca is conical, with thick walls and has a small, slightly scooped aperture, while the other is barrel-shaped, with thin walls and has a broad, circular aperture with entire margin; upper third of the adaxial gonothecal wall with 1 (occasionally 2) sac-shaped nematotheca(e) with scooped rim on adaxial side.
</paragraph>
</subSubSection>
<caption id="DF63C1D61A5CFFA597AA704B5F4E83B9" ID-DOI="http://doi.org/10.5281/zenodo.3711008" ID-Zenodo-Dep="3711008" httpUri="https://zenodo.org/record/3711008/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1858,1884]" targetBox="[189,1396,493,1809]" targetPageId="6">
<paragraph id="8BA3915E1A5CFFA597AA704B5F4E83B9" blockId="6.[189,1399,1858,2025]" pageId="6" pageNumber="7">
<emphasis id="B9684D4C1A5CFFA597AA704B5B24830C" bold="true" box="[189,270,1858,1884]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="4C1CEADD1A5CFFA5960E704B5877830C" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[281,605,1858,1884]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="6" pageNumber="7" phylum="Cnidaria" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B9684D4C1A5CFFA5960E704B5877830C" box="[281,605,1858,1884]" italics="true" pageId="6" pageNumber="7">Cladocarpus asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A5CFFA5957E704B58E0830C" box="[617,714,1858,1884]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, from sample MNHN-IK-2015-530.
<emphasis id="B9684D4C1A5CFFA5939C704B5E8A830C" bold="true" box="[1163,1184,1858,1884]" pageId="6" pageNumber="7">A</emphasis>
. Portion of stem.
<emphasis id="B9684D4C1A5CFFA597AA706C5ADC832F" bold="true" box="[189,246,1893,1919]" pageId="6" pageNumber="7">B–H</emphasis>
. Hydrotheca in frontal (B), lateral (C) and apical (D) views; details of its proximal (E) and distal (F) parts seen laterally, and of the rim in frontal view (G); intrathecal septum in an apical view of the hydrotheca (H).
<emphasis id="B9684D4C1A5CFFA5966A70A55B8C8396" bold="true" box="[381,422,1964,1990]" pageId="6" pageNumber="7">I–J</emphasis>
. Portion of phylactocarp with gonothecae (I) and details of the nematothecae borne by the gonothecae (J). Scale bars: A = 500 µm; B–C, E–F, I = 200 µm; D, G–H = 100 µm; J = 50 µm.
</paragraph>
</caption>
<caption id="DF63C1D61A5DFFA497AA7605591B853C" pageId="7" pageNumber="8">
<paragraph id="8BA3915E1A5DFFA497AA7605591B853C" blockId="7.[189,1398,267,364]" pageId="7" pageNumber="8">
<emphasis id="B9684D4C1A5DFFA497AA76055B0A8575" bold="true" box="[189,288,267,294]" pageId="7" pageNumber="8">Table 1.</emphasis>
Nominal species of
<taxonomicName id="4C1CEADD1A5DFFA495057602597B8575" authority="Allman, 1874" authorityName="Allman" authorityYear="1874" box="[530,849,267,293]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A5DFFA49505760258808575" box="[530,682,267,293]" italics="true" pageId="7" pageNumber="8">Cladocarpus</emphasis>
Allman, 1874
</taxonomicName>
with (occasionally asymmetrical) hydrothecae provided with sigmoid (or nearly so) lateral ridges, similar to those of
<taxonomicName id="4C1CEADD1A5DFFA4930776265EFF8518" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[1040,1237,303,329]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B9684D4C1A5DFFA4930776265EFF8518" box="[1040,1237,303,329]" italics="true" pageId="7" pageNumber="8">C. asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A5DFFA493C876265F148519" box="[1247,1342,303,329]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, and their morphological differences with the new species.
