263 lines
34 KiB
XML
263 lines
34 KiB
XML
<document id="41B0E2FDB2F5BDC4466C08C4E92E98F7" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott" docDate="2021" docId="03A887D05F51FF9CFF3EEC76FD844B69" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella cyprusensis Motamedinia & Skevington, sp. nov." docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="20" updateTime="1699012478263" updateUser="plazi">
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<mods:mods id="D548714A668AFFFF44B35ABBB1607158" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E19CA0B22948B2675AD8D32CC5821F64">
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<mods:title id="B7586DF0EE3C3E496C50E849A4DAE8A5">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="106A083908AD4D027E02CECDF0F9FB52" type="personal">
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<mods:role id="17DB431E246A405124E0C905CDF59EE2">
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<mods:roleTerm id="9FA4E5F069301BC56090D3CE099EF436">Author</mods:roleTerm>
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<mods:namePart id="A579E601BA38E354C9C910CEB264388F">Motamedinia, Behnam</mods:namePart>
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<mods:roleTerm id="374E38EAF746E13F996F1FE69BCDED13">Author</mods:roleTerm>
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<mods:namePart id="3BD15E5014BB34FE9D7FC735013A40C8">Skevington, Jeffrey H.</mods:namePart>
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<mods:nameIdentifier id="97902F7D917EDC97FD3C5487DB761A69" type="ORCID">0000-0002-1445-9870</mods:nameIdentifier>
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<mods:nameIdentifier id="6F6197A72C59D228EDD57EDBB30BAE21" type="email">jhskevington@gmail.com</mods:nameIdentifier>
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<mods:name id="22539096C48D4179483FD27164767880" type="personal">
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<mods:roleTerm id="573248425A603CB947DC5F8928A5EA7A">Author</mods:roleTerm>
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<mods:namePart id="AC5FCC5981CC5CA1D6458186F3544249">Kelso, Scott</mods:namePart>
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<mods:nameIdentifier id="295ED22816CA12F8A764A1127F847AF8" type="ORCID">0000-0001-5559-9477</mods:nameIdentifier>
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<mods:nameIdentifier id="B5D6AA4ED8C60BFE1909D6FEAA5905BD" type="email">scott.kelso@canada.ca</mods:nameIdentifier>
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<mods:typeOfResource id="B98D3FD8495D2B627893F57645388A33">text</mods:typeOfResource>
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<mods:title id="0A78DA7BF7704A314DE2ECEEAA50F03F">Zootaxa</mods:title>
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<mods:part id="745B9C9AA7A35EFA8341C8B469E8626D">
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<mods:date id="B0E3926EA56EAC4474403FA903E58FB3">2021</mods:date>
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<mods:number id="4CD421E4AE995E5E9BD6C4229DE4FC81">2021-07-16</mods:number>
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<mods:number id="BE5A7A5AC5D2A362640CD83B5B66AD52">5002</mods:number>
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<mods:number id="9AD3A937005381DABC55919F8A390262">1</mods:number>
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<mods:start id="7397B1E5108F908AD2819DEEF8FD3911">1</mods:start>
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<mods:identifier id="6E36B553569F455E678F56E0CF4A5397" type="DOI">10.11646/zootaxa.5002.1.1</mods:identifier>
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<mods:identifier id="A4C712DA213EB23F2249041CDC2D2578" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A887D05F51FF9CFF3EEC76FD844B69" ID-DOI="http://doi.org/10.5281/zenodo.5122554" ID-Zenodo-Dep="5122554" LSID="urn:lsid:plazi:treatment:03A887D05F51FF9CFF3EEC76FD844B69" httpUri="http://treatment.plazi.org/id/03A887D05F51FF9CFF3EEC76FD844B69" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
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<subSubSection id="C31B654D5F51FF9AFF3EEC76FC0E487F" box="[151,929,1844,1871]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BBE36C65F51FF9AFF3EEC76FC0E487F" blockId="19.[151,929,1844,1908]" box="[151,929,1844,1871]" pageId="19" pageNumber="20">
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<heading id="D0F681AA5F51FF9AFF3EEC76FC0E487F" bold="true" box="[151,929,1844,1871]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="3">
