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<document id="5EA83FA967AF3235021EE85807B386FF" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629668466086" checkinUser="felipe" docAuthor="Pelsue, Frank W." docDate="2010" docId="03A887BDF2366670FF45FBFFFA268322" docLanguage="en" docName="zt02384p052.pdf" docOrigin="Zootaxa 2384" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Balaninus analis Marshall 1920" docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="FF91FFC5F2376672FFD2FFEDFFAD824E" masterDocTitle="Description of a neotype and designation of four lectotypes from Borneo in the genus Curculio (Coleoptera: Curculionidae)" masterLastPageNumber="52" masterPageNumber="44" pageNumber="45" updateTime="1699093124427" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="44550BB8B5EBADCCF93738EC0429D219">
<mods:title id="643B1E656F6D98BA35DDDAF9251DEAAD">Description of a neotype and designation of four lectotypes from Borneo in the genus Curculio (Coleoptera: Curculionidae)</mods:title>
</mods:titleInfo>
<mods:name id="BFAAF466DAC2FC0170C841763F7A577F" type="personal">
<mods:role id="F07CE521B391E3708F1B7AE312123538">
<mods:roleTerm id="4A1B839BBB1CD18BE85F5357F26FC7F6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FB1B2254C89BA1466F5360063BB5E2A">Pelsue, Frank W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D93019428E7BF93F472A20FF4E91FB60">text</mods:typeOfResource>
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<mods:title id="582F32F89D4AE2CFBFE2135CB6D18382">Zootaxa</mods:title>
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<mods:date id="86FBBA09EDA5F277ED8DD54EF75D99B1">2010</mods:date>
<mods:detail id="0A39BE2977337E1B16B555215A692001" type="volume">
<mods:number id="2431B74C920BB623522475A350ACB505">2384</mods:number>
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<mods:classification id="AF30121BA80C47FD312C4B0E20C3C2BA">journal article</mods:classification>
<mods:identifier id="BA198B8BB3C0C8D7A46467AB7847C63F" type="ISSN">1175-5326</mods:identifier>
</mods:mods>
<treatment id="03A887BDF2366670FF45FBFFFA268322" ID-DOI="http://doi.org/10.5281/zenodo.5314966" ID-Zenodo-Dep="5314966" LSID="urn:lsid:plazi:treatment:03A887BDF2366670FF45FBFFFA268322" httpUri="http://treatment.plazi.org/id/03A887BDF2366670FF45FBFFFA268322" lastPageId="2" lastPageNumber="46" pageId="1" pageNumber="45">
<subSubSection id="C31B6520F2366673FF45FBFFFDF08662" box="[151,605,1042,1068]" pageId="1" pageNumber="45" type="nomenclature">
<paragraph id="8BBE36ABF2366673FF45FBFFFDF08662" blockId="1.[151,605,1042,1068]" box="[151,605,1042,1068]" pageId="1" pageNumber="45">
<heading id="D0F681C7F2366673FF45FBFFFDF08662" bold="true" box="[151,605,1042,1068]" fontSize="11" level="1" pageId="1" pageNumber="45" reason="1">
<treatmentCitationGroup id="AB111185F2366673FF45FBFFFDF08662" box="[151,605,1042,1068]" pageId="1" pageNumber="45">
<treatmentCitation id="0AA010BAF2366673FF45FBFFFDF08662" author="Marshall, G. A. K." box="[151,605,1042,1068]" page="387" pageId="1" pageNumber="45" year="1919">
<taxonomicName id="4C014D28F2366673FF45FBFFFDF08662" ID-CoL="KFCX" authority="Marshall 1919: 387" authorityName="Marshall" authorityPageNumber="387" authorityYear="1920" box="[151,605,1042,1068]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B975EAB9F2366673FF45FBFFFDF08662" bold="true" box="[151,605,1042,1068]" pageId="1" pageNumber="45">
<emphasis id="B975EAB9F2366673FF45FBFFFEC88662" bold="true" box="[151,357,1042,1068]" italics="true" pageId="1" pageNumber="45">Balaninus analis</emphasis>
<bibRefCitation id="EF904B5AF2366673FEBEFBFFFDF08662" author="Marshall, G. A. K." box="[364,605,1042,1068]" pageId="1" pageNumber="45" pagination="365 - 397" refId="ref3734" refString="Marshall, G. A. K. (1919) On the Species of Balaninus occurring in Borneo (Coleoptera, Curculionidae). Proceedings of the Zoological Society of London, 16, 365 - 397." type="journal article" year="1919">Marshall 1919: 387</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B6520F2366673FF45FBB7FBB38734" pageId="1" pageNumber="45" type="materials_examined">
<paragraph id="8BBE36ABF2366673FF45FBB7FEA786F1" blockId="1.[151,1436,1114,2036]" pageId="1" pageNumber="45">
<emphasis id="B975EAB9F2366673FF45FBB7FEF5863A" bold="true" box="[151,344,1114,1140]" pageId="1" pageNumber="45">Type material:</emphasis>
<materialsCitation id="3B693CF6F2366673FEB0FBB7FD0F86D4" collectorName="G. E. Bryant" country="Malaysia" location="Sarawak" pageId="1" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B975EAB9F2366673FEB0FBB7FE4E863A" bold="true" box="[354,483,1114,1140]" pageId="1" pageNumber="45">
<typeStatus id="54BA8809F2366673FEB0FBB7FE4E863A" box="[354,483,1114,1140]" pageId="1" pageNumber="45" type="lectotype">Lectotype</typeStatus>
</emphasis>
<specimenCount id="9D07FD22F2366673FE3FFBB7FDED863A" box="[493,576,1114,1140]" pageId="1" pageNumber="45" type="male">1 male</specimenCount>
(present designation) “Puak, /
<collectingCountry id="F316763BF2366673FC14FBB7FB83863A" box="[966,1070,1114,1140]" name="Malaysia" pageId="1" pageNumber="45">Sarawak</collectingCountry>
/
<collectorName id="26F4537DF2366673FB9BFBB7FB4E863A" box="[1097,1251,1114,1140]" pageId="1" pageNumber="45">G. E. Bryant</collectorName>
/ 30.v.14” and “
<taxonomicName id="4C014D28F2366673FF73FB6DFEB386D4" box="[161,286,1152,1178]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">Balaninus</taxonomicName>
/
<taxonomicName id="4C014D28F2366673FEEBFB6DFE7D86D4" authority="Mshl." authorityName="Marshall" authorityYear="1920" box="[313,464,1152,1178]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="analis">analis Mshl.</taxonomicName>
/ COTYPE” and
</materialsCitation>
<materialsCitation id="3B693CF6F2366673FD78FB6DFEAB86F1" collectionCode="BMNH" country="Malaysia" pageId="1" pageNumber="45" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54BA8809F2366673FD78FB6DFCFB86D4" box="[682,854,1152,1178]" pageId="1" pageNumber="45" type="lectotype">LECTOTYPE</typeStatus>
“
<taxonomicName id="4C014D28F2366673FCBFFB6DFB1B86D4" authority="(Marshall)" baseAuthorityName="Marshall" box="[877,1206,1152,1178]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B975EAB9F2366673FCBFFB6DFB8186D7" box="[877,1068,1152,1177]" italics="true" pageId="1" pageNumber="45">Curculio analis</emphasis>
(Marshall)
</taxonomicName>
des.
<bibRefCitation id="EF904B5AF2366673FB2BFB6DFA3C86D4" author="Pelsue, F. W." box="[1273,1425,1152,1178]" pageId="1" pageNumber="45" pagination="295 - 304" refId="ref3773" refString="Pelsue, F. W. (2009) Five new Species of the genus Indocurculio Panji et al., and three new combinations (Coleoptera: Curculionidae: Curculioninae). Koleopterologische Rundschau, 79, 295 - 304." type="journal article" year="2009">Pelsue 2009</bibRefCitation>
” (
<collectionCode id="ED10AE6EF2366673FF4DFB48FF5386F1" box="[159,254,1189,1215]" pageId="1" pageNumber="45">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBE36ABF2366673FF16FB27FCD586AA" blockId="1.[151,1436,1114,2036]" box="[196,888,1226,1252]" pageId="1" pageNumber="45">
<materialsCitation id="3B693CF6F2366673FF16FB27FCD586AA" box="[196,888,1226,1252]" collectorName="J. E. A. Lewis." pageId="1" pageNumber="45" specimenCount="4" specimenCount-male="4" typeStatus="syntype">
<typeStatus id="54BA8809F2366673FF16FB27FE9986AA" box="[196,308,1226,1252]" pageId="1" pageNumber="45" type="syntype">Syntypes</typeStatus>
:
<specimenCount id="9D07FD22F2366673FEEDFB27FE3486AA" box="[319,409,1226,1252]" pageId="1" pageNumber="45" type="male">4 males</specimenCount>
”Kuching. /
<collectorName id="26F4537DF2366673FDE2FB27FD7386AA" box="[560,734,1226,1252]" pageId="1" pageNumber="45">J. E. A. Lewis.</collectorName>
/ 1910-116.”
</materialsCitation>
</paragraph>
<paragraph id="8BBE36ABF2366673FF16FB1DFC598761" blockId="1.[151,1436,1114,2036]" pageId="1" pageNumber="45">
<materialsCitation id="3B693CF6F2366673FF16FB1DFC598761" collectorName="J. E. A. Lewis." pageId="1" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D07FD22F2366673FF16FB1DFE998744" box="[196,308,1264,1290]" pageId="1" pageNumber="45" type="female">1 female</specimenCount>
“
<typeStatus id="54BA8809F2366673FE98FB1DFE218744" box="[330,396,1264,1290]" pageId="1" pageNumber="45">Type</typeStatus>
/ (BM red-rimmed
<typeStatus id="54BA8809F2366673FD43FB1DFD658744" box="[657,712,1264,1290]" pageId="1" pageNumber="45">type</typeStatus>
disc)” and “Kuching. /
<collectorName id="26F4537DF2366673FBD7FB1DFB1B8744" box="[1029,1206,1264,1290]" pageId="1" pageNumber="45">J.E.A. Lewis.</collectorName>
/ 1910-116.” and “
<collectorName id="26F4537DF2366673FF70FAF8FE918761" box="[162,316,1301,1327]" pageId="1" pageNumber="45">J.E.A.Lewis.</collectorName>
/ 1910-116.” and “
<taxonomicName id="4C014D28F2366673FDCBFAF8FD3C8761" box="[537,657,1301,1327]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">Balaninus</taxonomicName>
/ analis / Mshl /
<typeStatus id="54BA8809F2366673FC84FAF8FC3D8761" box="[854,912,1301,1327]" pageId="1" pageNumber="45">Type</typeStatus>
female”
</materialsCitation>
</paragraph>
<paragraph id="8BBE36ABF2366673FF16FAD7FD53871A" blockId="1.[151,1436,1114,2036]" box="[196,766,1338,1364]" pageId="1" pageNumber="45">
<materialsCitation id="3B693CF6F2366673FF16FAD7FD53871A" box="[196,766,1338,1364]" collectorName="J. E. A. Lewis." pageId="1" pageNumber="45" specimenCount="8" specimenCount-female="8">
<specimenCount id="9D07FD22F2366673FF16FAD7FE98871A" box="[196,309,1338,1364]" pageId="1" pageNumber="45" type="female">8 females</specimenCount>
“Kuching. /
<collectorName id="26F4537DF2366673FE1EFAD7FDC9871A" box="[460,612,1338,1364]" pageId="1" pageNumber="45">J.E.A.Lewis.</collectorName>
/ 1910-116.”
</materialsCitation>
</paragraph>
<paragraph id="8BBE36ABF2366673FF16FA8DFBB38734" blockId="1.[151,1436,1114,2036]" box="[196,1054,1376,1402]" pageId="1" pageNumber="45">
<specimenCount id="9D07FD22F2366673FF16FA8DFE988734" box="[196,309,1376,1402]" pageId="1" pageNumber="45" type="female">4 females</specimenCount>
“Kuching / 11.5.09, and “Kuching. / J.E.A.Lewis. / 1910-116.”
</paragraph>
</subSubSection>
<subSubSection id="C31B6520F2366670FF16FA68FAC78309" lastPageId="2" lastPageNumber="46" pageId="1" pageNumber="45" type="diagnosis">
<paragraph id="8BBE36ABF2366673FF16FA68FD4B855A" blockId="1.[151,1436,1114,2036]" pageId="1" pageNumber="45">
<emphasis id="B975EAB9F2366673FF16FA68FEEB87D1" bold="true" box="[196,326,1413,1439]" pageId="1" pageNumber="45">Diagnosis:</emphasis>
Male 4.0–
<quantity id="4CF99B4EF2366673FE17FA68FDB087D1" box="[453,541,1413,1439]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="45" unit="mm" value="5.3">5.3 mm</quantity>
long; piceous except for legs which are fuscorufous; rostrum more than half the body length in male, but longer than body in female, gradually emerging from frons, evenly deflexed from base to apex; pronotum with anterior basolateral macula with arms reaching past dorsal margin of eye in lateral view and just visible in dorsal view, consisting of tightly packed recumbent, clavate, cretaceous scales; basal margin of pronotum with lateral macula of similar scales and in dorsal view line up with basal fascia on elytra forming V with piceous scutellum set in middle of V. Mesoventral intercoxal process prominent. Elytra with submedian fascia of recumbent, clavate, cretaceous scales and distal vittae of cretaceous scales at apex on interval 1 along margin of crosshatch seta-like scales; venter clothed with decumbent, clavate, cretaceous scales with 2 pronounced tufts of seta-like cretaceous scales on apex of ventrite 5 with apical margin emarginated; metafemur exceeding apex of elytra, with medium tooth; mesofemur with basal margin of long white seta-like scales appearing like fringe of hair.
</paragraph>
<paragraph id="8BBE36ABF2366670FF16F8F2FAC78309" blockId="1.[151,1436,1114,2036]" lastBlockId="2.[151,1436,152,364]" lastPageId="2" lastPageNumber="46" pageId="1" pageNumber="45">
<emphasis id="B975EAB9F2366673FF16F8F2FE708577" bold="true" box="[196,477,1823,1849]" pageId="1" pageNumber="45">Differential diagnosis:</emphasis>
This species can be confused with
<taxonomicName id="4C014D28F2366673FC55F8CDFB4A8574" authority="(Marshall)" baseAuthorityName="Marshall" box="[903,1255,1824,1850]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="deceptor">
<emphasis id="B975EAB9F2366673FC55F8CDFBF38577" box="[903,1118,1824,1849]" italics="true" pageId="1" pageNumber="45">Curculio deceptor</emphasis>
(Marshall)
</taxonomicName>
,
<taxonomicName id="4C014D28F2366673FB26F8CDFF558511" authority="(Kirsch)" baseAuthorityName="Kirsch" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="B975EAB9F2366673FB26F8CDFA318577" box="[1268,1436,1824,1849]" italics="true" pageId="1" pageNumber="45">C. interruptus</emphasis>
(Kirsch)
</taxonomicName>
and
<taxonomicName id="4C014D28F2366673FEE0F8A8FE1E8510" authorityName="Marshall" authorityYear="1920" box="[306,435,1861,1886]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="imitator">
<emphasis id="B975EAB9F2366673FEE0F8A8FE1E8510" box="[306,435,1861,1886]" italics="true" pageId="1" pageNumber="45">C. imitator</emphasis>
</taxonomicName>
due to the very similar fascia extending from the base of the pronotum to the base of the elytra forming a white V on a black background. It can be distinguished from
<taxonomicName id="4C014D28F2366673FBA2F886FB5685CA" box="[1136,1275,1899,1924]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="deceptor">
<emphasis id="B975EAB9F2366673FBA2F886FB5685CA" box="[1136,1275,1899,1924]" italics="true" pageId="1" pageNumber="45">C. deceptor</emphasis>
</taxonomicName>