</paragraph>
</caption>
<paragraph id="8BA3915E1A5DFFA497AA76B35E2283D1" pageId="7" pageNumber="8">
<table id="F91C63FE1A5D005C97AA76A15F5C83D1" box="[189,1398,424,1921]" gridcols="2" gridrows="5" pageId="7" pageNumber="8">
<tr id="352C931C1A5D005C97AA76A15F5C85B6" box="[189,1398,424,486]" gridrow="0" pageId="7" pageNumber="8">
<th id="76FDFA601A5D005C97AA76A15BAC85B6" box="[189,390,424,486]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B9684D4C1A5DFFA497AA76B35BAC8584" bold="true" box="[189,390,442,468]" pageId="7" pageNumber="8">Nominal species</emphasis>
</th>
<th id="76FDFA601A5D005C96BA76A15F5C85B6" box="[429,1398,424,486]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="B9684D4C1A5DFFA496BA76A1586B85B6" bold="true" pageId="7" pageNumber="8">Brief description, with emphasis on the distinctive features, and geographical distribution</emphasis>
</th>
</tr>
<tr id="352C931C1A5D005C97AA76F75F5C87EF" box="[189,1398,510,959]" gridrow="1" pageId="7" pageNumber="8">
<td id="76FDFA601A5D005C97AA76F75BAC87EF" box="[189,390,510,959]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">
<taxonomicName id="4C1CEADD1A5DFFA497AA75A75B7F8698" authorityName="Allman" authorityYear="1874" box="[189,341,686,712]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A5DFFA497AA75A75B7F8698" box="[189,341,686,712]" italics="true" pageId="7" pageNumber="8">Cladocarpus</emphasis>
</taxonomicName>
<emphasis id="B9684D4C1A5DFFA497AA75DB5B0586BC" box="[189,303,722,748]" italics="true" pageId="7" pageNumber="8">elongatus</emphasis>
Bedot, 1921
</td>
<td id="76FDFA601A5D005C96BA76F75F5C87EF" box="[429,1398,510,959]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">
Colonies rather tall (up to 15 cm high), occasionally branched, fascicled; stem internodes with one frontal and a pair of axillar nematothecae, flanking the cladial apophyses, mamelon
<emphasis id="B9684D4C1A5DFFA495A7754C58D7860F" box="[688,765,581,607]" italics="true" pageId="7" pageNumber="8">absent</emphasis>
; cormidia with 7–8 intranodal septa, of which 2 are abaxial on proximal part; hydrothecae elongated, aperture
<emphasis id="B9684D4C1A5DFFA4937575615ED786D2" box="[1122,1277,616,642]" italics="true" pageId="7" pageNumber="8">slightly tilted</emphasis>
forwards, margin with
<emphasis id="B9684D4C1A5DFFA495597582590C86F5" box="[590,806,651,677]" italics="true" pageId="7" pageNumber="8">11 rounded cusps</emphasis>
, of which the median, abaxial one is the most developed; mesial nematotheca adnate for a
<emphasis id="B9684D4C1A5DFFA494AF75A65E048698" box="[952,1070,686,712]" italics="true" pageId="7" pageNumber="8">very short</emphasis>
portion of the basal abaxial wall of hydrotheca, laterals not overtopping the hydrothecal aperture; internal septum
<emphasis id="B9684D4C1A5DFFA496BA75FC582F875F" box="[429,517,757,783]" italics="true" pageId="7" pageNumber="8">oblique</emphasis>
rather than decidedly sigmoid, insertions on lateral walls of hydrotheca at different levels; phylactocarp borne on first cormidium,
<emphasis id="B9684D4C1A5DFFA4934374105E868762" box="[1108,1196,793,818]" italics="true" pageId="7" pageNumber="8">pinnate</emphasis>
, branchlets with successive pairs of opposite to subopposite nematothecae; gonothecae
<emphasis id="B9684D4C1A5DFFA4923A74325F448705" box="[1325,1390,827,853]" italics="true" pageId="7" pageNumber="8">ovoid</emphasis>
, borne
<emphasis id="B9684D4C1A5DFFA496EE745658B38728" box="[505,665,862,888]" italics="true" pageId="7" pageNumber="8">on apophyses</emphasis>
of branchlets, male with apical, rounded, operculate aperture
<emphasis id="B9684D4C1A5DFFA496BA748B582687CC" box="[429,524,898,924]" italics="true" pageId="7" pageNumber="8">inclined</emphasis>
to one side, female bigger, with broad, ovoid,
<emphasis id="B9684D4C1A5DFFA49336748B5F2287CC" box="[1057,1288,898,924]" italics="true" pageId="7" pageNumber="8">lateral, subterminal</emphasis>
aperture. Distribution: off the Atlantic coast of Morocco (
<bibRefCitation id="EF8DECAF1A5DFFA494F774AC5E4787EF" author="Bedot M." box="[992,1133,933,959]" firstAuthor="Bedot" journalOrPublisher="Resultats des Campagnes scientifiques accomplies sur son Yacht par Albert Ier Prince souverain de Monaco" pageId="7" pageNumber="8" pagination="1 - 74" part="60" refId="ref21597" refString="Bedot M. 1921. Hydroides provenant des campagnes des Yachts Hirondelle et Princesse-Alice (1887 - 1912). Resultats des Campagnes scientifiques accomplies sur son Yacht par Albert Ier Prince souverain de Monaco 60: 1 - 74." title="Hydroides provenant des campagnes des Yachts Hirondelle et Princesse-Alice (1887 - 1912)" type="journal article" year="1921">Bedot 1921</bibRefCitation>
;
<bibRefCitation id="EF8DECAF1A5DFFA4936E74AC5F6C87EF" accessDate="2020-03-06" author="Ramil F. &amp; Vervoort W. &amp; Ansin Agis J." box="[1145,1350,933,959]" etAl="et al." firstAuthor="Ramil" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="7" pageNumber="8" pagination="1 - 42" part="322" publicationUrl="https://www.repository.naturalis.nl/document/149163" refId="ref23271" refString="Ramil F., Vervoort W. &amp; Ansin Agis J. 1998. Report on the Haleciidae and Plumularioidea (Cnidaria, Hydrozoa) collected by the French Seamount 1 Expedition. Zoologische Verhandelingen, Leiden 322: 1 - 42. Available from https: // www. repository. naturalis. nl / document / 149163 [accessed 6 Mar. 2020]." title="Report on the Haleciidae and Plumularioidea (Cnidaria, Hydrozoa) collected by the French Seamount 1 Expedition" type="journal article" year="1998">
Ramil
<emphasis id="B9684D4C1A5DFFA493DF74AF5F2987EF" box="[1224,1283,933,959]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
1998
</bibRefCitation>
).