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<emphasis id="B975EAD45F51FF9AFF3EEC76FC0E487F" bold="true" box="[151,929,1844,1871]" pageId="19" pageNumber="20">
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<taxonomicName id="4C014D455F51FF9AFF3EEC76FC93487E" ID-CoL="9HZPB" authority="Motamedinia & Skevington" authorityName="Motamedinia & Skevington" box="[151,828,1844,1871]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cyprusensis" status="sp. nov.">
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<emphasis id="B975EAD45F51FF9AFF3EEC76FE76487F" bold="true" box="[151,473,1844,1871]" italics="true" pageId="19" pageNumber="20">Tomosvaryella cyprusensis</emphasis>
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Motamedinia & Skevington
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</taxonomicName>
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<taxonomicNameLabel id="A24657AF5F51FF9AFCEAEC77FC0E487F" box="[835,929,1845,1871]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31B654D5F51FF9AFF3EEC1BFE364843" box="[151,409,1881,1908]" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BBE36C65F51FF9AFF3EEC1BFE364843" blockId="19.[151,929,1844,1908]" box="[151,409,1881,1908]" pageId="19" pageNumber="20">
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<figureCitation id="133A2A435F51FF9AFF3EEC1BFEBD4843" box="[151,274,1881,1907]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="19" pageNumber="20">Figs 8A–E</figureCitation>
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,
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<figureCitation id="133A2A435F51FF9AFEB7EC1BFE934844" box="[286,316,1881,1908]" captionStart="FIGURE 50" captionStartId="85.[151,250,1898,1922]" captionTargetBox="[151,1436,1232,1874]" captionTargetId="figure-107@85.[151,1436,1232,1874]" captionTargetPageId="85" captionText="FIGURE 50. Distribution of Tomosvaryella angulata, T. argyrata, T. argyratoides, T. cyprusensis sp. nov. in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122771" httpUri="https://zenodo.org/record/5122771/files/figure.png" pageId="19" pageNumber="20">50</figureCitation>
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,
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<figureCitation id="133A2A435F51FF9AFEE1EC1BFE364843" box="[328,409,1881,1907]" captionStart="FIGURE 66" captionStartId="94.[151,250,1518,1542]" captionTargetBox="[151,1436,182,1494]" captionTargetId="figure-31@94.[151,1436,181,1494]" captionTargetPageId="94" captionText="FIGURE 66. Male habitus of Tomosvaryella species A–B) T. bistounensis sp. nov., holotype (JSS51939, CNC) A) in dorsal view B) in lateral view, C–D) T. cyprusensis sp. nov., paratype (JSS52349, CNC) C) in dorsal view D) in lateral view, E–F) T. ellipiensis sp. nov., holotype (JSS51945, CNC): E) in dorsal view F) in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122803" httpUri="https://zenodo.org/record/5122803/files/figure.png" pageId="19" pageNumber="20">66C–D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B654D5F51FF9DFF3EECE3FB2C4FC9" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="8BBE36C65F51FF9DFF3EECE3FB2C4FC9" blockId="19.[151,1437,1953,2015]" lastBlockId="20.[151,1436,151,249]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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<emphasis id="B975EAD45F51FF9AFF3EECE3FEBB488B" bold="true" box="[151,276,1953,1979]" pageId="19" pageNumber="20">Diagnosis:</emphasis>
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This species can be recognized by the shape of surstyli in lateral view, elongated and straight (
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<figureCitation id="133A2A435F51FF9AFA91ECE3FF0448EF" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="19" pageNumber="20">Fig. 8D– E</figureCitation>
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); epandrium longer than wide in dorsal view (
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<figureCitation id="133A2A435F51FF9AFD11EC87FCBC48EF" box="[696,787,1989,2015]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="19" pageNumber="20">Fig. 8A</figureCitation>
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); left gonopod higher than right one in ventral view (
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<figureCitation id="133A2A435F51FF9DFAC6EC87FF144F81" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">Fig. 8B</figureCitation>
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). Based on the shape of surstyli in dorsal view, this species resembles species in the kuthyi group (
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<taxonomicName id="4C014D455F56FF9DFB51EBD5FE6B4FE5" authority="De Meyer, 1995" authorityName="De Meyer" authorityYear="1995" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="freidbergi">
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<emphasis id="B975EAD45F56FF9DFB51EBD5FEAA4FE5" italics="true" pageId="20" pageNumber="21">Tomosvaryella freidbergi</emphasis>
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<bibRefCitation id="EF904B375F56FF9DFEA4EBF9FE6B4FE5" author="De Meyer, M." box="[269,452,186,213]" pageId="20" pageNumber="21" pagination="283 - 319" refId="ref56694" refString="De Meyer, M. (1995) The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana, 18, 283 - 319." type="journal article" year="1995">De Meyer, 1995</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C014D455F56FF9DFE7BEBF9FCB24FE5" authority="De Meyer, 1997" authorityName="De Meyer" authorityYear="1997" box="[466,797,187,213]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hispanica">
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<emphasis id="B975EAD45F56FF9DFE7BEBF9FDF34FE5" box="[466,604,187,213]" italics="true" pageId="20" pageNumber="21">T. hispanica</emphasis>
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<bibRefCitation id="EF904B375F56FF9DFDCAEBF9FCB24FE5" author="De Meyer, M." box="[611,797,187,213]" pageId="20" pageNumber="21" pagination="421 - 450" refId="ref56757" refString="De Meyer, M. (1997) Contribution to the Pipunculidae fauna of Spain: (Diptera). Beitrage zur Entomologie, 47, 421 - 450." type="journal article" year="1997">De Meyer, 1997</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C014D455F56FF9DFC80EBF9FBD04FE5" authority="De Meyer, 1995" authorityName="De Meyer" authorityYear="1995" box="[809,1151,186,213]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="israelensis">
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<emphasis id="B975EAD45F56FF9DFC80EBF9FC124FE5" box="[809,957,187,213]" italics="true" pageId="20" pageNumber="21">T. israelensis</emphasis>
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<bibRefCitation id="EF904B375F56FF9DFC6CEBF9FBD04FE5" author="De Meyer, M." box="[965,1151,186,213]" pageId="20" pageNumber="21" pagination="283 - 319" refId="ref56694" refString="De Meyer, M. (1995) The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana, 18, 283 - 319." type="journal article" year="1995">De Meyer, 1995</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C014D455F56FF9DFB23EBF9FAD34FE5" authority="Aczel, 1944" authorityName="Aczel" authorityYear="1944" box="[1162,1404,187,213]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kuthyi">
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<emphasis id="B975EAD45F56FF9DFB23EBF9FB454FE5" box="[1162,1258,187,213]" italics="true" pageId="20" pageNumber="21">T. kuthyi</emphasis>
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<bibRefCitation id="EF904B375F56FF9DFB58EBF9FAD34FE5" author="Aczel, M. L." box="[1265,1404,187,213]" pageId="20" pageNumber="21" pagination="75 - 130" refId="ref56188" refString="Aczel, M. L. (1944) Die gattung Tomosvaryella Acz. (Dipt.). (Dorylaiden-Studien VIII). Annales Historico-Naturales Musei Nationalis Hungarici, 37, 75 - 130." type="journal article" year="1944">Aczél, 1944</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C014D455F56FF9DFA21EBF9FE7E4FC9" authority="De Meyer, 1995" authorityName="De Meyer" authorityYear="1995" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="parakuthyi">
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<emphasis id="B975EAD45F56FF9DFA21EBF9FEBF4FC9" italics="true" pageId="20" pageNumber="21">T. parakuthyi</emphasis>
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<bibRefCitation id="EF904B375F56FF9DFEBEEB9DFE7E4FC9" author="De Meyer, M." box="[279,465,222,249]" pageId="20" pageNumber="21" pagination="283 - 319" refId="ref56694" refString="De Meyer, M. (1995) The pipunculid flies of Israel and the Sinai (Insecta, Diptera, Pipunculidae). Spixiana, 18, 283 - 319." type="journal article" year="1995">De Meyer, 1995</bibRefCitation>
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</taxonomicName>
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). It differs by the straight surstyli in lateral view (
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<figureCitation id="133A2A435F56FF9DFC55EB9DFBD94FC9" box="[1020,1142,223,249]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="20" pageNumber="21">Fig. 8D–E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF7E664E5F56FF9DFF3EEC04FDFF48B2" ID-DOI="http://doi.org/10.5281/zenodo.5122685" ID-Zenodo-Dep="5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1862,1886]" targetBox="[170,1418,273,1831]" targetPageId="20">
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<paragraph id="8BBE36C65F56FF9DFF3EEC04FDFF48B2" blockId="20.[151,1436,1862,1922]" pageId="20" pageNumber="21">
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<emphasis id="B975EAD45F56FF9DFF3EEC04FEBC486E" bold="true" box="[151,275,1862,1886]" pageId="20" pageNumber="21">FIGURE 8.</emphasis>
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Male genitalia of
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<taxonomicName id="4C014D455F56FF9DFE65EC04FD73486E" box="[460,732,1862,1886]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cyprusensis" status="sp. nov.">
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<emphasis id="B975EAD45F56FF9DFE65EC04FD73486E" box="[460,732,1862,1886]" italics="true" pageId="20" pageNumber="21">Tomosvaryella cyprusensis</emphasis>
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</taxonomicName>
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<emphasis id="B975EAD45F56FF9DFD4BEC04FC9D486E" bold="true" box="[738,818,1862,1886]" pageId="20" pageNumber="21">
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<taxonomicNameLabel id="A24657AF5F56FF9DFD4BEC04FC9D486E" box="[738,818,1862,1886]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.
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</paragraph>
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</caption>
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<subSubSection id="C31B654D5F57FF9CFF6EEBD4FB6F4C94" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BBE36C65F57FF9CFF6EEBD4FB6F4C94" blockId="21.[151,1437,150,1113]" pageId="21" pageNumber="22">
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<emphasis id="B975EAD45F57FF9CFF6EEBD4FE1A4F80" bold="true" box="[199,437,150,176]" pageId="21" pageNumber="22">Description: MALE</emphasis>
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: Body length (excluding antennae):
|
||
<quantity id="4CF99B235F57FF9CFCE4EBD4FC094F81" box="[845,934,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="mm" value="2.5">2.5 mm</quantity>
|
||
.
|
||
<emphasis id="B975EAD45F57FF9CFC19EBD4FC414F81" bold="true" box="[944,1006,150,177]" italics="true" pageId="21" pageNumber="22">Head</emphasis>
|
||
. Scape, pedicel and arista dark brown, flagellum light brown. Pedicel with three short, pale upper setae; flagellum tapering and gray white pollinose (LF: WF = 2.6). Eyes meeting for a distance of 5–6 facets. Frons dark silver-gray pollinose. Vertex black, lacking pollinosity. Occiput dark and gray pollinose.
|
||
<emphasis id="B975EAD45F57FF9CFDCFEA40FD6C4E2C" box="[614,707,258,285]" italics="true" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFDCFEA40FD134E2D" bold="true" box="[614,700,258,285]" italics="true" pageId="21" pageNumber="22">Thorax</emphasis>
|
||
.
|
||
</emphasis>
|
||
Postpronotal lobe light yellow, gray pollinose with 6–7 yellowish setae along the upper margin (up to
|
||
<quantity id="4CF99B235F57FF9CFD90EA64FD0C4E71" box="[569,675,294,321]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="22" unit="mm" value="0.08">0.08 mm</quantity>
|
||
). Prescutum, scutum, scutellum black. Scutum with two uniseriate rows of conspicuous intra-alar setae and two uniseriate rows of dorsocentral setae. Postalar callus with two dark setae. Scutellum gray pollinose without setae along lateral margin. Subscutellum black, gray pollinose. Pleura dark, gray pollinose.
|
||
<emphasis id="B975EAD45F57FF9CFEECEAD0FE2D4E9C" bold="true" box="[325,386,402,428]" italics="true" pageId="21" pageNumber="22">Wing</emphasis>
|
||
. Length:
|
||
<quantity id="4CF99B235F57FF9CFE59EAD0FDE64E9D" box="[496,585,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="mm" value="2.2">2.2 mm</quantity>
|
||
. LW:MWW = 3.05. Wing almost entirely covered with microtrichia. Crossvein r-m reaches dm at the middle. M
|
||
<subScript id="178534835F57FF9CFD90EA8BFDEE4EE7" attach="left" box="[569,577,457,471]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
|
||
moderately undulating in middle. Halter length:
|
||
<quantity id="4CF99B235F57FF9CFBCAEAF4FB654EE1" box="[1123,1226,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="21" pageNumber="22" unit="mm" value="0.37">0.37 mm</quantity>
|
||
. Whitish, brown at base.
|
||
<emphasis id="B975EAD45F57FF9CFF71EA99FEA14EC5" bold="true" box="[216,270,475,501]" italics="true" pageId="21" pageNumber="22">Legs</emphasis>
|
||
. Coxae dark, gray pollinose. Mid coxa with three dark anterior setae. Trochanters dark, gray pollinose, smooth. Femora dark brown, gray pollinose. All femora with 1–2 wrinkled indentations at base. Tibiae brown but light brown in basal half, with two ventral rows of short setae on anterior and three rows on posterior side. Hind tibia with two wrinkled indentations in middle. Tarsi brown with scattered brown setae at anterior margin. Pulvilli yellow. Claws light brown with black tips.
|
||
<emphasis id="B975EAD45F57FF9CFDB6E928FD3A4DB4" box="[543,661,618,645]" italics="true" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFDB6E928FD224DB5" bold="true" box="[543,653,618,645]" italics="true" pageId="21" pageNumber="22">Abdomen</emphasis>
|
||
.
|
||
</emphasis>
|
||
Ground color dark brown. Tergite 1 densely gray pollinose, with 6–8 brown lateral setae (up to
|
||
<quantity id="4CF99B235F57FF9CFE1FE9CCFDB24D99" box="[438,541,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="21" pageNumber="22" unit="mm" value="0.13">0.13 mm</quantity>
|
||
). Tergites 2 densely gray pollinose in anterior margin.
|
||
<emphasis id="B975EAD45F57FF9CFBDEE9CCFB454D98" box="[1143,1258,654,681]" italics="true" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFBDEE9CCFB4D4D99" bold="true" box="[1143,1250,654,681]" italics="true" pageId="21" pageNumber="22">Genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Genital capsule in dorsal view: epandrium and surstyli light brown, gray pollinose. Epandrium longer than wide (MLE:MWE = 1.33). Surstyli rather symmetrical. Both surstyli elongated, left longer than right one (
|
||
<figureCitation id="133A2A435F57FF9CFBCAE994FB114DC0" box="[1123,1214,726,752]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8A</figureCitation>
|
||
). Both surstyli bent towards each other apically (
|
||
<figureCitation id="133A2A435F57FF9CFE76E9B8FD944C24" box="[479,571,762,788]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8A</figureCitation>
|
||
). Genital capsule in ventral view: gonopods unequal in height, left is higher than right one (
|
||
<figureCitation id="133A2A435F57FF9CFE97E85CFE3A4C08" box="[318,405,798,824]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8B</figureCitation>
|
||
). Subepandrial sclerite elongated (
|
||
<figureCitation id="133A2A435F57FF9CFCBAE85CFCC54C08" box="[787,874,798,824]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8B</figureCitation>
|
||
). Genital capsule in lateral view: both surstyli long and straight, slightly broader at base (
|
||
<figureCitation id="133A2A435F57FF9CFDEBE800FD124C6C" box="[578,701,834,860]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8D–E</figureCitation>
|
||
). Phallic guide strong with distinct patch of setae dorsally before apex (
|
||
<figureCitation id="133A2A435F57FF9CFF77E824FEF54CB0" box="[222,346,870,896]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8D–E</figureCitation>
|
||
), phallus with three small ejaculatory ducts, one with some small teeth in basal half (
|
||
<figureCitation id="133A2A435F57FF9CFA9DE824FA204CB0" box="[1332,1423,870,896]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8B</figureCitation>
|
||
). Ejaculatory apodeme tube-like, bent, with a bulb in its middle (
|
||
<figureCitation id="133A2A435F57FF9CFCF2E8C8FC1B4C94" box="[859,948,906,932]" captionStart="FIGURE 8" captionStartId="20.[151,250,1862,1886]" captionTargetBox="[170,1418,273,1831]" captionTargetId="figure-91@20.[151,1436,273,1838]" captionTargetPageId="20" captionText="FIGURE 8. Male genitalia of Tomosvaryella cyprusensis sp. nov., holotype (JSS52302, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122685" httpUri="https://zenodo.org/record/5122685/files/figure.png" pageId="21" pageNumber="22">Fig. 8C</figureCitation>
|
||
).
|
||
<emphasis id="B975EAD45F57FF9CFC61E8C8FBE64C94" bold="true" box="[968,1097,906,932]" pageId="21" pageNumber="22">FEMALE:</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F57FF9CFF6EE8ECFD2C4B04" pageId="21" pageNumber="22" type="etymology">
|
||
<paragraph id="8BBE36C65F57FF9CFF6EE8ECFABE4CF8" blockId="21.[151,1437,150,1113]" box="[199,1297,942,969]" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFF6EE8ECFEFD4CF8" bold="true" box="[199,338,942,968]" pageId="21" pageNumber="22">Etymology:</emphasis>
|
||
The specific epithet refers to
|
||
<collectingCountry id="F31676565F57FF9CFD09E8ECFD5B4CF8" box="[672,756,942,968]" name="Cyprus" pageId="21" pageNumber="22">Cyprus</collectingCountry>
|
||
, from where the specimens have been collected.
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F57FF9CFF6EE890FD2C4B04" blockId="21.[151,1437,150,1113]" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFF6EE890FE654CDC" bold="true" box="[199,458,978,1004]" pageId="21" pageNumber="22">Specimens examined:</emphasis>
|
||
<materialsCitation id="3B693C9B5F57FF9CFE7BE890FD914B20" accessionNumber="JSS52302" collectingDate="2017-07-30" collectingDateMax="2017-08-13" collectingDateMin="2017-07-30" collectionCode="CNC" collectorName="O. Ozden" country="Cyprus" latitude="35.35" location="Kyrenia" longLatPrecision="1193" longitude="33.15" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Kyrenia" typeStatus="holotype">
|
||
<emphasis id="B975EAD45F57FF9CFE7BE890FDD54CDC" bold="true" box="[466,634,978,1004]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54BA88645F57FF9CFE7BE890FDDA4CDC" box="[466,629,978,1004]" pageId="21" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="F31676565F57FF9CFD2BE890FD5C4CDC" box="[642,755,978,1004]" name="Cyprus" pageId="21" pageNumber="22">CYPRUS</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C5F8245F57FF9CFD57E890FCF44CDC" box="[766,859,978,1004]" country="Cyprus" name="Kyrenia" pageId="21" pageNumber="22">Kyrenia</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE3550015F57FF9CFCC1E890FC644CDD" box="[872,971,978,1005]" degrees="35" direction="north" minutes="21" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="35.35">35°21’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3550015F57FF9CFC7EE890FB984CDC" box="[983,1079,978,1004]" degrees="33" direction="east" minutes="09" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="33.15">33°09’E</geoCoordinate>
|
||
,
|
||
<date id="FFBF10065F57FF9CFBEDE890FAB34CDC" box="[1092,1308,978,1004]" pageId="21" pageNumber="22" value="2017-07-30" valueMax="2017-08-13" valueMin="2017-07-30">
|
||
<collectingDate id="EFFBE9EE5F57FF9CFBEDE890FAB34CDC" box="[1092,1308,978,1004]" pageId="21" pageNumber="22" value="2017-07-30" valueMax="2017-08-13" valueMin="2017-07-30">30.vii–13.viii.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F453105F57FF9CFA80E890FA374CDC" box="[1321,1432,978,1004]" pageId="21" pageNumber="22">O. Ozden</collectorName>
|
||
,
|
||
<collectingMethod id="52404ED15F57FF9CFF3EE8B4FE854B21" box="[151,298,1014,1041]" pageId="21" pageNumber="22">Malaise trap</collectingMethod>
|
||
, JSS52302 (
|
||
<specimenCount id="9D07FD4F5F57FF9CFE6BE8B4FE494B20" box="[450,486,1014,1040]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED10AE035F57FF9CFE5EE8B4FD9A4B20" box="[503,565,1014,1040]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="21" pageNumber="22">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B693C9B5F57FF9CFDE7E8B4FDD04B04" accessionNumber="JSS52349" collectingDate="2017-09-10" collectingDateMax="2017-09-17" collectingDateMin="2017-09-10" collectionCode="CNC" collectorName="O. Ozden" country="Cyprus" latitude="35.35" location="Kyrenia" longLatPrecision="1193" longitude="33.15" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Kyrenia" typeStatus="paratype">
|
||
<emphasis id="B975EAD45F57FF9CFDE7E8B4FD434B20" bold="true" box="[590,748,1014,1040]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54BA88645F57FF9CFDE7E8B4FD484B20" box="[590,743,1014,1040]" pageId="21" pageNumber="22" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry id="F31676565F57FF9CFD51E8B4FCC64B20" box="[760,873,1014,1040]" name="Cyprus" pageId="21" pageNumber="22">CYPRUS</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C5F8245F57FF9CFCD0E8B4FC794B20" box="[889,982,1014,1040]" country="Cyprus" name="Kyrenia" pageId="21" pageNumber="22">Kyrenia</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE3550015F57FF9CFC4EE8B4FBE54B21" box="[999,1098,1014,1041]" degrees="35" direction="north" minutes="21" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="35.35">35°21’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3550015F57FF9CFBF2E8B4FB144B20" box="[1115,1211,1014,1040]" degrees="33" direction="east" minutes="09" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="33.15">33°09’E</geoCoordinate>
|
||
,
|
||
<date id="FFBF10065F57FF9CFB65E8B4FADF4B20" box="[1228,1392,1014,1040]" pageId="21" pageNumber="22" value="2017-09-10" valueMax="2017-09-17" valueMin="2017-09-10">
|
||
<collectingDate id="EFFBE9EE5F57FF9CFB65E8B4FADF4B20" box="[1228,1392,1014,1040]" pageId="21" pageNumber="22" value="2017-09-10" valueMax="2017-09-17" valueMin="2017-09-10">10–17.ix.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26F453105F57FF9CFA28E8B4FF4C4B04" pageId="21" pageNumber="22">O. Ozden</collectorName>
|
||
,
|
||
<collectingMethod id="52404ED15F57FF9CFF47EF58FED34B05" box="[238,380,1050,1077]" pageId="21" pageNumber="22">Malaise trap</collectingMethod>
|
||
, JSS52349 (
|
||
<specimenCount id="9D07FD4F5F57FF9CFDA0EF58FD824B04" box="[521,557,1050,1076]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED10AE035F57FF9CFD90EF58FDD84B04" box="[569,631,1050,1076]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="21" pageNumber="22">CNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B654D5F57FF9CFF6EEF7CFD844B69" box="[199,555,1086,1113]" pageId="21" pageNumber="22" type="distribution">
|
||
<paragraph id="8BBE36C65F57FF9CFF6EEF7CFD844B69" blockId="21.[151,1437,150,1113]" box="[199,555,1086,1113]" pageId="21" pageNumber="22">
|
||
<emphasis id="B975EAD45F57FF9CFF6EEF7CFECC4B68" bold="true" box="[199,355,1086,1112]" pageId="21" pageNumber="22">Distribution:</emphasis>
|
||
<collectingCountry id="F31676565F57FF9CFEC3EF7CFE144B68" box="[362,443,1086,1112]" name="Cyprus" pageId="21" pageNumber="22">Cyprus</collectingCountry>
|
||
(
|
||
<figureCitation id="133A2A435F57FF9CFE63EF7CFDB14B69" box="[458,542,1086,1113]" captionStart="FIGURE 50" captionStartId="85.[151,250,1898,1922]" captionTargetBox="[151,1436,1232,1874]" captionTargetId="figure-107@85.[151,1436,1232,1874]" captionTargetPageId="85" captionText="FIGURE 50. Distribution of Tomosvaryella angulata, T. argyrata, T. argyratoides, T. cyprusensis sp. nov. in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122771" httpUri="https://zenodo.org/record/5122771/files/figure.png" pageId="21" pageNumber="22">Fig. 50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |