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<document id="A301CDC72EF1E96AE42F2A0F6084F6A3" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A96B196FAFE9FEA6445F6048" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus cracentis Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="71" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="916C117C53E044C1C0B0B5D8FA45631B">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="03A887A1A96B196FAFE9FEA6445F6048" ID-DOI="http://doi.org/10.5281/zenodo.4417466" ID-GBIF-Taxon="176581836" ID-Zenodo-Dep="4417466" LSID="urn:lsid:plazi:treatment:03A887A1A96B196FAFE9FEA6445F6048" httpUri="http://treatment.plazi.org/id/03A887A1A96B196FAFE9FEA6445F6048" lastPageId="11" lastPageNumber="72" pageId="10" pageNumber="71">
<subSubSection id="C31B653CA96B196EAFE9FEA6453B65BD" box="[151,560,366,393]" pageId="10" pageNumber="71" type="nomenclature">
<paragraph id="8BBE36B7A96B196EAFE9FEA6453B65BD" blockId="10.[151,560,366,429]" box="[151,560,366,393]" pageId="10" pageNumber="71">
<heading id="D0F681DBA96B196EAFE9FEA6453B65BD" bold="true" box="[151,560,366,393]" fontSize="11" level="1" pageId="10" pageNumber="71" reason="1">
<emphasis id="B975EAA5A96B196EAFE9FEA6453B65BD" bold="true" box="[151,560,366,393]" pageId="10" pageNumber="71">
<taxonomicName id="4C014D34A96B196EAFE9FEA6469A65BC" ID-CoL="8Y4PS" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,401,366,393]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="species" species="cracentis" status="sp. nov.">
<emphasis id="B975EAA5A96B196EAFE9FEA6469A65BC" bold="true" box="[151,401,366,393]" italics="true" pageId="10" pageNumber="71">Hydrochus cracentis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA96B196EAEDEFEA7453B65BD" box="[416,560,366,393]" pageId="10" pageNumber="71" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B653CA96B196EAFE9FE5B47F46598" box="[151,255,403,429]" pageId="10" pageNumber="71" type="description">
<paragraph id="8BBE36B7A96B196EAFE9FE5B47F46598" blockId="10.[151,560,366,429]" box="[151,255,403,429]" pageId="10" pageNumber="71">
<figureCitation id="133A2A32A96B196EAFE9FE5B47E86598" box="[151,227,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="10" pageNumber="71">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A96B196EAF8FFE5B47F46598" box="[241,255,403,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,2004,2028]" captionTargetBox="[164,1421,461,1966]" captionTargetId="figure-66@10.[151,1435,453,1981]" captionTargetPageId="10" captionText="FIGURE 7. Hydrochus cracentis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417297" httpUri="https://zenodo.org/record/4417297/files/figure.png" pageId="10" pageNumber="71">7</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF7E663FA96B196EAFE9F81C44C963D9" ID-DOI="http://doi.org/10.5281/zenodo.4417297" ID-Zenodo-Dep="4417297" box="[151,962,2004,2028]" httpUri="https://zenodo.org/record/4417297/files/figure.png" pageId="10" pageNumber="71" startId="10.[151,250,2004,2028]" targetBox="[164,1421,461,1966]" targetPageId="10">
<paragraph id="8BBE36B7A96B196EAFE9F81C44C963D9" blockId="10.[151,962,2004,2028]" box="[151,962,2004,2028]" pageId="10" pageNumber="71">
<emphasis id="B975EAA5A96B196EAFE9F81C461863D9" bold="true" box="[151,275,2004,2028]" pageId="10" pageNumber="71">FIGURE 7.</emphasis>
<taxonomicName id="4C014D34A96B196EAE67F81C46E063D9" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,491,2004,2028]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="species" species="cracentis" status="sp. nov.">