</td>
</tr>
<tr id="352C931C1A5D005C97AA74DE5F5C8102" box="[189,1398,983,1362]" gridrow="2" pageId="7" pageNumber="8">
<td id="76FDFA601A5D005C97AA74DE5BAC8102" box="[189,390,983,1362]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<taxonomicName id="4C1CEADD1A5DFFA497AA735A5B748086" authority="Vervoort" authorityName="Vervoort" authorityYear="2003" class="Hydrozoa" family="Aglaopheniidae" genus="Carpocladus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="species" species="fertilis">
<emphasis id="B9684D4C1A5DFFA497AA735A5B2080C0" box="[189,341,1107,1168]" italics="true" pageId="7" pageNumber="8">Carpocladus fertilis</emphasis>
<bibRefCitation id="EF8DECAF1A5DFFA497AA73935B748086" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="7" pageNumber="8" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson, 2003</bibRefCitation>
</taxonomicName>
</td>
<td id="76FDFA601A5D005C96BA74DE5F5C8102" box="[429,1398,983,1362]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">
Colonies rather tall (ca 12 cm high), with unbranched, fascicled stems; internodes with 1
<emphasis id="B9684D4C1A5DFFA4951874F258748045" box="[527,606,1019,1045]" italics="true" pageId="7" pageNumber="8">frontal</emphasis>
and a fronto-axillar nematotheca, apophyses alternate,
<emphasis id="B9684D4C1A5DFFA4920A74F25F5F8045" box="[1309,1397,1019,1045]" italics="true" pageId="7" pageNumber="8">without</emphasis>
mamelons; cormidia with ca 7 intranodal septa, of which 3 are abaxial, below the hydrotheca, the remaining 4 on adaxial side of the hydrotheca; abaxial wall with
<emphasis id="B9684D4C1A5DFFA496F9736D58B4802E" box="[494,670,1124,1150]" italics="true" pageId="7" pageNumber="8">median carina</emphasis>
, forming a conspicuous
<emphasis id="B9684D4C1A5DFFA494C4736D5F2D802E" box="[979,1287,1124,1150]" italics="true" pageId="7" pageNumber="8">dagger-shaped projection</emphasis>
on front of median marginal cusp; aperture
<emphasis id="B9684D4C1A5DFFA49477738E59D480F1" box="[864,1022,1159,1185]" italics="true" pageId="7" pageNumber="8">slightly tilted</emphasis>
forwards, rim with 4 pairs of lateral cusps; a strong adaxial ledge projecting into hydrothecal lumen, prolonged upwards sigmoidally on lateral walls; mesial nematotheca covering basal third of abaxial hydrothecal wall, laterals just over-reaching margin; phylactocarps borne on first cormidia, each internode with a gonotheca and a
<emphasis id="B9684D4C1A5DFFA49350721D5EB8817E" box="[1095,1170,1300,1326]" italics="true" pageId="7" pageNumber="8">forked</emphasis>
costa comprising a
<emphasis id="B9684D4C1A5DFFA496BA723E581A8101" box="[429,560,1335,1361]" italics="true" pageId="7" pageNumber="8">hydrotheca</emphasis>
. Distribution: Tasman Sea (
<bibRefCitation id="EF8DECAF1A5DFFA4946072315E898102" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[887,1187,1336,1362]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="7" pageNumber="8" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson 2003</bibRefCitation>
).
