337 lines
38 KiB
XML
337 lines
38 KiB
XML
<document id="F087F14B07D3517DE1F680E8EF96DDBC" ID-DOI="10.5281/zenodo.156784" ID-GBIF-Dataset="7450e6f5-2d35-43f7-9cbc-5aa4eab735b6" ID-ISSN="11755326" ID-Zenodo-Dep="156784" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459597271218" checkinUser="plazi" docAuthor="Kohlmann, Bert & Delgado, Enio Cano And Leonardo" docDate="2003" docId="03A887EBFFD3FFECFEFBFCCCB4377FE6" docLanguage="en" docName="zt00167.pdf" docOrigin="Zootaxa 167" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Copris nubilosus Kohlmann, Cano and Delgado, sp. nov." docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="FF91FF93FFD0FFE9FFF3FFD8B6687C38" masterDocTitle="New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America" masterLastPageNumber="16" masterPageNumber="1" pageNumber="4" updateTime="1698202608495" updateUser="plazi">
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<mods:mods id="3E9942EE6BAD27A4760E6E764463066E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2485E3E241038AFB79F5814CC03BA67A">
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<mods:title id="ED7881EFEA87177B7C22DF3C881F67AB">New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America</mods:title>
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</mods:titleInfo>
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<mods:name id="B3DB396A3AB467CD07F44DAC640CD80B" type="personal">
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<mods:roleTerm id="3CE592A5D4B4C82FB75E8DB085D355DB">Author</mods:roleTerm>
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<mods:namePart id="16AFFE5BC984C287552B0CE0DDD015E8">Kohlmann, Bert</mods:namePart>
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<mods:name id="986C3960C8E4598E189E001C2E13277A" type="personal">
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<mods:roleTerm id="48E013182E5DB354608BE57011378483">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B42A8616B0E406C654E1DECA9CF8A1D1">Delgado, Enio Cano And Leonardo</mods:namePart>
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<mods:typeOfResource id="5753CEEA508B6A879EEA960FFCECE1A5">text</mods:typeOfResource>
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<mods:title id="378445499314A345E1A89FBF5DA8158E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AAB453D2BC88B79535596FA3F5B3BE75">
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<mods:date id="D2D5577461FB91EA3055F18D5890A288">2003</mods:date>
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<mods:detail id="6DCCFA456E0070F3A5485D548D9CF21E" type="volume">
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<mods:number id="35AB534EB32E3D83BCF348C849C22F88">167</mods:number>
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<mods:classification id="3AFD5DC4A7A53FDEBF028B67603A84F4">journal article</mods:classification>
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<mods:identifier id="E3B58E094F0C8DA007566B3B550BE2B3" type="DOI">10.5281/zenodo.156784</mods:identifier>
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<mods:identifier id="DF8BA49985A1A2679B7BDB129925294B" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="B462449DF9EC90EAF79DDCC525660D4E" type="Zenodo-Dep">156784</mods:identifier>
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<treatment id="03A887EBFFD3FFECFEFBFCCCB4377FE6" ID-DOI="http://doi.org/10.5281/zenodo.6276529" ID-GBIF-Taxon="119331250" ID-Zenodo-Dep="6276529" LSID="urn:lsid:plazi:treatment:03A887EBFFD3FFECFEFBFCCCB4377FE6" httpUri="http://treatment.plazi.org/id/03A887EBFFD3FFECFEFBFCCCB4377FE6" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<subSubSection id="C31B6576FFD3FFEAFEFBFCCCB21A7F15" box="[264,1138,787,814]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BBE36FDFFD3FFEAFEFBFCCCB21A7F15" blockId="3.[264,1138,787,814]" box="[264,1138,787,814]" pageId="3" pageNumber="4">
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<heading id="D0F68191FFD3FFEAFEFBFCCCB21A7F15" bold="true" box="[264,1138,787,814]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="B975EAEFFFD3FFEAFEFBFCCCB21A7F15" bold="true" box="[264,1138,787,814]" pageId="3" pageNumber="4">
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<taxonomicName id="4C014D7EFFD3FFEAFEFBFCCCB5317F15" ID-CoL="7ZML8" authority="Kohlmann, Cano and Delgado" authorityName="Kohlmann, Cano and Delgado" box="[264,857,787,814]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nubilosus" status="sp. nov.">
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<emphasis id="B975EAEFFFD3FFEAFEFBFCCCB7BC7F16" bold="true" box="[264,468,788,814]" italics="true" pageId="3" pageNumber="4">Copris nubilosus</emphasis>
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Kohlmann, Cano and Delgado