by ventrite 5 having a pair of large tufts of hair on the apex and the second funicular antennomere as long as the first and appears less robust across the elytral humeri; mesofemur has a basal fringe of long setae forming a beard; basal macular median margin reaching interval 4, whereas in
<taxonomicName id="4C014D28F2366673FCBEF836FC5985BA" box="[876,1012,2011,2036]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="deceptor">
<emphasis id="B975EAB9F2366673FCBEF836FC5985BA" box="[876,1012,2011,2036]" italics="true" pageId="1" pageNumber="45">C. deceptor</emphasis>
</taxonomicName>
this reaches interval 5 of the elytra. It can be distinguished from
<taxonomicName id="4C014D28F2356670FE2BFF75FD0882FF" box="[505,677,152,177]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="B975EAB9F2356670FE2BFF75FD0882FF" box="[505,677,152,177]" italics="true" pageId="2" pageNumber="46">C. interruptus</emphasis>
</taxonomicName>
by the vestiture of the scutellum which is black in
<taxonomicName id="4C014D28F2356670FACFFF75FACA82FF" box="[1309,1383,152,177]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B975EAB9F2356670FACFFF75FACA82FF" box="[1309,1383,152,177]" italics="true" pageId="2" pageNumber="46">analis</emphasis>
</taxonomicName>
and clothed with white scales in
<taxonomicName id="4C014D28F2356670FE38FF50FDC08298" box="[490,621,189,214]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="B975EAB9F2356670FE38FF50FDC08298" box="[490,621,189,214]" italics="true" pageId="2" pageNumber="46">interruptus</emphasis>
</taxonomicName>
. With
<taxonomicName id="4C014D28F2356670FD6CFF50FCED8298" authorityName="Marshall" authorityYear="1920" box="[702,832,189,214]" class="Insecta" family="Curculionidae" genus="Balaninus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="imitator">
<emphasis id="B975EAB9F2356670FD6CFF50FCED8298" box="[702,832,189,214]" italics="true" pageId="2" pageNumber="46">C. imitator</emphasis>
</taxonomicName>
the size is smaller, the anterior basilateral macular arm reaches only to the middle of the eye and is not visible in dorsal view, metafemur reaches apex of elytra or only slightly exceeds it; the basal pronotal macular median margin reaches to elytral interval 4 on both male and female; the mesofemur lacks the basal fringe of long setae-like hair that is present in
<taxonomicName id="4C014D28F2356670FB7FFEC0FABA8308" box="[1197,1303,301,326]" class="Insecta" family="Curculionidae" genus="Curculio" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="46" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B975EAB9F2356670FB7FFEC0FABA8308" box="[1197,1303,301,326]" italics="true" pageId="2" pageNumber="46">C. analis</emphasis>
</taxonomicName>
males.
</paragraph>
</subSubSection>
<subSubSection id="C31B6520F2356670FF16FEBFFA268322" box="[196,1419,338,364]" pageId="2" pageNumber="46" type="distribution">
<paragraph id="8BBE36ABF2356670FF16FEBFFA268322" blockId="2.[151,1436,152,364]" box="[196,1419,338,364]" pageId="2" pageNumber="46">
<emphasis id="B975EAB9F2356670FF16FEBFFECB8322" bold="true" box="[196,358,338,364]" pageId="2" pageNumber="46">Distribution:</emphasis>
This species is only known from Malaysian Borneo:
<collectingCountry id="F316763BF2356670FC08FEBFFBEF8322" box="[986,1090,338,364]" name="Malaysia" pageId="2" pageNumber="46">Sarawak</collectingCountry>
,
<collectingRegion id="49C5F849F2356670FB9FFEBFFB358322" box="[1101,1176,338,364]" country="Malaysia" name="Sabah" pageId="2" pageNumber="46">Sabah</collectingRegion>
, Kuching, and Puak.
</paragraph>
</subSubSection>
</treatment>
</document>
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