</td>
</tr>
<tr id="352C931C1A5D005C97AA72635F5C822B" box="[189,1398,1386,1659]" gridrow="3" pageId="7" pageNumber="8">
<td id="76FDFA601A5D005C97AA72635BAC822B" box="[189,390,1386,1659]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">
<taxonomicName id="4C1CEADD1A5DFFA497AA72CB5B5B8273" authority="(Allman, 1877)" baseAuthorityName="Allman" baseAuthorityYear="1877" box="[189,369,1474,1571]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="species" species="sigma">
<emphasis id="B9684D4C1A5DFFA497AA72CB5B2981AF" box="[189,341,1474,1535]" italics="true" pageId="7" pageNumber="8">Cladocarpus sigma</emphasis>
(
<bibRefCitation id="EF8DECAF1A5DFFA497D271015B438273" author="Allman G. J." box="[197,361,1544,1571]" firstAuthor="Allman" journalOrPublisher="Memoirs of the Museum of Comparative Zoology at Harvard College" pageId="7" pageNumber="8" pagination="1 - 66" part="5" refId="ref21156" refString="Allman G. J. 1877. Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoology at Harvard College 5 (2): 1 - 66. https: // doi. org / 10.5962 / bhl. title. 10420" title="Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey" type="journal article" year="1877">Allman, 1877</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76FDFA601A5D005C96BA72635F5C822B" box="[429,1398,1386,1659]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">
Stems
<emphasis id="B9684D4C1A5DFFA496EA7263580E81D4" box="[509,548,1386,1412]" italics="true" pageId="7" pageNumber="8">tall</emphasis>
(ca 15 cm high), unbranched, fascicled (
<bibRefCitation id="EF8DECAF1A5DFFA4931F72635E8381D4" author="Allman G. J." box="[1032,1193,1386,1412]" firstAuthor="Allman" journalOrPublisher="Memoirs of the Museum of Comparative Zoology at Harvard College" pageId="7" pageNumber="8" pagination="1 - 66" part="5" refId="ref21156" refString="Allman G. J. 1877. Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoology at Harvard College 5 (2): 1 - 66. https: // doi. org / 10.5962 / bhl. title. 10420" title="Report on the Hydroida collected during the exploration of the Gulf Stream by L. F. de Pourtales, Assistant United States Coast Survey" type="journal article" year="1877">Allman 1877</bibRefCitation>
); internodes with 1 frontal nematotheca and 2 axillar nematothecae, conical nematotheca
<emphasis id="B9684D4C1A5DFFA4923672845F4481F7" box="[1313,1390,1421,1447]" italics="true" pageId="7" pageNumber="8">absent</emphasis>
; cormidia with 5–7 intranodal septa; hydrotheca
<emphasis id="B9684D4C1A5DFFA4931D72B95EBB819A" box="[1034,1169,1456,1482]" italics="true" pageId="7" pageNumber="8">cup-shaped</emphasis>
, with two lateral, intrathecal, sigmoidal ridges arising at different levels; margin with median abaxial cusps and
<emphasis id="B9684D4C1A5DFFA4953872FE58AF8241" box="[559,645,1527,1553]" italics="true" pageId="7" pageNumber="8">5 pairs</emphasis>
of lateral cusps; mesial nematotheca
<emphasis id="B9684D4C1A5DFFA4934472FE5F5F8241" box="[1107,1397,1527,1553]" italics="true" pageId="7" pageNumber="8">short, barely surpassing</emphasis>
hydrothecal base; lateral nematothecae
<emphasis id="B9684D4C1A5DFFA4946E7113599F8264" box="[889,949,1562,1588]" italics="true" pageId="7" pageNumber="8">short</emphasis>
, not always reaching hydrothecal rim; phylactocarps
<emphasis id="B9684D4C1A5DFFA4954B7137589E8207" box="[604,692,1598,1623]" italics="true" pageId="7" pageNumber="8">pinnate</emphasis>
, an
<emphasis id="B9684D4C1A5DFFA495F87137598C8207" box="[751,934,1597,1623]" italics="true" pageId="7" pageNumber="8">ovate gonothec</emphasis>
a at base of each costa (
<bibRefCitation id="EF8DECAF1A5DFFA493C171345F428208" author="Bogle M. A." box="[1238,1384,1597,1624]" firstAuthor="Bogle" journalOrPublisher="MSc Thesis, University of Miami, Coral Gables, USA" pageId="7" pageNumber="8" refId="ref21715" refString="Bogle M. A. 1975. A Review and Preliminary Revision of the Aglaopheniinae (Hydroida: Plumulariidae) of the Tropical Western Atlantic. MSc Thesis, University of Miami, Coral Gables, USA." title="A Review and Preliminary Revision of the Aglaopheniinae (Hydroida: Plumulariidae) of the Tropical Western Atlantic" type="book" year="1975">Bogle 1975</bibRefCitation>
). Distribution: tropical and subtropical western Atlantic (
<bibRefCitation id="EF8DECAF1A5DFFA4932271685ECE822B" accessDate="2020-03-06" author="Vervoort W." box="[1077,1252,1633,1659]" firstAuthor="Vervoort" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="7" pageNumber="8" pagination="1 - 247" part="120" publicationUrl="https://www.repository.naturalis.nl/document/149147" refId="ref24013" refString="Vervoort W. 1972. Hydroids from the Theta, Vema and Yelcho cruises of the Lamont-Doherty Gological Observatory. Zoologische Verhandelingen, Leiden 120: 1 - 247. Available from https: // www. repository. naturalis. nl / document / 149147 [accessed 6 Mar. 2020]." title="Hydroids from the Theta, Vema and Yelcho cruises of the Lamont-Doherty Gological Observatory" type="journal article" year="1972">Vervoort 1972</bibRefCitation>
).