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2465794FFD3FFEAFC95FCCBB5AB7F15" box="[870,963,787,813]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="133A2A78FFD3FFEAFC29FCCBB2467F15" box="[986,1070,787,813]" captionStart="FIGURE. 2" captionStartId="7.[264,370,1696,1720]" captionTargetBox="[391,1181,414,1621]" captionTargetId="figure@7.[365,1223,326,1673]" captionTargetPageId="7" captionText="FIGURE. 2. Dorsal view of Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168712/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
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,
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<figureCitation id="133A2A78FFD3FFEAFBCFFCCBB2247F15" box="[1084,1100,787,813]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
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,
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<figureCitation id="133A2A78FFD3FFEAFBAAFCCBB2007F15" box="[1113,1128,787,813]" captionStart="FIGURE 5" captionStartId="12.[264,367,1467,1491]" captionTargetBox="[291,1299,810,1440]" captionTargetId="figure@12.[288,1299,809,1440]" captionTargetPageId="12" captionText="FIGURE 5. Distribution of Copris cited in the work (except the widely distributed C. lugubris and the Mexican C. sallei) in Central America. White area is above 800 m. Black stars = C. nubilosus Kohlmann, Cano and Delgado, sp. nov.; black triangle = C. caliginosus Kohlmann, Cano and Delgado, sp. nov.; black squares = C. aspericollis; black circle = C. matthewsi pacificus; white triangle = C. costaricensis dolichocerus; white rhombus = C. laeviceps." httpUri="https://zenodo.org/record/156788/files/figure.png" pageId="3" pageNumber="4">5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B6576FFD3FFEAFEFBFC8EB7327859" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BBE36FDFFD3FFEAFEFBFC8EB7327859" blockId="3.[264,1324,854,1880]" pageId="3" pageNumber="4">
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<emphasis id="B975EAEFFFD3FFEAFEFBFC8EB7E07F48" bold="true" box="[264,392,854,880]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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This species differs from the
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<taxonomicName id="4C014D7EFFD3FFEAFD11FC8FB5E27F48" box="[738,906,855,880]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caliginosus">
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<emphasis id="B975EAEFFFD3FFEAFD11FC8FB5E27F48" box="[738,906,855,880]" italics="true" pageId="3" pageNumber="4">C. caliginosus</emphasis>
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</taxonomicName>
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in the structure of the male anterior pronotal angle. The anterior margin forms a small, downwardly directed median point and the anterolateral angles are separated from it by a more or less level area (
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<figureCitation id="133A2A78FFD3FFEAFB9FFC7FB2DC7FF9" box="[1132,1204,935,961]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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); whereas in
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<taxonomicName id="4C014D7EFFD3FFEAFED9FC17B7D87FD0" box="[298,432,975,1000]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caliginosus">
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<emphasis id="B975EAEFFFD3FFEAFED9FC17B7D87FD0" box="[298,432,975,1000]" italics="true" pageId="3" pageNumber="4">caliginosus</emphasis>
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</taxonomicName>
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the margin does not form this median point and the anterolateral angles occupying this space and an excavated area forms behind the angles (
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<figureCitation id="133A2A78FFD3FFEAFBA7FC2FB2C87829" box="[1108,1184,1015,1041]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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). Also, this species has reddishbrown antennal lamellae, whereas in C.
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<taxonomicName id="4C014D7EFFD3FFEAFC1FFBC7B21A7800" box="[1004,1138,1055,1080]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caliginosus">
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<emphasis id="B975EAEFFFD3FFEAFC1FFBC7B21A7800" box="[1004,1138,1055,1080]" italics="true" pageId="3" pageNumber="4">caliginosus</emphasis>
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</taxonomicName>
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they are dark brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B6576FFD3FFEDFECBFBB6B5197896" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
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<paragraph id="8BBE36FDFFD3FFEAFECBFBB6B7557889" blockId="3.[264,1324,854,1880]" pageId="3" pageNumber="4">
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<emphasis id="B975EAEFFFD3FFEAFECBFBB6B55D78B0" bold="true" box="[312,821,1134,1160]" pageId="3" pageNumber="4">
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Description.
