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<document id="E5FCBAE9382B1F6226ECA8E58A05CF46" ID-DOI="10.5281/zenodo.214470" ID-GBIF-Dataset="36bfe153-09d4-48c1-be84-9ce1765005c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="214470" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460548210964" checkinUser="plazi" docAuthor="Gorochov, Andrej V. &amp; Tan, Ming Kai" docDate="2012" docId="03A887B9FFEEFFAA79AF689AFEE09C0D" docLanguage="en" docName="zt03525p034.pdf" docOrigin="Zootaxa 3525" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trellius (Protrellius) neesoon Gorochov et Tan, new species" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF91FFC1FFEFFFA979386934FF8B9F3A" masterDocTitle="New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore" masterLastPageNumber="34" masterPageNumber="18" pageNumber="19" updateTime="1698317987547" updateUser="plazi">
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<mods:title id="B505B7661201AB13BDA1F101D4D91371">New crickets of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from Singapore</mods:title>
</mods:titleInfo>
<mods:name id="3A383F5F14B7FB116CB8F0E33AE7A2E1" type="personal">
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<mods:namePart id="3B0B5061D959AAACAC8033F2E2B34481">Gorochov, Andrej V.</mods:namePart>
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<mods:namePart id="0DEFC48DCD9C6A4805C7B707439D540A">Tan, Ming Kai</mods:namePart>
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<mods:date id="65CAE8307A2344D7E4F58D93F92E277D">2012</mods:date>
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<treatment id="03A887B9FFEEFFAA79AF689AFEE09C0D" ID-DOI="http://doi.org/10.5281/zenodo.6178735" ID-GBIF-Taxon="119529245" ID-Zenodo-Dep="6178735" LSID="urn:lsid:plazi:treatment:03A887B9FFEEFFAA79AF689AFEE09C0D" httpUri="http://treatment.plazi.org/id/03A887B9FFEEFFAA79AF689AFEE09C0D" lastPageId="3" lastPageNumber="21" pageId="1" pageNumber="19">
<subSubSection id="C31B6524FFEEFFA879AF689AFE1C9ED3" pageId="1" pageNumber="19" type="nomenclature">
<paragraph id="8BBE36AFFFEEFFA879AF689AFCFB9EF2" blockId="1.[151,880,430,489]" box="[151,880,430,456]" pageId="1" pageNumber="19">
<heading id="D0F681C3FFEEFFA879AF689AFCFB9EF2" bold="true" box="[151,880,430,456]" fontSize="11" level="1" pageId="1" pageNumber="19" reason="1">
<emphasis id="B975EABDFFEEFFA879AF689AFCFB9EF2" bold="true" box="[151,880,430,456]" pageId="1" pageNumber="19">
<taxonomicName id="4C014D2CFFEEFFA879AF689AFD5F9EF2" authority="Gorochov et Tan" authorityName="Gorochov et Tan" box="[151,724,430,456]" class="Insecta" family="Haglotettigoniidae" genus="Trellius" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="species" species="neesoon" status="sp. nov." subGenus="Protrellius">
<emphasis id="B975EABDFFEEFFA879AF689AFF789EF2" bold="true" box="[151,243,430,456]" italics="true" pageId="1" pageNumber="19">Trellius</emphasis>
(
<emphasis id="B975EABDFFEEFFA8783D689AFE0D9EF2" bold="true" box="[261,390,430,456]" italics="true" pageId="1" pageNumber="19">Protrellius</emphasis>
)
<emphasis id="B975EABDFFEEFFA878AF689AFE719EF2" bold="true" box="[407,506,430,456]" italics="true" pageId="1" pageNumber="19">neesoon</emphasis>
Gorochov et Tan
</taxonomicName>
,
<taxonomicNameLabel id="A24657C6FFEEFFA87BD9689AFCFB9EF2" box="[737,880,430,456]" pageId="1" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE36AFFFEEFFA879AF68E5FE1C9ED3" blockId="1.[151,880,430,489]" box="[151,407,465,489]" pageId="1" pageNumber="19">
(
<figureCitation id="133A2A2AFFEEFFA879A768E5FE8F9ED3" box="[159,260,465,489]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Figs 16</figureCitation>
,
<figureCitation id="133A2A2AFFEEFFA8783768E6FEB09ED3" box="[271,315,465,489]" captionStart="FIGURES 7 16" captionStartId="4.[151,264,1397,1420]" captionTargetBox="[243,1339,214,1366]" captionTargetId="figure@4.[226,1360,193,1376]" captionTargetPageId="4" captionText="FIGURES 7 16. Phaloriinae and Pteroplistinae (schematically): 7, 8, Trellius neesoon sp. n. (7, holotype; 8, paratype); 9, 10, Tremellia timah sp. n. (9, paratype; 10, holotype); 11, 12, Phaloria? jerelynae sp. n., holotype; 13 16, Singapuriola separata sp. n., paratypes. Inner side of fore tibia (7, 9, 11, 13); male subgenital plate from below (8, 10, 12, 14); female subgenital plate with base of ovipositor from below (15); ovipositor from side (16). Scale bars 1 mm." httpUri="https://zenodo.org/record/214472/files/figure.png" pageId="1" pageNumber="19">7, 8</figureCitation>
,
<figureCitation id="133A2A2AFFEEFFA8787F68E5FE059ED3" box="[327,398,465,489]" captionStart="FIGURES 17 24" captionStartId="5.[151,264,1452,1475]" captionTargetBox="[250,1341,198,1406]" captionTargetId="figure@5.[226,1360,193,1431]" captionTargetPageId="5" captionText="FIGURES 17 24. Phaloriinae and Pteroplistinae, male (schematically): 17, 18, Trellius neesoon sp. n., holotype; 19, 20, Phaloria? jerelynae sp. n., paratype; 21 23, Singapuriola separata sp. n., paratype; 24, Tremellia timah sp. n., holotype. Genitalia from above (17, 19, 21) and from below (18, 20, 22); mold of spermatophore attachment plate from behind and slightly above (23); ectoparamere from below (24)." httpUri="https://zenodo.org/record/214473/files/figure.png" pageId="1" pageNumber="19">17, 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B6524FFEEFFA879AF6B2CFA019C33" pageId="1" pageNumber="19" type="materials_examined">
<paragraph id="8BBE36AFFFEEFFA879AF6B2CFB3A9D6F" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
<emphasis id="B975EABDFFEEFFA879AF6B2CFE029D0B" bold="true" box="[151,393,536,561]" pageId="1" pageNumber="19">Material examined.</emphasis>
<typeStatus id="54BA880DFFEEFFA878A06B2DFD8B9D0B" box="[408,512,537,561]" pageId="1" pageNumber="19" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B975EABDFFEEFFA87B576B2CFD639D0B" bold="true" box="[623,744,536,561]" pageId="1" pageNumber="19">
<collectingCountry id="F316763FFFEEFFA87B576B2CFD639D0B" box="[623,744,536,561]" name="Singapore" pageId="1" pageNumber="19">Singapore</collectingCountry>
</emphasis>
, Central Catchment Nature Reserve, Nee Soon pipeline, secondary forest, near primary swamp forest,
<date id="FFBF106FFFEEFFA87BA06B09FCBD9D6F" box="[664,822,573,597]" pageId="1" pageNumber="19" value="2011-04-20">20 April 2011</date>
, coll. M.K. Tan &amp; M.R.B. Ismail.
</paragraph>
<paragraph id="8BBE36AFFFEEFFA879FF6B56FA019C33" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
<typeStatus id="54BA880DFFEEFFA879FF6B56FEB39D40" box="[199,312,610,634]" pageId="1" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B975EABDFFEEFFA8787E6B55FE349D40" bold="true" box="[326,447,609,634]" pageId="1" pageNumber="19">
<collectingCountry id="F316763FFFEEFFA8787E6B55FE349D40" box="[326,447,609,634]" name="Singapore" pageId="1" pageNumber="19">Singapore</collectingCountry>
</emphasis>
:
<specimenCount id="9D07FD26FFEEFFA878E96B55FDA99D43" box="[465,546,609,633]" pageId="1" pageNumber="19" type="male">1 male</specimenCount>
, Bukit
<taxonomicName id="4C014D2CFFEEFFA87B426B55FD499D43" box="[634,706,609,633]" class="Insecta" family="Haglotettigoniidae" genus="Tremellia" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="timah" status="sp. nov.">Timah</taxonomicName>
Nature Reserve, degraded hill forest,
<date id="FFBF106FFFEEFFA87DB96B55FA909D43" box="[1153,1307,609,633]" pageId="1" pageNumber="19" value="1967-07-25">25 July 1967</date>
, coll. D.H. Murphy;
<specimenCount id="9D07FD26FFEEFFA8783C6BB1FEDD9DA7" box="[260,342,645,669]" pageId="1" pageNumber="19" type="male">1 male</specimenCount>
, same reserve, Jungle Fall Valley, along stream in secondary forest,
<date id="FFBF106FFFEEFFA87D466BB1FA929DA7" box="[1150,1305,645,669]" pageId="1" pageNumber="19" value="1979-07-18">18 July 1979</date>
, coll. D.H. Murphy;
<specimenCount id="9D07FD26FFEEFFA8783A6B9DFED99DFB" box="[258,338,681,705]" pageId="1" pageNumber="19" type="male">1 male</specimenCount>
, MacRitchie Catchment Area, Sime Road forest, under log,
<date id="FFBF106FFFEEFFA87D3E6B9DFB169DFB" box="[1030,1181,681,705]" pageId="1" pageNumber="19" value="1975-04-04">4 April 1975</date>
, coll. D.H. Murphy;
<specimenCount id="9D07FD26FFEEFFA87CB66B9DFF5B9DDF" pageId="1" pageNumber="19" type="male">1 male</specimenCount>
, labelled “new road by racecourse”, actual locality unknown but suspect to be along Old Upper Thomson Road bordering Central Catchment Nature Reserve, under bark of felled trees,
<date id="FFBF106FFFEEFFA87D316BC5FB229C33" box="[1033,1193,753,777]" pageId="1" pageNumber="19" value="1976-04-25">25 April 1976</date>
, coll. D.H. Murphy.
</paragraph>
</subSubSection>
<subSubSection id="C31B6524FFEEFFA879FF6A21FC8B9CDB" pageId="1" pageNumber="19" type="diagnosis">
<paragraph id="8BBE36AFFFEEFFA879FF6A21FC8B9CDB" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
<emphasis id="B975EABDFFEEFFA879FF6A21FECA9C14" bold="true" box="[199,321,789,814]" pageId="1" pageNumber="19">Diagnosis.</emphasis>
The new species is most similar to
<taxonomicName id="4C014D2CFFEEFFA87BE46A21FC789C17" authority="Gorochov" authorityName="Gorochov" box="[732,1011,789,813]" class="Insecta" family="Haglotettigoniidae" genus="Trellius" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="species" species="curup">
<emphasis id="B975EABDFFEEFFA87BE46A21FCF09C17" box="[732,891,789,813]" italics="true" pageId="1" pageNumber="19">Trellius curup</emphasis>
Gorochov
</taxonomicName>
from Sumatra (
<bibRefCitation id="EF904B5EFFEEFFA87D906A21FAE89C17" author="Gorochov" box="[1192,1379,789,813]" pageId="1" pageNumber="19" refString="Gorochov A. V. (2010) New cricket taxa of the subfamilies Phaloriinae and Pteroplistinae (Orthoptera: Gryllidae) from South- East Asia. Proceedings of the Zoological Institute RAS, 314 (2), 184 - 190." type="journal article" year="2010">Gorochov, 2010</bibRefCitation>
), but distinguished from it by the distinctly shorter dorsal lobules of epiphallus. From all the other species of this subgenus, the new species differs in the presence of short dorsal epiphallic lobules (versus absence of dorsal epiphallic lobules or presence of long spine-like processes instead them) in combination with the lateral epiphallic processes longer and/or narrower, additional processes and spinules on these processes absent, ectoparameres and guiding rod small, and distal part of this rod spine-like.
</paragraph>
</subSubSection>
<subSubSection id="C31B6524FFEEFFAB79FF6AD9FD8A98DC" lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="19" type="description">
<paragraph id="8BBE36AFFFEEFFA879FF6AD9FBC09B83" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
<emphasis id="B975EABDFFEEFFA879FF6AD9FED29B3C" bold="true" box="[199,345,1005,1030]" pageId="1" pageNumber="19">Description.</emphasis>
Male
<typeStatus id="54BA880DFFEEFFA878906AD9FD859B3F" box="[424,526,1005,1029]" pageId="1" pageNumber="19" type="holotype">holotype</typeStatus>
. General appearance typical of this genus (
<figureCitation id="133A2A2AFFEEFFA87D326AD9FBDB9B3F" box="[1034,1104,1005,1029]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 1</figureCitation>
). Head with frontal rostrum between antennal cavities approximately as wide as scape. Pronotum moderately transverse (1.3 times wider than long), weakly widening posteriorly (
<figureCitation id="133A2A2AFFEEFFA87B7F6D01FD059B77" box="[583,654,1077,1101]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 1</figureCitation>
). Fore tibiae with rather large, oval tympana on both side; inner tympanum slightly larger than outer one and hardly immersed (
<figureCitation id="133A2A2AFFEEFFA87A566D6DFC3F9B4B" box="[878,948,1113,1137]" captionStart="FIGURES 7 16" captionStartId="4.[151,264,1397,1420]" captionTargetBox="[243,1339,214,1366]" captionTargetId="figure@4.[226,1360,193,1376]" captionTargetPageId="4" captionText="FIGURES 7 16. Phaloriinae and Pteroplistinae (schematically): 7, 8, Trellius neesoon sp. n. (7, holotype; 8, paratype); 9, 10, Tremellia timah sp. n. (9, paratype; 10, holotype); 11, 12, Phaloria? jerelynae sp. n., holotype; 13 16, Singapuriola separata sp. n., paratypes. Inner side of fore tibia (7, 9, 11, 13); male subgenital plate from below (8, 10, 12, 14); female subgenital plate with base of ovipositor from below (15); ovipositor from side (16). Scale bars 1 mm." httpUri="https://zenodo.org/record/214472/files/figure.png" pageId="1" pageNumber="19">Fig. 7</figureCitation>
); hind tibiae with 4 pairs of long movable dorsal spines (length of longest inner distal spine
<quantity id="4CF99B4AFFEEFFA87BF06D49FCA99BAF" box="[712,802,1149,1173]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="19" unit="mm" value="2.6">2.6 mm</quantity>
), not very numerous small immovable spinules between them, and 6 long spurs (=apical spines) (length of longest inner dorsal spur
<quantity id="4CF99B4AFFEEFFA87ADD6D95FBB79B83" box="[997,1084,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="19" unit="mm" value="2.3">2.3 mm</quantity>
).
</paragraph>
<paragraph id="8BBE36AFFFEEFFA879FF6DF1FA099AAB" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
Tegmina reaching distal third of hind femora, with 7 oblique veins (2 proximal veins small) in dorsal field; mirror with 2 dividing veins gently converging medially (
<figureCitation id="133A2A2AFFEEFFA87A266DDDFCE89A3B" box="[798,867,1257,1281]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 2</figureCitation>
); lateral tegminal field with 18 branches of Sc and 9 crossveins between R and M (
<figureCitation id="133A2A2AFFEEFFA878C66C39FDC89A1F" box="[510,579,1293,1317]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 2</figureCitation>
); stridulatory vein of right tegmen short and sinuous as shown in
<figureCitation id="133A2A2AFFEEFFA87C1C6C39FAED9A1F" box="[1316,1382,1293,1317]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 3</figureCitation>
, and with 19 stridulatory teeth; hind wings distinctly exposed behind tegmina. Anal (=supra-anal) plate of abdomen simple but with median groove, lateral ridge, and apex truncate; subgenital plate rectangular (longer than wide), with apical part truncate and with median groove, as well as with shallow depression on distal half (as in
<figureCitation id="133A2A2AFFEEFFA87C096C4DFAFF9AAB" box="[1329,1396,1401,1425]" captionStart="FIGURES 7 16" captionStartId="4.[151,264,1397,1420]" captionTargetBox="[243,1339,214,1366]" captionTargetId="figure@4.[226,1360,193,1376]" captionTargetPageId="4" captionText="FIGURES 7 16. Phaloriinae and Pteroplistinae (schematically): 7, 8, Trellius neesoon sp. n. (7, holotype; 8, paratype); 9, 10, Tremellia timah sp. n. (9, paratype; 10, holotype); 11, 12, Phaloria? jerelynae sp. n., holotype; 13 16, Singapuriola separata sp. n., paratypes. Inner side of fore tibia (7, 9, 11, 13); male subgenital plate from below (8, 10, 12, 14); female subgenital plate with base of ovipositor from below (15); ovipositor from side (16). Scale bars 1 mm." httpUri="https://zenodo.org/record/214472/files/figure.png" pageId="1" pageNumber="19">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BBE36AFFFEEFFA879FF6CA9FEFA99EF" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
Genitalia (
<figureCitation id="133A2A2AFFEEFFA878056CA9FE2F9A8F" box="[317,420,1437,1461]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Figs 46</figureCitation>
,
<figureCitation id="133A2A2AFFEEFFA878886CA9FE719A8F" box="[432,506,1437,1461]" captionStart="FIGURES 17 24" captionStartId="5.[151,264,1452,1475]" captionTargetBox="[250,1341,198,1406]" captionTargetId="figure@5.[226,1360,193,1431]" captionTargetPageId="5" captionText="FIGURES 17 24. Phaloriinae and Pteroplistinae, male (schematically): 17, 18, Trellius neesoon sp. n., holotype; 19, 20, Phaloria? jerelynae sp. n., paratype; 21 23, Singapuriola separata sp. n., paratype; 24, Tremellia timah sp. n., holotype. Genitalia from above (17, 19, 21) and from below (18, 20, 22); mold of spermatophore attachment plate from behind and slightly above (23); ectoparamere from below (24)." httpUri="https://zenodo.org/record/214473/files/figure.png" pageId="1" pageNumber="19">17, 18</figureCitation>
): each posterolateral part of epiphallus with rather short ventral spine, long lateral process directed backwards and having narrowly rounded apex, two dorsal lobules (one directed backwards and slightly bilobate at apex, and one directed partly medially and with rounded apex), and medial ribbon almost reaching small median sclerite of epiphallus; last sclerite with rather wide lateral apodemes and small posteromedian spine; ectoparameres small and with rather thin posterior part; endoparameres with posterolateral arms fused with ectoparameres, wide (long) median bridge, and rather short anterior apodemes; guiding rod rather small, with oval plate-like proximal part and with spine-like distal part situated just under posteromedian spine of median epiphallic sclerite; mold of spermatophore attachment plate not wide, but its apodeme long and directed backwards (
<figureCitation id="133A2A2AFFEEFFA878266F89FEE899EF" box="[286,355,1725,1749]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[234,1351,205,1766]" captionTargetId="figure@2.[226,1361,193,1782]" captionTargetPageId="2" captionText="FIGURES 1 6. Trellius neesoon sp. n. (Phaloriinae), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 4 6, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3." httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="1" pageNumber="19">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BBE36AFFFEEFFA879FF6FD5FD8F9897" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">Colouration. Brown with following marks: head with dark patterns on vertex, with median and lateral ocelli white, and with mouthparts light brown; fore and middle femora light brown with 2 brown bands (one near middle, and one near knee); fore and middle tibiae with 2 whitish bands (one just below tympanum, and one near apex); hind tibiae with numerous light brown bands (some of which situated between spurs); tegminal lateral field with membranes between R and M light yellowish brown, and with membranes between Sc branches transparent; abdomen with cerci light brown.</paragraph>
<paragraph id="8BBE36AFFFEEFFA879FF6E8DFB9E98CF" blockId="1.[151,1436,536,2037]" pageId="1" pageNumber="19">
Variations. Right tegmina of
<typeStatus id="54BA880DFFEEFFA87B366E8EFDF198E8" box="[526,634,1978,2002]" pageId="1" pageNumber="19" type="paratype">paratypes</typeStatus>
with 67 oblique veins (2 proximal veins small) in dorsal field; lateral tegminal field with 1820 branches of Sc and 89 crossveins between R and M.
</paragraph>
<caption id="DF7E6627FFEDFFAB79AF6E3FFEA89847" httpUri="https://zenodo.org/record/214471/files/figure.png" pageId="2" pageNumber="20" targetBox="[234,1351,205,1766]" targetPageId="2">
<paragraph id="8BBE36AFFFEDFFAB79AF6E3FFEA89847" blockId="2.[151,1436,1803,1917]" pageId="2" pageNumber="20">
<emphasis id="B975EABDFFEDFFAB79AF6E3FFECB981B" bold="true" box="[151,320,1803,1826]" pageId="2" pageNumber="20">FIGURES 16.</emphasis>
<taxonomicName id="4C014D2CFFEDFFAB78746E38FE739818" box="[332,504,1804,1826]" class="Insecta" family="Haglotettigoniidae" genus="Trellius" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="neesoon" status="sp. nov.">
<emphasis id="B975EABDFFEDFFAB78746E38FE739818" box="[332,504,1804,1826]" italics="true" pageId="2" pageNumber="20">Trellius neesoon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657C6FFEDFFAB7B3C6E39FDCA9818" box="[516,577,1805,1826]" pageId="2" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
(
<taxonomicName id="4C014D2CFFEDFFAB7B6C6E3FFD46981B" box="[596,717,1803,1825]" class="Insecta" family="Phalangopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Phaloriinae">Phaloriinae</taxonomicName>
), male holotype: 1, general view from above; 2, right tegmen; 3, stridulatory vein of right tegmen from below; 46, genitalia and mold of spermatophore attachment plate (different photographs combined) from above (4); genitalia without this mold from below (5) and from side (6). Scale bars 5 mm for FIGURES 1, 2; 1 mm for Fig 3.
</paragraph>
</caption>
<paragraph id="8BBE36AFFFEDFFAB79FF6E9EFE1698F8" blockId="2.[199,513,1962,2022]" box="[199,413,1962,1986]" pageId="2" pageNumber="20">Female. Unknown.</paragraph>
<paragraph id="8BBE36AFFFEDFFAB79FF6EFBFD8A98DC" blockId="2.[199,513,1962,2022]" box="[199,513,1998,2022]" pageId="2" pageNumber="20">
Measurements. See
<tableCitation id="C6830314FFEDFFAB789F6EFAFE7098DC" box="[423,507,1998,2022]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetBox="[165,1418,209,661]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Trellius (Protrellius) neesoon (in mm, mean values in brackets)." httpUri="http://table.plazi.org/id/DF7E6627FFECFFAA79AF69A3FBF19F95" pageId="2" pageNumber="20" tableUuid="DF7E6627FFECFFAA79AF69A3FBF19F95">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF7E6627FFECFFAA79AF69A3FBF19F95" ID-Table-UUID="DF7E6627FFECFFAA79AF69A3FBF19F95" box="[151,1146,151,175]" httpUri="http://table.plazi.org/id/DF7E6627FFECFFAA79AF69A3FBF19F95" pageId="3" pageNumber="21" targetBox="[165,1418,209,661]" targetIsTable="true" targetPageId="3">
<paragraph id="8BBE36AFFFECFFAA79AF69A3FBF19F95" blockId="3.[151,1146,151,175]" box="[151,1146,151,175]" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA79AF69A3FE819F95" bold="true" box="[151,266,151,175]" pageId="3" pageNumber="21">TABLE 1.</emphasis>
Measurements of
<taxonomicName id="4C014D2CFFECFFAA78F769ACFC839F95" box="[463,776,152,175]" class="Insecta" family="Haglotettigoniidae" genus="Trellius" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="neesoon" subGenus="Protrellius">
<emphasis id="B975EABDFFECFFAA78F769ACFD959F95" box="[463,542,152,175]" italics="true" pageId="3" pageNumber="21">Trellius</emphasis>
(
<emphasis id="B975EABDFFECFFAA7B1769ACFD2A9F95" box="[559,673,152,175]" italics="true" pageId="3" pageNumber="21">Protrellius</emphasis>
)
<emphasis id="B975EABDFFECFFAA7B8969ACFC839F95" box="[689,776,152,175]" italics="true" pageId="3" pageNumber="21">neesoon</emphasis>
</taxonomicName>
(in mm, mean values in brackets).
</paragraph>
</caption>
<paragraph id="8BBE36AFFFECFFAA78A569E5FA019D49" pageId="3" pageNumber="21">
<table id="F901C40FFFEC0056799D69E5FA019DAF" box="[165,1418,209,661]" gridcols="9" gridrows="7" pageId="3" pageNumber="21">
<tr id="353134EDFFEC0056799D69E5FA019FDC" box="[165,1418,209,230]" gridrow="0" pageId="3" pageNumber="21" rowspan-0="1">
<th id="76E05D91FFEC0056784169E5FE559FDC" box="[377,478,209,230]" gridcol="1" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA78A569E5FE359FDC" bold="true" box="[413,446,209,230]" pageId="3" pageNumber="21">BL</emphasis>
</th>
<th id="76E05D91FFEC00567B3869E5FDEF9FDC" box="[512,612,209,230]" gridcol="2" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7B2F69E5FDD99FDC" bold="true" box="[535,594,209,230]" pageId="3" pageNumber="21">BWL</emphasis>
</th>
<th id="76E05D91FFEC00567BBE69E5FD5A9FDC" box="[646,721,209,230]" gridcol="3" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7B9269E5FD419FDC" bold="true" box="[682,714,209,230]" pageId="3" pageNumber="21">PL</emphasis>
</th>
<th id="76E05D91FFEC00567A3469E5FCD39FDC" box="[780,856,209,230]" gridcol="4" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7A1469E5FCDF9FDC" bold="true" box="[812,852,209,230]" pageId="3" pageNumber="21">PW</emphasis>
</th>
<th id="76E05D91FFEC00567AAA69E5FC7C9FDC" box="[914,1015,209,230]" gridcol="5" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7A8E69E5FC539FDC" bold="true" box="[950,984,209,230]" pageId="3" pageNumber="21">TL</emphasis>
</th>
<th id="76E05D91FFEC00567D2069E5FBEF9FDC" box="[1048,1124,209,230]" gridcol="6" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7D0F69E5FBEA9FDC" bold="true" box="[1079,1121,209,230]" pageId="3" pageNumber="21">TW</emphasis>
</th>
<th id="76E05D91FFEC00567DA669E5FA889FDC" box="[1182,1283,209,230]" gridcol="7" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7D8169E5FB669FDC" bold="true" box="[1209,1261,209,230]" pageId="3" pageNumber="21">HFL</emphasis>
</th>
<th id="76E05D91FFEC00567C1C69E5FA019FDC" box="[1316,1418,209,230]" gridcol="8" gridrow="0" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA7C0669E5FAF89FDC" bold="true" box="[1342,1395,209,230]" pageId="3" pageNumber="21">HTL</emphasis>
</th>
</tr>
<tr id="353134EDFFEC0056799D69C8FA019E28" box="[165,1418,252,274]" gridrow="1" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D69C8FEDD9E28" box="[165,342,252,274]" gridcol="0" gridrow="1" pageId="3" pageNumber="21">Male holotype</th>
<td id="76E05D91FFEC0056784169C8FE559E28" box="[377,478,252,274]" gridcol="1" gridrow="1" pageId="3" pageNumber="21">13.5</td>
<td id="76E05D91FFEC00567B3869C8FDEF9E28" box="[512,612,252,274]" gridcol="2" gridrow="1" pageId="3" pageNumber="21">24.2</td>
<td id="76E05D91FFEC00567BBE69C8FD5A9E28" box="[646,721,252,274]" gridcol="3" gridrow="1" pageId="3" pageNumber="21">2.9</td>
<td id="76E05D91FFEC00567A3469C8FCD39E28" box="[780,856,252,274]" gridcol="4" gridrow="1" pageId="3" pageNumber="21">3.9</td>
<td id="76E05D91FFEC00567AAA69C8FC7C9E28" box="[914,1015,252,274]" gridcol="5" gridrow="1" pageId="3" pageNumber="21">17.4</td>
<td id="76E05D91FFEC00567D2069C8FBEF9E28" box="[1048,1124,252,274]" gridcol="6" gridrow="1" pageId="3" pageNumber="21">-</td>
<td id="76E05D91FFEC00567DA669C8FA889E28" box="[1182,1283,252,274]" gridcol="7" gridrow="1" pageId="3" pageNumber="21">11.7</td>
<td id="76E05D91FFEC00567C1C69C8FA019E28" box="[1316,1418,252,274]" gridcol="8" gridrow="1" pageId="3" pageNumber="21">11.5</td>
</tr>
<tr id="353134EDFFEC0056799D681CFA019E65" box="[165,1418,296,351]" gridrow="2" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D681CFEDD9E65" box="[165,342,296,351]" gridcol="0" gridrow="2" pageId="3" pageNumber="21">Male paratype 25 Jul.1967</th>
<td id="76E05D91FFEC00567841681CFE559E65" box="[377,478,296,351]" gridcol="1" gridrow="2" pageId="3" pageNumber="21">13.0</td>
<td id="76E05D91FFEC00567B38681CFDEF9E65" box="[512,612,296,351]" gridcol="2" gridrow="2" pageId="3" pageNumber="21">22.7</td>
<td id="76E05D91FFEC00567BBE681CFD5A9E65" box="[646,721,296,351]" gridcol="3" gridrow="2" pageId="3" pageNumber="21">2.8</td>
<td id="76E05D91FFEC00567A34681CFCD39E65" box="[780,856,296,351]" gridcol="4" gridrow="2" pageId="3" pageNumber="21">4.0</td>
<td id="76E05D91FFEC00567AAA681CFC7C9E65" box="[914,1015,296,351]" gridcol="5" gridrow="2" pageId="3" pageNumber="21">15.2</td>
<td id="76E05D91FFEC00567D20681CFBEF9E65" box="[1048,1124,296,351]" gridcol="6" gridrow="2" pageId="3" pageNumber="21">6.0</td>
<td id="76E05D91FFEC00567DA6681CFA889E65" box="[1182,1283,296,351]" gridcol="7" gridrow="2" pageId="3" pageNumber="21">11.3</td>
<td id="76E05D91FFEC00567C1C681CFA019E65" box="[1316,1418,296,351]" gridcol="8" gridrow="2" pageId="3" pageNumber="21">11.1</td>
</tr>
<tr id="353134EDFFEC0056799D6841FA019E97" box="[165,1418,373,429]" gridrow="3" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D6841FEDD9E97" box="[165,342,373,429]" gridcol="0" gridrow="3" pageId="3" pageNumber="21">Male paratype 4 Apr.1975</th>
<td id="76E05D91FFEC005678416841FE559E97" box="[377,478,373,429]" gridcol="1" gridrow="3" pageId="3" pageNumber="21">12.5</td>
<td id="76E05D91FFEC00567B386841FDEF9E97" box="[512,612,373,429]" gridcol="2" gridrow="3" pageId="3" pageNumber="21">23.1</td>
<td id="76E05D91FFEC00567BBE6841FD5A9E97" box="[646,721,373,429]" gridcol="3" gridrow="3" pageId="3" pageNumber="21">3.0</td>
<td id="76E05D91FFEC00567A346841FCD39E97" box="[780,856,373,429]" gridcol="4" gridrow="3" pageId="3" pageNumber="21">4.1</td>
<td id="76E05D91FFEC00567AAA6841FC7C9E97" box="[914,1015,373,429]" gridcol="5" gridrow="3" pageId="3" pageNumber="21">16.9</td>
<td id="76E05D91FFEC00567D206841FBEF9E97" box="[1048,1124,373,429]" gridcol="6" gridrow="3" pageId="3" pageNumber="21">-</td>
<td id="76E05D91FFEC00567DA66841FA889E97" box="[1182,1283,373,429]" gridcol="7" gridrow="3" pageId="3" pageNumber="21">12.2</td>
<td id="76E05D91FFEC00567C1C6841FA019E97" box="[1316,1418,373,429]" gridcol="8" gridrow="3" pageId="3" pageNumber="21">11.8</td>
</tr>
<tr id="353134EDFFEC0056799D68F6FA019EC0" box="[165,1418,450,506]" gridrow="4" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D68F6FEDD9EC0" box="[165,342,450,506]" gridcol="0" gridrow="4" pageId="3" pageNumber="21">Male paratype 25 Apr.1976</th>
<td id="76E05D91FFEC0056784168F6FE559EC0" box="[377,478,450,506]" gridcol="1" gridrow="4" pageId="3" pageNumber="21">13.3</td>
<td id="76E05D91FFEC00567B3868F6FDEF9EC0" box="[512,612,450,506]" gridcol="2" gridrow="4" pageId="3" pageNumber="21">23.5</td>
<td id="76E05D91FFEC00567BBE68F6FD5A9EC0" box="[646,721,450,506]" gridcol="3" gridrow="4" pageId="3" pageNumber="21">2.9</td>
<td id="76E05D91FFEC00567A3468F6FCD39EC0" box="[780,856,450,506]" gridcol="4" gridrow="4" pageId="3" pageNumber="21">4.0</td>
<td id="76E05D91FFEC00567AAA68F6FC7C9EC0" box="[914,1015,450,506]" gridcol="5" gridrow="4" pageId="3" pageNumber="21">16.0</td>
<td id="76E05D91FFEC00567D2068F6FBEF9EC0" box="[1048,1124,450,506]" gridcol="6" gridrow="4" pageId="3" pageNumber="21">6.6</td>
<td id="76E05D91FFEC00567DA668F6FA889EC0" box="[1182,1283,450,506]" gridcol="7" gridrow="4" pageId="3" pageNumber="21">11.8</td>
<td id="76E05D91FFEC00567C1C68F6FA019EC0" box="[1316,1418,450,506]" gridcol="8" gridrow="4" pageId="3" pageNumber="21">-</td>
</tr>
<tr id="353134EDFFEC0056799D6B24FA019D7D" box="[165,1418,528,583]" gridrow="5" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D6B24FEDD9D7D" box="[165,342,528,583]" gridcol="0" gridrow="5" pageId="3" pageNumber="21">Male paratype 18 Jul.1979</th>
<td id="76E05D91FFEC005678416B24FE559D7D" box="[377,478,528,583]" gridcol="1" gridrow="5" pageId="3" pageNumber="21">12.4</td>
<td id="76E05D91FFEC00567B386B24FDEF9D7D" box="[512,612,528,583]" gridcol="2" gridrow="5" pageId="3" pageNumber="21">23.3</td>
<td id="76E05D91FFEC00567BBE6B24FD5A9D7D" box="[646,721,528,583]" gridcol="3" gridrow="5" pageId="3" pageNumber="21">2.9</td>
<td id="76E05D91FFEC00567A346B24FCD39D7D" box="[780,856,528,583]" gridcol="4" gridrow="5" pageId="3" pageNumber="21">3.6</td>
<td id="76E05D91FFEC00567AAA6B24FC7C9D7D" box="[914,1015,528,583]" gridcol="5" gridrow="5" pageId="3" pageNumber="21">15.6</td>
<td id="76E05D91FFEC00567D206B24FBEF9D7D" box="[1048,1124,528,583]" gridcol="6" gridrow="5" pageId="3" pageNumber="21">6.2</td>
<td id="76E05D91FFEC00567DA66B24FA889D7D" box="[1182,1283,528,583]" gridcol="7" gridrow="5" pageId="3" pageNumber="21">11.1</td>
<td id="76E05D91FFEC00567C1C6B24FA019D7D" box="[1316,1418,528,583]" gridcol="8" gridrow="5" pageId="3" pageNumber="21">10.8</td>
</tr>
<tr id="353134EDFFEC0056799D6B69FA019DAF" box="[165,1418,605,661]" gridrow="6" pageId="3" pageNumber="21">
<th id="76E05D91FFEC0056799D6B69FEDD9DAF" box="[165,342,605,661]" gridcol="0" gridrow="6" pageId="3" pageNumber="21">Males (n = 5)</th>
<td id="76E05D91FFEC005678416B69FE559DAF" box="[377,478,605,661]" gridcol="1" gridrow="6" pageId="3" pageNumber="21">12.413.5 (12.9)</td>
<td id="76E05D91FFEC00567B386B69FDEF9DAF" box="[512,612,605,661]" gridcol="2" gridrow="6" pageId="3" pageNumber="21">22.724.2 (23.4)</td>
<td id="76E05D91FFEC00567BBE6B69FD5A9DAF" box="[646,721,605,661]" gridcol="3" gridrow="6" pageId="3" pageNumber="21">2.83.0 (2.9)</td>
<td id="76E05D91FFEC00567A346B69FCD39DAF" box="[780,856,605,661]" gridcol="4" gridrow="6" pageId="3" pageNumber="21">3.64.1 (3.9)</td>
<td id="76E05D91FFEC00567AAA6B69FC7C9DAF" box="[914,1015,605,661]" gridcol="5" gridrow="6" pageId="3" pageNumber="21">15.217.4 (16.2)</td>
<td id="76E05D91FFEC00567D206B69FBEF9DAF" box="[1048,1124,605,661]" gridcol="6" gridrow="6" pageId="3" pageNumber="21">6.06.6</td>
<td id="76E05D91FFEC00567DA66B69FA889DAF" box="[1182,1283,605,661]" gridcol="7" gridrow="6" pageId="3" pageNumber="21">11.112.2 (11.6)</td>
<td id="76E05D91FFEC00567C1C6B69FA019DAF" box="[1316,1418,605,661]" gridcol="8" gridrow="6" pageId="3" pageNumber="21">10.811.8</td>
</tr>
</table>
</paragraph>
<subSubSection id="C31B6524FFECFFAA79FF6BE3FEE09C0D" pageId="3" pageNumber="21" type="etymology">
<paragraph id="8BBE36AFFFECFFAA79FF6BE3FEE09C0D" blockId="3.[151,1437,727,823]" pageId="3" pageNumber="21">
<emphasis id="B975EABDFFECFFAA79FF6BE3FEC59DD5" bold="true" box="[199,334,727,751]" pageId="3" pageNumber="21">Etymology.</emphasis>
The species is named after
<typeStatus id="54BA880DFFECFFAA7BB06BECFD339DCA" box="[648,696,728,752]" pageId="3" pageNumber="21">type</typeStatus>
locality, environs of Nee Soon pipeline. Nee Soon pipeline refers to a trail within the Central Catchment Nature Reserve, which is located adjacent to Singapores remaining primary freshwater swamp.
</paragraph>
</subSubSection>
</treatment>
</document>