</td>
</tr>
<tr id="352C931C1A5D005C97AA719A5F5C83D1" box="[189,1398,1683,1921]" gridrow="4" pageId="7" pageNumber="8">
<td id="76FDFA601A5D005C97AA719A5BAC83D1" box="[189,390,1683,1921]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">
<taxonomicName id="4C1CEADD1A5DFFA497AA71D05B7F82A3" authorityName="Allman" authorityYear="1874" box="[189,341,1753,1779]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="7" pageNumber="8" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A5DFFA497AA71D05B7F82A3" box="[189,341,1753,1779]" italics="true" pageId="7" pageNumber="8">Cladocarpus</emphasis>
</taxonomicName>
<emphasis id="B9684D4C1A5DFFA497AA71F45B6E8347" box="[189,324,1789,1815]" italics="true" pageId="7" pageNumber="8">unilateralis</emphasis>
<bibRefCitation id="EF8DECAF1A5DFFA497AA70295B56836B" author="Schuchert P." box="[189,380,1824,1851]" firstAuthor="Schuchert" journalOrPublisher="Revue suisse de Zoologie" pageId="7" pageNumber="8" pagination="325 - 370" part="122" refId="ref23682" refString="Schuchert P. 2015. On some hydroids (Cnidaria, Hydrozoa) from the Okinawa Islands, Japan. Revue suisse de Zoologie 122 (2): 325 - 370. https: // doi. org / 10.5281 / zenodo. 30004" title="On some hydroids (Cnidaria, Hydrozoa) from the Okinawa Islands, Japan" type="journal article" year="2015">Schuchert, 2015</bibRefCitation>
</td>
<td id="76FDFA601A5D005C96BA719A5F5C83D1" box="[429,1398,1683,1921]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">
Colony
<emphasis id="B9684D4C1A5DFFA4951B719A581982FD" box="[524,563,1683,1709]" italics="true" pageId="7" pageNumber="8">tall</emphasis>
(ca 40 cm high), stem
<emphasis id="B9684D4C1A5DFFA49451719A5E3A82FD" box="[838,1040,1683,1709]" italics="true" pageId="7" pageNumber="8">loosely branched</emphasis>
, fascicled, axis
<emphasis id="B9684D4C1A5DFFA493D8719A5F3582FD" box="[1231,1311,1683,1709]" italics="true" pageId="7" pageNumber="8">helical</emphasis>
; cladia
<emphasis id="B9684D4C1A5DFFA496BA71BF58AD8280" box="[429,647,1718,1744]" italics="true" pageId="7" pageNumber="8">spirally arranged</emphasis>
; hydrotheca longitudinally asymmetrical, with two “loop- shaped ridges” on its lateral walls,
<emphasis id="B9684D4C1A5DFFA4944D71D35E2082A3" box="[858,1034,1754,1779]" italics="true" pageId="7" pageNumber="8">not connecting</emphasis>
ab- and adaxial walls; mesial nematotheca short,
<emphasis id="B9684D4C1A5DFFA495B671F458CD8347" box="[673,743,1789,1815]" italics="true" pageId="7" pageNumber="8">below</emphasis>
the hydrothecal base; a lateral
<emphasis id="B9684D4C1A5DFFA4939671F45F3D8347" box="[1153,1303,1789,1815]" italics="true" pageId="7" pageNumber="8">nematotheca</emphasis>
on the cormidial internode; hydrothecal rim
<emphasis id="B9684D4C1A5DFFA4947070295996836A" box="[871,956,1824,1850]" italics="true" pageId="7" pageNumber="8">smooth</emphasis>
, except for the median, abaxial cusp; lateral nematothecae
<emphasis id="B9684D4C1A5DFFA495BD704A5998830D" box="[682,946,1859,1885]" italics="true" pageId="7" pageNumber="8">exceedingly elongated</emphasis>
, projecting
<emphasis id="B9684D4C1A5DFFA49355704A5E8C830D" box="[1090,1190,1859,1885]" italics="true" pageId="7" pageNumber="8">frontally</emphasis>
,
<emphasis id="B9684D4C1A5DFFA493A0704A5F5F830D" box="[1207,1397,1859,1885]" italics="true" pageId="7" pageNumber="8">almost reaching</emphasis>
abaxial cusp. Distribution: Japan (
<bibRefCitation id="EF8DECAF1A5DFFA4942A706F59D683D1" author="Schuchert P." box="[829,1020,1894,1921]" firstAuthor="Schuchert" journalOrPublisher="Revue suisse de Zoologie" pageId="7" pageNumber="8" pagination="325 - 370" part="122" refId="ref23682" refString="Schuchert P. 2015. On some hydroids (Cnidaria, Hydrozoa) from the Okinawa Islands, Japan. Revue suisse de Zoologie 122 (2): 325 - 370. https: // doi. org / 10.5281 / zenodo. 30004" title="On some hydroids (Cnidaria, Hydrozoa) from the Okinawa Islands, Japan" type="journal article" year="2015">Schuchert 2015</bibRefCitation>
).
</td>
</tr>
</table>
</paragraph>
<subSubSection id="C306C2D51A52FFAB97AA76025EFF8002" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BA3915E1A52FFAB97AA76025B058575" blockId="8.[189,303,267,293]" box="[189,303,267,293]" pageId="8" pageNumber="9">
<heading id="D0EB26321A52FFAB97AA76025B058575" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9684D4C1A52FFAB97AA76025B058575" bold="true" box="[189,303,267,293]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A52FFAB97AA76375F748615" blockId="8.[189,1399,318,581]" pageId="8" pageNumber="9">
The relationships between several related aglaopheniid genera, viz.
<taxonomicName id="4C1CEADD1A52FFAB930876375F5B8508" authority="Allman, 1874" authorityName="Allman" authorityYear="1874" box="[1055,1393,318,344]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB930876375E9D8508" box="[1055,1207,318,344]" italics="true" pageId="8" pageNumber="9">Cladocarpus</emphasis>
Allman, 1874
</taxonomicName>
,
<taxonomicName id="4C1CEADD1A52FFAB97AA766C581B852F" authority="Fewkes, 1881" authorityName="Fewkes" authorityYear="1881" box="[189,561,357,383]" class="Hydrozoa" family="Aglaopheniidae" genus="Aglaophenopsis" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB97AA766C5B52852F" box="[189,376,357,383]" italics="true" pageId="8" pageNumber="9">Aglaophenopsis</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB9692766C581B852F" accessDate="2020-03-06" author="Fewkes J. W." box="[389,561,357,383]" firstAuthor="Fewkes" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, at Harvard College" pageId="8" pageNumber="9" pagination="127 - 140" part="8" publicationUrl="https://www.biodiversitylibrary.org/page/28870347" refId="ref22175" refString="Fewkes J. W. 1881. Report on the Acalephae. In: Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea, in 1878, 1879, and along the Atlantic coast of the United States, during the summer of 1880, by the U. S. Coast Survey Steamer Blake. Bulletin of the Museum of Comparative Zoology, at Harvard College 8 (7): 127 - 140. Available from https: // www. biodiversitylibrary. org / page / 28870347 [accessed 6 Mar. 2020]." title="Report on the Acalephae" type="journal article" volumeTitle="Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Caribbean Sea, in 1878, 1879, and along the Atlantic coast of the United States, during the summer of 1880, by the U. S. Coast Survey Steamer Blake" year="1881">Fewkes, 1881</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1CEADD1A52FFAB9554766C59BC852F" authority="Allman, 1883" authorityName="Allman" authorityYear="1883" box="[579,918,357,383]" class="Hydrozoa" family="Aglaopheniidae" genus="Streptocaulus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB9554766C58C9852F" box="[579,739,357,383]" italics="true" pageId="8" pageNumber="9">Streptocaulus</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB95F9766C59BC852F" author="Allman G. J." box="[750,918,357,383]" firstAuthor="Allman" journalOrPublisher="Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology" pageId="8" pageNumber="9" pagination="1 - 55" part="7" refId="ref21222" refString="Allman G. J. 1883. Report on the Hydroida dredged by H. M. S. Challenger during the years 1873 - 76. Part I. - Plumularidae. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology 7 (20): 1 - 55. https: // doi. org / 10.5962 / bhl. title. 11299" title="Report on the Hydroida dredged by H. M. S. Challenger during the years 1873 - 76. Part I. - Plumularidae" type="journal article" year="1883">Allman, 1883</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1CEADD1A52FFAB94CD766C5F07852F" authority="Broch, 1918" authorityName="Broch" authorityYear="1918" box="[986,1325,357,383]" class="Hydrozoa" family="Aglaopheniidae" genus="Nematophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB94CD766C5EAD852F" box="[986,1159,357,383]" italics="true" pageId="8" pageNumber="9">Nematophorus</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB9383766C5F07852F" author="Broch H." box="[1172,1325,357,383]" firstAuthor="Broch" journalOrPublisher="Danish Ingolf-Expedition" pageId="8" pageNumber="9" pagination="1 - 206" part="5" refId="ref21849" refString="Broch H. 1918. Hydroida (Part II). Danish Ingolf-Expedition 5 (7): 1 - 206." title="Hydroida (Part II)" type="journal article" year="1918">Broch, 1918</bibRefCitation>
</taxonomicName>
, have been discussed and interpreted by a number of authors (e.g.,
<bibRefCitation id="EF8DECAF1A52FFAB94B876845ECA85F7" accessDate="2020-03-06" author="Ramil F. &amp; Vervoort W." box="[943,1248,397,423]" firstAuthor="Ramil" journalOrPublisher="Scientia Marina" pageId="8" pageNumber="9" pagination="171 - 176" part="56" publicationUrl="http://scimar.icm.csic.es/scimar/pdf/56/sm56n2171.pdf" refId="ref23140" refString="Ramil F. &amp; Vervoort W. 1992 a. Some consideration concerning the genus Cladocarpus (Cnidaria: Hydrozoa). Scientia Marina 56 (2 - 3): 171 - 176. Available from http: // scimar. icm. csic. es / scimar / pdf / 56 / sm 56 n 2171. pdf [accessed 6 Mar. 2020]." title="Some consideration concerning the genus Cladocarpus (Cnidaria: Hydrozoa)" type="journal article" year="1992" yearSuffix="a">Ramil &amp; Vervoort 1992a</bibRefCitation>
;
<bibRefCitation id="EF8DECAF1A52FFAB93F876845B6B859E" accessDate="2020-03-06" author="Ansin Agis J. &amp; Ramil F. &amp; Vervoort W." etAl="et al." firstAuthor="Ansin Agis" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="8" pageNumber="9" pagination="1 - 268" part="333" publicationUrl="https://repository.naturalis.nl/document/46286" refId="ref21371" refString="Ansin Agis J., Ramil F. &amp; Vervoort W. 2001. Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, the Netherlands. Zoologische Verhandelingen, Leiden 333: 1 - 268. Available from https: // repository. naturalis. nl / document / 46286 [accessed 6 Mar. 2020]." title="Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, the Netherlands" type="journal article" year="2001">
Ansín Agís
<emphasis id="B9684D4C1A52FFAB97AA76BC5AD3859E" box="[189,249,436,462]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation id="EF8DECAF1A52FFAB965A76BD5802859E" author="Altuna A. &amp; Murillo F. J. &amp; Calder D. R." box="[333,552,436,462]" etAl="et al." firstAuthor="Altuna" journalOrPublisher="Zootaxa" pageId="8" pageNumber="9" pagination="501 - 537" part="3757" refId="ref21302" refString="Altuna A., Murillo F. J. &amp; Calder D. R. 2013. Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from bathyal waters of the Flemish Cap, Flemish Pass, and Grand Banks of Newfoundland (NW Atlantic). Zootaxa 3757 (5): 501 - 537. https: // doi. org / 10.11646 / zootaxa. 3737.5.1" title="Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from bathyal waters of the Flemish Cap, Flemish Pass, and Grand Banks of Newfoundland (NW Atlantic)" type="journal article" year="2013">
Altuna
<emphasis id="B9684D4C1A52FFAB96B176BC5BC9859E" box="[422,483,436,462]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2013
</bibRefCitation>
) on the account of morphological characters alone. To these should be added other genera, such as
<taxonomicName id="4C1CEADD1A52FFAB951976D559CB85A6" authority="Vervoort &amp; Watson, 2003" authorityName="Vervoort &amp; Watson" authorityYear="2003" box="[526,993,476,502]" class="Hydrozoa" family="Aglaopheniidae" genus="Carpocladus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB951976D5588C85A6" box="[526,678,476,502]" italics="true" pageId="8" pageNumber="9">Carpocladus</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB95B976D559CB85A6" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[686,993,476,502]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="8" pageNumber="9" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson, 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1CEADD1A52FFAB94F876D55F6E85A6" authority="Bogle, 1984" authorityName="Bogle" authorityYear="1984" box="[1007,1348,476,502]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpoides" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB94F876D55E8085A6" box="[1007,1194,476,502]" italics="true" pageId="8" pageNumber="9">Cladocarpoides</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB93A476D55F6E85A6" author="Bogle M. A." box="[1203,1348,476,502]" firstAuthor="Bogle" journalOrPublisher="Bulletin of Marine Science" pageId="8" pageNumber="9" pagination="135 - 140" part="34" refId="ref21753" refString="Bogle M. A. 1984. Cladocarpoides yucatanicus, a new genus and species of Aglaopheniinae (Coelenterata: Hydroida: Plumulariidae) from Arrowsmith Bank, Yucatan Channel. Bulletin of Marine Science 34 (1): 135 - 140." title="Cladocarpoides yucatanicus, a new genus and species of Aglaopheniinae (Coelenterata: Hydroida: Plumulariidae) from Arrowsmith Bank, Yucatan Channel" type="journal article" year="1984">Bogle, 1984</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1CEADD1A52FFAB97AA750A5887864D" authority="Vervoort &amp; Watson, 2003" authorityName="Vervoort &amp; Watson" authorityYear="2003" box="[189,685,515,542]" class="Hydrozoa" family="Aglaopheniidae" genus="Wanglaophenia" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB97AA750A5B5E864D" box="[189,372,515,541]" italics="true" pageId="8" pageNumber="9">Wanglaophenia</emphasis>
<bibRefCitation id="EF8DECAF1A52FFAB966B750D5887864D" accessDate="2020-03-06" author="Vervoort W. &amp; Watson J. E." box="[380,685,515,542]" firstAuthor="Vervoort" journalOrPublisher="NIWA Biodiversity Memoir" pageId="8" pageNumber="9" pagination="1 - 538" part="119" publicationUrl="https://archive.org/details/niwabiodiversitymemoir01192003" refId="ref24067" refString="Vervoort W. &amp; Watson J. E. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). NIWA Biodiversity Memoir 119: 1 - 538. Available from https: // archive. org / details / niwabiodiversitymemoir 01192003 [accessed 6 Mar. 2020]." title="The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids)" type="journal article" year="2003">Vervoort &amp; Watson, 2003</bibRefCitation>
</taxonomicName>
, that make the case even more complex and, taken globally, it becomes evident that only a genetic study can be expected to clarify their relationships objectively.
</paragraph>
<paragraph id="8BA3915E1A52FFAB97AA7579597C8778" blockId="8.[189,1399,624,809]" pageId="8" pageNumber="9">
The present hydroid, as well as
<taxonomicName id="4C1CEADD1A52FFAB953C7579590E86DA" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[555,804,624,650]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="partitus" status="sp. nov.">
<emphasis id="B9684D4C1A52FFAB953C7579590E86DA" box="[555,804,624,650]" italics="true" pageId="8" pageNumber="9">Cladocarpus partitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A52FFAB943D757859AE86DB" box="[810,900,625,651]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1CEADD1A52FFAB94AD75785E6F86DA" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[954,1093,625,651]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="pennatus" status="sp. nov.">
<emphasis id="B9684D4C1A52FFAB94AD75785E6F86DA" box="[954,1093,625,651]" italics="true" pageId="8" pageNumber="9">C. pennatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A52FFAB935C75785E8F86DB" box="[1099,1189,625,651]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
(see below), have gonosomes that do not conform accurately to any diagnosis of the abovementioned genera, displaying only some of their characters, as well as additional, distinctive ones. For this reason, these three new species are provisionally accommodated in the genus
<taxonomicName id="4C1CEADD1A52FFAB942075EE59E58751" authorityName="Allman" authorityYear="1874" box="[823,975,743,769]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB942075EE59E58751" box="[823,975,743,769]" italics="true" pageId="8" pageNumber="9">Cladocarpus</emphasis>
</taxonomicName>
, the oldest available name, pending a future phylogenetic study based on molecular markers.
</paragraph>
<paragraph id="8BA3915E1A52FFAB97AA745D587487ED" blockId="8.[189,1399,852,958]" pageId="8" pageNumber="9">
A few congeners have hydrothecae resembling those of
<taxonomicName id="4C1CEADD1A52FFAB9456745C5E29873E" authority="Galea, 2020" authorityName="Galea" authorityYear="2020" box="[833,1027,853,879]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B9684D4C1A52FFAB9456745C5E29873E" box="[833,1027,853,879]" italics="true" pageId="8" pageNumber="9">C. asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25BF0371A52FFAB931E745C5E4F873F" box="[1033,1125,853,879]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, notably with respect to their elongated appearance and the presence of two lateral, oblique, intrathecal ridges. Their distinctive features are summarized in
<tableCitation id="C69EA4E51A52FFAB951674AA587387ED" box="[513,601,931,957]" captionStart="Table 1" captionStartId="7.[189,258,268,294]" captionText="Table 1. Nominal species of Cladocarpus Allman, 1874 with (occasionally asymmetrical) hydrothecae provided with sigmoid (or nearly so) lateral ridges, similar to those of C. asymmetricus sp. nov., and their morphological differences with the new species." pageId="8" pageNumber="9">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BA3915E1A52FFAB97AA74E05EFF8002" blockId="8.[189,1398,1001,1106]" pageId="8" pageNumber="9">
The new species is also highly distinctive in that its gonothecae are provided with nematothecae, a situation unknown in any of the
<taxonomicName id="4C1CEADD1A52FFAB953D731958E8807A" authorityName="Allman" authorityYear="1874" box="[554,706,1040,1066]" class="Hydrozoa" family="Aglaopheniidae" genus="Cladocarpus" kingdom="Animalia" order="Leptothecata" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">
<emphasis id="B9684D4C1A52FFAB953D731958E8807A" box="[554,706,1040,1066]" italics="true" pageId="8" pageNumber="9">Cladocarpus</emphasis>
</taxonomicName>
-like species described so far. Moreover, these nematothecae have three different morphologies and are present at different positions on the gonotheca.
</paragraph>
</subSubSection>
<subSubSection id="C306C2D51A52FFAB97AA737459DF809B" box="[189,1013,1149,1227]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BA3915E1A52FFAB97AA73745B7C80C7" blockId="8.[189,1013,1149,1227]" box="[189,342,1149,1175]" pageId="8" pageNumber="9">
<heading id="D0EB26321A52FFAB97AA73745B7C80C7" bold="true" box="[189,342,1149,1175]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9684D4C1A52FFAB97AA73745B7C80C7" bold="true" box="[189,342,1149,1175]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3915E1A52FFAB97AA73B859DF809B" blockId="8.[189,1013,1149,1227]" box="[189,1013,1200,1227]" pageId="8" pageNumber="9">
Known only from its
<typeStatus id="54A72FFC1A52FFAB96AD73B85BC7809B" box="[442,493,1201,1227]" pageId="8" pageNumber="9">type</typeStatus>
locality, off
<collectingCountry id="F30BD1CE1A52FFAB959673B8591D809A" box="[641,823,1200,1227]" name="New Caledonia" pageId="8" pageNumber="9">New Caledonia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>
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