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<typeStatus id="54BA885FFFD3FFEAFE29FBB6B42678B0" box="[474,590,1134,1160]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
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. Male (
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<figureCitation id="133A2A78FFD3FFEAFD5EFBB6B56B78B0" box="[685,771,1134,1160]" captionStart="FIGURE. 2" captionStartId="7.[264,370,1696,1720]" captionTargetBox="[391,1181,414,1621]" captionTargetId="figure@7.[365,1223,326,1673]" captionTargetPageId="7" captionText="FIGURE. 2. Dorsal view of Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168712/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
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,
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<figureCitation id="133A2A78FFD3FFEAFCE0FBB6B54D78B0" box="[787,805,1134,1160]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
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):
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</emphasis>
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Total length:
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<quantity id="4CF99B18FFD3FFEAFC2DFBB7B22478B1" box="[990,1100,1135,1161]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="3" pageNumber="4" unit="mm" value="15.1">15.1 mm</quantity>
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. Elytral width:
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<quantity id="4CF99B18FFD3FFEAFAF5FBB7B7517889" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="4" unit="mm" value="8.6">8.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BBE36FDFFD3FFEAFECBFB67B45B7940" blockId="3.[264,1324,854,1880]" pageId="3" pageNumber="4">Head armed. Clypeus with two remote, small teeth, the margin between them curved inward in a broad, shallow arc. Posterior angles of genae acute. Upper surface of head umbilicopunctate, with the exception of the base and along genal suture. Head horn is long and strongly curved backwards. Demarcation between gula and submentum arcuate. Antennae reddishbrown.</paragraph>
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<paragraph id="8BBE36FDFFD3FFEAFECBFA5EB2617B60" blockId="3.[264,1324,854,1880]" pageId="3" pageNumber="4">
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Pronotum armed. Anterolateral angles acute, immediately followed by a sharply curved margin. Lateral carina rounded, issuing from margin. Anterior margin behind gena forming a small, downwardly directed median point and separated from the anterolateral acute angles by a more or less level area, not forming any excavation (
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<figureCitation id="133A2A78FFD3FFEAFB82FA26B2D67A20" box="[1137,1214,1534,1560]" captionStart="FIGURE. 4" captionStartId="9.[264,370,1708,1732]" captionTargetBox="[525,1064,323,1678]" captionTargetId="figure@9.[533,1053,793,1144]" captionTargetPageId="9" captionText="FIGURE. 4. Lateral views of head and pronotum of Copris caliginosus Kohlmann, Cano and Delgado, sp. nov. (holotype, top), Copris nubilosus Kohlmann, Cano and Delgado, sp. nov. (holotype, middle) and Copris sallei (bottom)." httpUri="https://zenodo.org/record/168713/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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). Frontal declivity with a carina running through its middle. Median longitudinal sulcus impressed, umbilicopunctate. Punctation of pronotum as follows: coarsely umbilicopunctate in all depressions and margins; raised areas of pronotal base, prominences and most of anterior declivities contrastingly impunctate; no simple punctures. Lateral pronotal prominences small and pointing parallel to the body axis. Median pronotal prominences with a wide base and two diverging teeth pointing upwards. Anterior prosternal margin with a pointed tooth; sternellum umbilicopunctate. Median lobe of metasternum umbilicopunctate anteriorly and laterally; median longitudinal impressed line complete.
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</paragraph>
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<paragraph id="8BBE36FDFFD4FFEDFECBFEFCB2977DB6" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">Elytra with 8th stria incomplete; 9th stria arising at anterior third of elytron; 10th complete. Striae coarsely punctate, punctures circular, umbilical, and separated by a distance equal to about their diameter. Interstriae slightly convex and microscopically punctate.</paragraph>
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<paragraph id="8BBE36FDFFD4FFEDFECBFE44B4497DE6" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">Pygidium with complete margin, irregularly umbilicopunctate, punctation bearing short, stiff golden setae.</paragraph>
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<paragraph id="8BBE36FDFFD4FFEDFECBFE34B5A47E9E" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">Ventral surface of profemur setigerous umbilicopunctate on posterior longitudinal half, microscopically punctate on anterior. Protibial apical spur linear and curved downwards tapering to a blunt end. Coxa with some umbilical punctures on outer surface. Mesofemora with umbilicate punctures setigerous toward apex and rear margin, rest of the surface microscopically punctate. Metafemur as mesofemur.</paragraph>
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<paragraph id="8BBE36FDFFD4FFEDFECBFD6BB78D7836" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">
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<emphasis id="B975EAEFFFD4FFEDFECBFD6BB4707EF5" bold="true" box="[312,536,691,717]" pageId="4" pageNumber="5">Allotype. Female:</emphasis>
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Total length:
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||
<quantity id="4CF99B18FFD4FFEDFD34FD6CB55E7EF6" box="[711,822,692,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="4" pageNumber="5" unit="mm" value="13.8">13.8 mm</quantity>
|
||
. Elytral width:
|
||
<quantity id="4CF99B18FFD4FFEDFC05FD6CB23F7EF6" box="[1014,1111,692,718]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="4" pageNumber="5" unit="mm" value="7.1">7.1 mm</quantity>
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||
. Differs from the
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<typeStatus id="54BA885FFFD4FFEDFEFBFD04B7067ECE" box="[264,366,732,758]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
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||
by the following characters: Head with a slightly raised horn, wider than long, apex excavated posterodorsally. Clypeal margin with teeth more extended and rounded, with an open medial notch. Antennae reddishbrown. Pronotum armed with a faintly developed central carina and two lateral tubercles behind declivity, carina wavelike, emarginated at middle. Pronotal disc and area near the base and posterior angles impunctate, remaining punctures umbilicate. Median longitudinal sulcus impressed and umbilicopunctate. Anterior angles forming a small tooth. Anterolateral regions of metasternal lobe umbilicopunctate.
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||
</paragraph>
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<paragraph id="8BBE36FDFFD4FFEDFECBFBC3B5197896" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">
|
||
<emphasis id="B975EAEFFFD4FFEDFECBFBC3B7DA780D" bold="true" box="[312,434,1051,1077]" pageId="4" pageNumber="5">Variation.</emphasis>
|
||
Total length:
|
||
<quantity id="4CF99B18FFD4FFEDFD97FBC4B57A780E" box="[612,786,1052,1078]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.69" metricValueMin="1.38" pageId="4" pageNumber="5" unit="mm" value="15.35" valueMax="16.9" valueMin="13.8">13.816.9 mm</quantity>
|
||
. Elytral width:
|
||
<quantity id="4CF99B18FFD4FFEDFC21FBC4B20B780E" box="[978,1123,1052,1078]" metricMagnitude="-3" metricUnit="m" metricValue="7.8999999999999995" metricValueMax="8.7" metricValueMin="7.1" pageId="4" pageNumber="5" unit="mm" value="7.8999999999999995" valueMax="8.7" valueMin="7.1">7.18.7 mm</quantity>
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. Less developed males have a small thin horn, that does not curve back; whereas in minor males the horn is reduced to a small pointed tubercle. Some male specimens present a faint frontal carina running through the middle of the pronotal declivity.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B6576FFD4FFEDFECBFB63B50D7A2E" pageId="4" pageNumber="5" type="materials_examined">
|
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<paragraph id="8BBE36FDFFD4FFEDFECBFB63B50D7A2E" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">
|
||
<emphasis id="B975EAEFFFD4FFEDFECBFB63B44278ED" bold="true" box="[312,554,1211,1237]" pageId="4" pageNumber="5">Examined material</emphasis>
|
||
(
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<specimenCount id="9D07FD74FFD4FFEDFDCEFB64B4F578EE" box="[573,669,1212,1238]" pageId="4" pageNumber="5" type="male">9 males</specimenCount>
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||
,
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<specimenCount id="9D07FD74FFD4FFEDFD58FB64B57C78EE" box="[683,788,1212,1238]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
|
||
).
|
||
<typeStatus id="54BA885FFFD4FFEDFCDEFB63B5F778ED" box="[813,927,1211,1237]" pageId="4" pageNumber="5" type="holotype">
|
||
<emphasis id="B975EAEFFFD4FFEDFCDEFB63B5F778ED" bold="true" box="[813,927,1211,1237]" pageId="4" pageNumber="5">Holotype</emphasis>
|
||
</typeStatus>
|
||
, male:
|
||
<collectingCountry id="F316766DFFD4FFEDFC08FB64B2DD78EE" box="[1019,1205,1212,1238]" name="Guatemala" pageId="4" pageNumber="5">GUATEMALA</collectingCountry>
|
||
.
|
||
<emphasis id="B975EAEFFFD4FFEDFB38FB64B34378ED" box="[1227,1323,1212,1237]" italics="true" pageId="4" pageNumber="5">Zacapa.</emphasis>
|
||
La Unión,
|
||
<quantity id="4CF99B18FFD4FFEDFE67FB3CB7BF78C6" box="[404,471,1252,1278]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="km" value="2.0">2 km</quantity>
|
||
Norte,
|
||
<date id="FFBF103DFFD4FFEDFDCEFB3CB4B678C6" box="[573,734,1252,1278]" pageId="4" pageNumber="5" value="1993-05-23" valueMax="1993-05-24" valueMin="1993-05-23">2324.V.1993</date>
|
||
,
|
||
<quantity id="4CF99B18FFD4FFEDFD02FB3CB52D78C6" box="[753,837,1252,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="m" value="1400.0">1400m</quantity>
|
||
, J. Monzón,
|
||
<emphasis id="B975EAEFFFD4FFEDFC1CFB3BB23178C5" bold="true" box="[1007,1113,1251,1277]" pageId="4" pageNumber="5">Allotype</emphasis>
|
||
, female:
|
||
<emphasis id="B975EAEFFFD4FFEDFB20FB3CB34B78C5" box="[1235,1315,1252,1277]" italics="true" pageId="4" pageNumber="5">ibidem</emphasis>
|
||
,
|
||
<date id="FFBF103DFFD4FFEDFEFBFAD4B71B791E" box="[264,371,1292,1318]" pageId="4" pageNumber="5" value="1994-09-25">25.IX.94</date>
|
||
, C. Estrada.