<emphasis id="B975EAA5A96B196EAE67F81C46E063D9" box="[281,491,2004,2028]" italics="true" pageId="10" pageNumber="71">Hydrochus cracentis</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A96B196EAE8FF81C452163D9" bold="true" box="[497,554,2004,2028]" pageId="10" pageNumber="71">
<taxonomicNameLabel id="A24657DEA96B196EAE8FF81C452163D9" box="[497,554,2004,2028]" pageId="10" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<subSubSection id="C31B653CA96A196FAFE9FF5F464F64CC" pageId="11" pageNumber="72" type="materials_examined">
<paragraph id="8BBE36B7A96A196FAFE9FF5F464F64CC" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFE9FF5F46426485" bold="true" box="[151,329,150,177]" pageId="11" pageNumber="72">Type Material.</emphasis>
<materialsCitation id="3B693CEAA96A196FAE2CFF5F464F64CC" ID-GBIF-Occurrence="3015416349" collectingDate="1989-06-21" collectionCode="SEMC" collectorName="J. S. Ashe &amp; R. A. Leschen" country="Peru" county="Tambopata Prov." elevation="200" latitude="-12.55" location="Reserva" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" pageId="11" pageNumber="72" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B975EAA5A96A196FAE2CFF5F45176485" bold="true" box="[338,540,150,177]" pageId="11" pageNumber="72">
<typeStatus id="54BA8815A96A196FAE2CFF5F46B46484" box="[338,447,151,177]" pageId="11" pageNumber="72" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A96A196FAD51FF5F45766484" box="[559,637,151,177]" name="Peru" pageId="11" pageNumber="72">PERU</collectingCountry>
:
<collectingCounty id="62DF4E3BA96A196FADF6FF5F44416484" box="[648,842,151,177]" pageId="11" pageNumber="72">Tambopata Prov.</collectingCounty>
,
<collectingRegion id="49C5F855A96A196FAC28FF5F44F46484" box="[854,1023,151,177]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="72">Madre de Dios</collectingRegion>
<collectingMunicipality id="6BDAACCDA96A196FAB79FF5F43496484" box="[1031,1090,151,177]" pageId="11" pageNumber="72">Dpto</collectingMunicipality>
,
<locationDeviation id="9A0C2F95A96A196FAB31FF5E42936484" box="[1103,1432,150,177]" location="Puerto Maldonado" pageId="11" pageNumber="72">
<quantity id="4CF99B52A96A196FAB31FF5E43846484" box="[1103,1167,150,177]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="72" unit="km" value="15.0">15km</quantity>
NE Puerto Maldonado
</locationDeviation>
,
<location id="8EDE606CA96A196FAFE9FF7347FA64E0" LSID="urn:lsid:plazi:treatment:03A887A1A96B196FAFE9FEA6445F6048:8EDE606CA96A196FAFE9FF7347FA64E0" box="[151,241,187,213]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Reserva" pageId="11" pageNumber="72" stateProvince="Madre de Dios">Reserva</location>
<collectingRegion id="49C5F855A96A196FAF87FF73464A64E0" box="[249,321,187,213]" country="Peru" name="Cusco" pageId="11" pageNumber="72">Cuzco</collectingRegion>
<location id="8EDE606CA96A196FAE39FF7346C064E0" LSID="urn:lsid:plazi:treatment:03A887A1A96B196FAFE9FEA6445F6048:8EDE606CA96A196FAE39FF7346C064E0" box="[327,459,187,213]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Amazonico" pageId="11" pageNumber="72" stateProvince="Madre de Dios">Amazonico</location>
,
<geoCoordinate id="EE355070A96A196FAEA8FF73453F64E0" box="[470,564,187,213]" degrees="12" direction="south" minutes="33" orientation="latitude" pageId="11" pageNumber="72" precision="925" value="-12.55">12°33S</geoCoordinate>
,
<geoCoordinate id="EE355070A96A196FAD3FFF73459164E0" box="[577,666,187,213]" degrees="69" direction="west" minutes="3" orientation="longitude" pageId="11" pageNumber="72" precision="925" value="-69.05">69°3W</geoCoordinate>
,
<quantity id="4CF99B52A96A196FADD8FF7345E264E0" box="[678,745,187,213]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="72" unit="m" value="200.0">
<elevation id="002CD184A96A196FADD8FF7345E264E0" box="[678,745,187,213]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="72" unit="m" value="200.0">200m</elevation>
</quantity>
, camp,
<date id="FFBF1077A96A196FAC3EFF7344D264E0" box="[832,985,187,213]" pageId="11" pageNumber="72" value="1989-06-21">
<collectingDate id="EFFBE99FA96A196FAC3EFF7344D264E0" box="[832,985,187,213]" pageId="11" pageNumber="72" value="1989-06-21">21 June 1989</collectingDate>
</date>
,
<collectorName id="26F45361A96A196FAC9BFF73435C64E0" box="[997,1111,187,213]" pageId="11" pageNumber="72">J. S. Ashe</collectorName>
,
<collectorName id="26F45361A96A196FAB1DFF73420D64E0" box="[1123,1286,187,213]" pageId="11" pageNumber="72">R. A. Leschen</collectorName>
, #213, ex., at light” (
<collectionCode id="ED10AE72A96A196FAF98FF17463364CC" box="[230,312,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="72" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A196FAFB9FECA43B06550" pageId="11" pageNumber="72" type="diagnosis">
<paragraph id="8BBE36B7A96A196FAFB9FECA43B06550" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFB9FECA46D86529" bold="true" box="[199,467,258,284]" pageId="11" pageNumber="72">Differential Diagnosis.</emphasis>
Differentiated from other Peruvian species by the combination of the small body size (ca.
<quantity id="4CF99B52A96A196FAFB8FEEF46276574" box="[198,300,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="11" pageNumber="72" unit="mm" value="2.11">2.11 mm</quantity>
), the narrow body form (width/length ca. 0.33), the numerous dorsal microsetae, and the male genitalia (
<figureCitation id="133A2A32A96A196FAFDEFE8347EF6550" box="[160,228,331,357]" captionStart="FIGURE 7" captionStartId="10.[151,250,2004,2028]" captionTargetBox="[164,1421,461,1966]" captionTargetId="figure-66@10.[151,1435,453,1981]" captionTargetPageId="10" captionText="FIGURE 7. Hydrochus cracentis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417297" httpUri="https://zenodo.org/record/4417297/files/figure.png" pageId="11" pageNumber="72">Fig. 7</figureCitation>
; described below). Completely reliable determinations will include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A196FAFB9FEA643BF606C" pageId="11" pageNumber="72" type="description">
<paragraph id="8BBE36B7A96A196FAFB9FEA642556608" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFB9FEA6465B65BD" bold="true" box="[199,336,366,392]" pageId="11" pageNumber="72">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A96A196FAD23FEA745A965BC" box="[605,674,367,393]" captionStart="FIGURE 7" captionStartId="10.[151,250,2004,2028]" captionTargetBox="[164,1421,461,1966]" captionTargetId="figure-66@10.[151,1435,453,1981]" captionTargetPageId="10" captionText="FIGURE 7. Hydrochus cracentis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417297" httpUri="https://zenodo.org/record/4417297/files/figure.png" pageId="11" pageNumber="72">Fig. 7</figureCitation>
). Size:
<typeStatus id="54BA8815A96A196FAD86FEA7445265BC" box="[760,857,367,393]" pageId="11" pageNumber="72" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.11/0.71; head width 0.54; pronotum l/w 0.49/0.47; PA 0.47; PB 0.40; elytra 1.33/0.71. Body size small (ca.
<quantity id="4CF99B52A96A196FAA13FE5B47CD65E4" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="11" pageNumber="72" unit="mm" value="2.11">2.11 mm</quantity>
), very narrow. Dorsum brown to copper (
<typeStatus id="54BA8815A96A196FADE8FE7F45F265E4" box="[662,761,439,465]" pageId="11" pageNumber="72" type="holotype">holotype</typeStatus>
teneral), with slight to moderately strong iridescence, elytra with black spots. Dorsum with numerous microsetae. Legs testaceous, with tibiofemoral joints and basal ½ of femora darker. Punctation on elytra ca. 23x that of pronotum. Elytra interstriae ca. 0.51x strial puncture diameter. Interstria 9
<superScript id="7C749BFFA96A196FAFDBFDEA47B96605" attach="left" box="[165,178,546,560]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
more convex than others, overhanging 10
<superScript id="7C749BFFA96A196FADFBFDEA45996605" attach="left" box="[645,658,546,560]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA96A196FAB62FDEA43226605" attach="left" box="[1052,1065,546,560]" fontSize="6" pageId="11" pageNumber="72">th</superScript>
very weakly if at all raised.
</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FD8F4561669C" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">Pronotum very slightly longer than wide, widest at anterior margin, narrowed at base, sides very weakly arcuate, almost straight, smooth; anterior margin arcuate; depressions moderately deep, coarsely densely punctate, reliefs between depressions less punctate.</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FD7B445166C4" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">Elytra with spaces between strial punctures ca. 0.51x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FD3343FC6720" blockId="11.[151,1437,150,1149]" box="[199,1271,763,789]" pageId="11" pageNumber="72">Ventral characters: Mentum densely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FCD7428C67FC" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">
Male genitalia (
<figureCitation id="133A2A32A96A196FAE09FCD746B1670C" box="[375,442,799,825]" captionStart="FIGURE 7" captionStartId="10.[151,250,2004,2028]" captionTargetBox="[164,1421,461,1966]" captionTargetId="figure-66@10.[151,1435,453,1981]" captionTargetPageId="10" captionText="FIGURE 7. Hydrochus cracentis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417297" httpUri="https://zenodo.org/record/4417297/files/figure.png" pageId="11" pageNumber="72">Fig. 7</figureCitation>
) general characters: genitalia very small, wider in dorsal/ventral views than lateral view; paramere tips extending slightly beyond tip of aedeagus, in dorsal/ventral views slightly widened, arcuate toward midline, with apical portions separated one from the other, in lateral view tips straight, distal end rounded; aedeagus broad in both dorsal/ventral and lateral views, in distal ½ much wider than parameres in dorsal/ventral views; basal piece longer and slightly wider than parameres; basal piece much wider in dorsal/ventral views than lateral view.
</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FC1B429267D8" blockId="11.[151,1437,150,1149]" box="[199,1433,979,1005]" pageId="11" pageNumber="72">Dorsal surface: pdmm slightly separated from alm in distal portion, contiguous or nearly so for remainder of length.</paragraph>
<paragraph id="8BBE36B7A96A196FAFB9FC3F43BF606C" blockId="11.[151,1437,150,1149]" pageId="11" pageNumber="72">Ventral surface: ventrally parameres in distal ½ distinctly wider than dorsal part of parameres, pvmm in this area very thin and nearly straight, overlapping alm; pvmm in proximal ½ becoming narrowly separate one from the other and with thin, finely striated ridge; pseu small, appear to be between parameres and aedeagus.</paragraph>
</subSubSection>
<subSubSection id="C31B653CA96A196FAFB9FBAB445F6048" box="[199,852,1123,1149]" pageId="11" pageNumber="72" type="etymology">
<paragraph id="8BBE36B7A96A196FAFB9FBAB445F6048" blockId="11.[151,1437,150,1149]" box="[199,852,1123,1149]" pageId="11" pageNumber="72">
<emphasis id="B975EAA5A96A196FAFB9FBAB46456048" bold="true" box="[199,334,1123,1149]" pageId="11" pageNumber="72">Etymology.</emphasis>
Named in reference to the slender body form.
</paragraph>
</subSubSection>
</treatment>
</document>