|
||
<emphasis id="B975EAEFFFD4FFEDFDFFFAD3B4F9791D" bold="true" box="[524,657,1291,1317]" pageId="4" pageNumber="5">
|
||
<typeStatus id="54BA885FFFD4FFEDFDFFFAD3B4E4791D" box="[524,652,1291,1317]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B975EAEFFFD4FFEDFD50FAD4B521791D" box="[675,841,1292,1317]" italics="true" pageId="4" pageNumber="5">Baja Verapaz.</emphasis>
|
||
Purulhá,
|
||
<date id="FFBF103DFFD4FFEDFC4EFAD4B229791E" box="[957,1089,1292,1318]" pageId="4" pageNumber="5" value="1995-04-30">30.IV.1995</date>
|
||
, X. Leiva,
|
||
<specimenCount id="9D07FD74FFD4FFEDFB34FAD4B34E791E" box="[1223,1318,1292,1318]" pageId="4" pageNumber="5" type="male">2 males</specimenCount>
|
||
; Purulhá, Biotopo del Quetzal,
|
||
<date id="FFBF103DFFD4FFEDFD86FAECB4817976" box="[629,745,1332,1358]" pageId="4" pageNumber="5" value="2000-08">VIII.2000</date>
|
||
heces humanas, A. Higueros,
|
||
<specimenCount id="9D07FD74FFD4FFEDFBAAFAECB2C67976" box="[1113,1198,1332,1358]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
|
||
.
|
||
<emphasis id="B975EAEFFFD4FFEDFB35FAECB706794D" italics="true" pageId="4" pageNumber="5">Huehuetenango.</emphasis>
|
||
Barillas, camino entre Nuevo San Mateo y San Juan de las Milpas, cerca de la Laguna Maxbal (
|
||
<emphasis id="B975EAEFFFD4FFEDFE21FA5CB79B79A5" box="[466,499,1412,1437]" italics="true" pageId="4" pageNumber="5">sic</emphasis>
|
||
, should be Muxbal),
|
||
<date id="FFBF103DFFD4FFEDFD1FFA5CB5E679A6" box="[748,910,1412,1438]" pageId="4" pageNumber="5" value="1998-05-28" valueMax="1998-05-30" valueMin="1998-05-28">2830.V.1998</date>
|
||
, bosque nuboso, E. Cano.
|
||
<emphasis id="B975EAEFFFD4FFEDFB38FA5CB34379A5" box="[1227,1323,1412,1437]" italics="true" pageId="4" pageNumber="5">Zacapa.</emphasis>
|
||
La Unión,
|
||
<quantity id="4CF99B18FFD4FFEDFE7BFA74B7AD79FE" box="[392,453,1452,1478]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="km" value="2.0">2 km</quantity>
|
||
Norte,
|
||
<date id="FFBF103DFFD4FFEDFDEDFA74B4A179FE" box="[542,713,1452,1478]" pageId="4" pageNumber="5" value="1992-04-12" valueMax="1992-04-14" valueMin="1992-04-12">1214.IV.1992</date>
|
||
,
|
||
<quantity id="4CF99B18FFD4FFEDFD24FA74B54279FE" box="[727,810,1452,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="m" value="1300.0">1300m</quantity>
|
||
, J. Monzón,
|
||
<specimenCount id="9D07FD74FFD4FFEDFC30FA74B27F79FE" box="[963,1047,1452,1478]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
|
||
;
|
||
<date id="FFBF103DFFD4FFEDFBD0FA74B2AD79FE" box="[1059,1221,1452,1478]" pageId="4" pageNumber="5" value="1993-05-23" valueMax="1993-05-24" valueMin="1993-05-23">2324.V.1993</date>
|
||
,
|
||
<quantity id="4CF99B18FFD4FFEDFB21FA74B34E79FE" box="[1234,1318,1452,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="m" value="1400.0">1400m</quantity>
|
||
, J. Monzón,
|
||
<specimenCount id="9D07FD74FFD4FFEDFE67FA0CB7B779D6" box="[404,479,1492,1518]" pageId="4" pageNumber="5" type="male">1male</specimenCount>
|
||
; La Unión,
|
||
<date id="FFBF103DFFD4FFEDFD9EFA0CB4B079D6" box="[621,728,1492,1518]" pageId="4" pageNumber="5" value="1994-09-25">25.IX.94</date>
|
||
, C. Estrada,
|
||
<specimenCount id="9D07FD74FFD4FFEDFC9CFA0CB5AB79D6" box="[879,963,1492,1518]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
|
||
.
|
||
<collectingCountry id="F316766DFFD4FFEDFC3CFA0CB21E79D6" box="[975,1142,1492,1518]" name="Honduras" pageId="4" pageNumber="5">HONDURAS</collectingCountry>
|
||
.
|
||
<emphasis id="B975EAEFFFD4FFEDFB79FA0CB28879D5" box="[1162,1248,1492,1517]" italics="true" pageId="4" pageNumber="5">Copán.</emphasis>
|
||
Cerro Azul, 4300 pies,
|
||
<date id="FFBF103DFFD4FFEDFE3CFA24B4367A2E" box="[463,606,1532,1558]" pageId="4" pageNumber="5" value="1993-04-05" valueMax="1993-04-06" valueMin="1993-04-05">56.IV.1993</date>
|
||
, W. González,
|
||
<specimenCount id="9D07FD74FFD4FFEDFCFDFA24B5097A2E" box="[782,865,1532,1558]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B6576FFD4FFECFECBF9FBB4147EA6" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BBE36FDFFD4FFEDFECBF9FBB28F7AB6" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">
|
||
<emphasis id="B975EAEFFFD4FFEDFECBF9FBB7D97A05" bold="true" box="[312,433,1571,1597]" pageId="4" pageNumber="5">Remarks.</emphasis>
|
||
This species belongs to the
|
||
<taxonomicName id="4C014D7EFFD4FFEDFCEDF9FCB5127A05" box="[798,890,1572,1597]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="remotus">
|
||
<emphasis id="B975EAEFFFD4FFEDFCEDF9FCB5127A05" box="[798,890,1572,1597]" italics="true" pageId="4" pageNumber="5">remotus</emphasis>
|
||
</taxonomicName>
|
||
complex, as defined by
|
||
<bibRefCitation id="EF904B0CFFD4FFEDFB4BF9FCB7327A5E" author="Matthews" pageId="4" pageNumber="16" refString="Matthews, E. G. (1961) A revision of the genus Copris Muller of the Western Hemisphere (Coleoptera, Scarabaeidae). Entomologica Americana (new series), XLI, 1 - 137." type="book chapter" year="1961">Matthews (1961)</bibRefCitation>
|
||
. This species will key to the
|
||
<taxonomicName id="4C014D7EFFD4FFEDFD4EF994B5CC7A5D" box="[701,932,1612,1637]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subSpecies" species="remotus" subSpecies="remotus">
|
||
<emphasis id="B975EAEFFFD4FFEDFD4EF994B5CC7A5D" box="[701,932,1612,1637]" italics="true" pageId="4" pageNumber="5">C. remotus remotus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C014D7EFFD4FFEDFC10F994B2EB7A5D" box="[995,1155,1612,1637]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B975EAEFFFD4FFEDFC10F994B2EB7A5D" box="[995,1155,1612,1637]" italics="true" pageId="4" pageNumber="5">C. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
dichotomy in Matthews’ key based on males, although it will not fit either of the two descriptions.
|
||
</paragraph>
|
||
<paragraph id="8BBE36FDFFD4FFEDFECBF944B3487B3E" blockId="4.[264,1324,284,1878]" pageId="4" pageNumber="5">
|
||
The only consistently reliable character for separating this species with C.
|
||
<taxonomicName id="4C014D7EFFD4FFEDFB56F944B3427A8D" box="[1189,1322,1692,1717]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caliginosus">
|
||
<emphasis id="B975EAEFFFD4FFEDFB56F944B3427A8D" box="[1189,1322,1692,1717]" italics="true" pageId="4" pageNumber="5">caliginosus</emphasis>
|
||
</taxonomicName>
|
||
is the antennal color.
|
||
<taxonomicName id="4C014D7EFFD4FFEDFDE5F91CB48B7AE5" box="[534,739,1732,1757]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nubilosus">
|
||
<emphasis id="B975EAEFFFD4FFEDFDE5F91CB48B7AE5" box="[534,739,1732,1757]" italics="true" pageId="4" pageNumber="5">Copris nubilosus</emphasis>
|
||
</taxonomicName>
|
||
has reddishbrown antennae, while those of
|
||
<taxonomicName id="4C014D7EFFD4FFEDFAE3F91CB7E67B3D" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caliginosus">
|
||
<emphasis id="B975EAEFFFD4FFEDFAE3F91CB7E67B3D" italics="true" pageId="4" pageNumber="5">C. caliginosus</emphasis>
|
||
</taxonomicName>
|
||
are dark brown. This species is closely related to
|
||
<taxonomicName id="4C014D7EFFD4FFEDFC24F934B2517B3D" box="[983,1081,1772,1797]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD4FFEDFC24F934B2517B3D" box="[983,1081,1772,1797]" italics="true" pageId="4" pageNumber="5">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C014D7EFFD4FFEDFB82F934B3717B3D" box="[1137,1305,1772,1797]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caliginosus">
|
||
<emphasis id="B975EAEFFFD4FFEDFB82F934B3717B3D" box="[1137,1305,1772,1797]" italics="true" pageId="4" pageNumber="5">C. caliginosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE36FDFFD4FFECFECBF8CCB4147EA6" blockId="4.[264,1324,284,1878]" lastBlockId="5.[264,1324,284,990]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C014D7EFFD4FFEDFECBF8CCB4697B15" box="[312,513,1812,1837]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nubilosus">
|
||
<emphasis id="B975EAEFFFD4FFEDFECBF8CCB4697B15" box="[312,513,1812,1837]" italics="true" pageId="4" pageNumber="5">Copris nubilosus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C014D7EFFD4FFEDFD52F8CCB56D7B15" box="[673,773,1812,1837]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD4FFEDFD52F8CCB56D7B15" box="[673,773,1812,1837]" italics="true" pageId="4" pageNumber="5">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
by having a much more strongly curved horn than
|
||
<emphasis id="B975EAEFFFD4FFEDFEB0F8E4B7C47B6D" box="[323,428,1852,1877]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C014D7EFFD4FFEDFEB0F8E4B7CF7B6D" box="[323,423,1852,1877]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">C. sallei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
antennae reddishbrown, whereas they are dark brown in
|
||
<taxonomicName id="4C014D7EFFD4FFEDFBAAF8E4B2D27B6D" box="[1113,1210,1852,1877]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD4FFEDFBAAF8E4B2D27B6D" box="[1113,1210,1852,1877]" italics="true" pageId="4" pageNumber="5">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
; the base of the midpronotal projections is evenly arched, whereas it forms a distinct gibbosity in
|
||
<taxonomicName id="4C014D7EFFD5FFECFAE3FEC4B72F7D65" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFAE3FEC4B72F7D65" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
; lateral pronotal prominences are small, whereas they are developed in
|
||
<taxonomicName id="4C014D7EFFD5FFECFB4EFE9CB34A7D65" box="[1213,1314,324,349]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFB4EFE9CB34A7D65" box="[1213,1314,324,349]" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
; pronotal face smooth and slightly concave with a faint carina running through its middle, umbilicopunctate only at its flanks, whereas the pronotal face in
|
||
<taxonomicName id="4C014D7EFFD5FFECFBDEFE4CB2FA7D95" box="[1069,1170,404,429]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFBDEFE4CB2FA7D95" box="[1069,1170,404,429]" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
is level and completely umbilicopunctate, no evidence of a central carina running through its middle; lateral pronotal prominences smooth, whereas grossly punctate in
|
||
<taxonomicName id="4C014D7EFFD5FFECFBE9FE3CB2157DC5" box="[1050,1149,484,509]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFBE9FE3CB2157DC5" box="[1050,1149,484,509]" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
; it differs also from
|
||
<taxonomicName id="4C014D7EFFD5FFECFEBAFDD4B7C27E1D" box="[329,426,524,549]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFEBAFDD4B7C27E1D" box="[329,426,524,549]" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
in having a well developed tooth on the anterior pronotal margin behind the gena and in having a feebly developed acute anterolateral margin; whereas
|
||
<taxonomicName id="4C014D7EFFD5FFECFB77FDECB28F7E75" box="[1156,1255,564,589]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B975EAEFFFD5FFECFB77FDECB28F7E75" box="[1156,1255,564,589]" italics="true" pageId="5" pageNumber="6">C. sallei</emphasis>
|
||
</taxonomicName>
|
||
has a very feeble tooth on the anterior pronotal margin behind the gena and a very acute and developed anterolateral margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B6576FFD5FFECFECBFD73B4037FB6" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="8BBE36FDFFD5FFECFECBFD73B4037FB6" blockId="5.[264,1324,284,990]" pageId="5" pageNumber="6">
|
||
<emphasis id="B975EAEFFFD5FFECFECBFD73B7B07EFD" bold="true" box="[312,472,683,709]" pageId="5" pageNumber="6">Distribution.</emphasis>
|
||
(
|
||
<figureCitation id="133A2A78FFD5FFECFE1BFD74B4587EFE" box="[488,560,684,710]" captionStart="FIGURE 5" captionStartId="12.[264,367,1467,1491]" captionTargetBox="[291,1299,810,1440]" captionTargetId="figure@12.[288,1299,809,1440]" captionTargetPageId="12" captionText="FIGURE 5. Distribution of Copris cited in the work (except the widely distributed C. lugubris and the Mexican C. sallei) in Central America. White area is above 800 m. Black stars = C. nubilosus Kohlmann, Cano and Delgado, sp. nov.; black triangle = C. caliginosus Kohlmann, Cano and Delgado, sp. nov.; black squares = C. aspericollis; black circle = C. matthewsi pacificus; white triangle = C. costaricensis dolichocerus; white rhombus = C. laeviceps." httpUri="https://zenodo.org/record/156788/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
|
||
). This species seems to be living in the mountainous areas of the Sierra de los Cuchumatanes, Sierra de las Minas and Sierra del Merendón in
|
||
<collectingCountry id="F316766DFFD5FFECFB5FFD0CB3437ED6" box="[1196,1323,724,750]" name="Guatemala" pageId="5" pageNumber="6">Guatemala</collectingCountry>
|
||
and
|
||
<collectingCountry id="F316766DFFD5FFECFECEFD24B7DA7F2E" box="[317,434,764,790]" name="Honduras" pageId="5" pageNumber="6">Honduras</collectingCountry>
|
||
, associated with cloud forest between
|
||
<quantity id="4CF99B18FFD5FFECFC75FD24B2777F2E" box="[902,1055,764,790]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.8" metricValueMin="1.35" pageId="5" pageNumber="6" unit="m" value="1575.0" valueMax="1800.0" valueMin="1350.0">13501800m</quantity>
|
||
, formed by species of
|
||
<taxonomicName id="4C014D7EFFD5FFECFEFBFCFCB7027F05" box="[264,362,804,829]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B975EAEFFFD5FFECFEFBFCFCB7027F05" box="[264,362,804,829]" italics="true" pageId="5" pageNumber="6">Quercus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C014D7EFFD5FFECFE88FCFCB7AF7F05" box="[379,455,804,829]" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B975EAEFFFD5FFECFE88FCFCB7AF7F05" box="[379,455,804,829]" italics="true" pageId="5" pageNumber="6">Clusia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C014D7EFFD5FFECFE24FCFCB43B7F05" box="[471,595,804,829]" class="Magnoliopsida" family="Lauraceae" genus="Nectandra" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B975EAEFFFD5FFECFE24FCFCB43B7F05" box="[471,595,804,829]" italics="true" pageId="5" pageNumber="6">Nectandra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B975EAEFFFD5FFECFD97FCFCB4C97F05" box="[612,673,804,829]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C014D7EFFD5FFECFD97FCFCB4F57F05" box="[612,669,804,829]" class="Insecta" family="Oecophoridae" genus="Inga" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Inga</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B975EAEFFFD5FFECFD2EFCFCB5317F05" box="[733,857,804,829]" italics="true" pageId="5" pageNumber="6">Hedyosum</emphasis>
|
||
. One would expect to find this species in the northern range of the Sierra Madre de Chiapas too, since Huehuetenango is but a few km away from the border.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B6576FFD5FFECFECBFC43B4377FE6" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8BBE36FDFFD5FFECFECBFC43B4377FE6" blockId="5.[264,1324,284,990]" pageId="5" pageNumber="6">
|
||
<emphasis id="B975EAEFFFD5FFECFECBFC43B7AD7F8D" bold="true" box="[312,453,923,949]" pageId="5" pageNumber="6">Etymology.</emphasis>
|
||
Latin adjective,
|
||
<taxonomicName id="4C014D7EFFD5FFECFD62FC44B5697F8D" box="[657,769,924,949]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nubilosus">
|
||
<emphasis id="B975EAEFFFD5FFECFD62FC44B5697F8D" box="[657,769,924,949]" italics="true" pageId="5" pageNumber="6">nubilosus</emphasis>
|
||
</taxonomicName>
|
||
, meaning cloudy; alluding to the fact that the species lives in cloud forests.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |